Bivalvia | Unionida | Hyrioidea | Hyriidae | Hyriinae | Hyridellini

genus Hyridella Swainson, 1840: cladomics

7 Recent valid species (10 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 7 (10 total)
analyzed in 46 cladograms in 28 publications
maximum number of terminals in an analysis = 3 (mean = 1.52, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 16
maximum number of included terminals = 3, (mean = 2.5 median = 2.5)
proportion of cladograms that support basic monophyly = 0.44

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyridella cladogram distances
PCoA Ordination

NJ Tree of Hyridella cladogram distances
NJ Tree

Cladograms

Graf et al., 2015 [Publ 3619], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.667 3 4 2 3 17 1 1 **
Cladomics cladogram Guyanhyrida Triplodon Castalia Hyridella depressa Hyridella drapeta Hyridella australis Cucumerunio

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

cst iT tT bi riS oT roT roS iD
0.333 3 20 20 3 18 1 1 19.0
Cladomics cladogram Neotrigonia Westralunio Diplodon Hyridella drapeta Hyridella depressa Hyridella australis Cucumerunio Echyridella Velesunio ambiguus Microdontia Lortiella Alathyria Velesunio angasi Guyanhyrida Triplodon Prisodon Castalia stevensi Callonaia Castalia ambigua 1 Castalia ambigua 4 Castalia ambigua 2 Castaliella Castalia ambigua 3

Santos-Neto et al., 2016 [Publ 3739], fig. 4

cst iT tT bi riS oT roT roS iD
0.333 3 13 11 3 18 1 1 9.0
Cladomics cladogram Guyanhyrida Triplodon Prisodon Castalia ambigua 4 Callonaia Castalia ambigua 3 Castalia stevensi Castalia ambigua 2 Castaliella Castalia ambigua 1 Westralunio Diplodon Hyridella drapeta Hyridella depressa Echyridella Cucumerunio Hyridella australis Velesunio ambiguus Alathyria Velesunio angasi Lortiella Microdontia

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.333 3 18 20 3 630 1 1 18.0
Cladomics cladogram Guyanhyrida Triplodon Prisodon Castalia ambigua 4 Callonaia Castalia stevensi Castalia ambigua 3 Castalia ambigua 1 Castaliella Castalia ambigua 2 Castalia Diplodon Hyridella depressa Hyridella drapeta Cucumerunio Hyridella australis Echyridella Velesunio 1 Alathyria jacksoni 1 Velesunio ambiguus Alathyria profuga Velesunio 3 Microdontia Lortiella Velesunio angasi Alathyria pertexta 2 Velesunio 2 Alathyria jacksoni 2 Alathyria pertexta 1

Lawrie et al., 2026 [Publ 4843], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 3 4 2 3 15 1 1 0
Cladomics cladogram Castalia Triplodon Hyridella depressa Hyridella drapeta Hyridella australis Cucumerunio Westralunio Velesunio 6 Alathyria jacksoni 2 Alathyria profuga Velesunio ambiguus Velesunio 5 Alathyria pertexta 2 Velesunio 4 Alathyria jacksoni 1 Alathyria pertexta 1 Lortiella Velesunio 3 Microdontia Velesunio angasi Velesunio 1 Velesunio 2

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 198 0 0 1.0
Cladomics cladogram Guyanhyrida Castalia Triplodon Hyridella australis Hyridella depressa Hyridella drapeta

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 196 0 0 1.0
Cladomics cladogram Guyanhyrida Castalia Triplodon Hyridella australis Hyridella depressa Hyridella drapeta

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 198 0 0 1.0
Cladomics cladogram Echyridella Westralunio Guyanhyrida Castalia Triplodon Hyridella australis Hyridella depressa Hyridella drapeta Velesunio Alathyria jacksoni Alathyria profuga Alathyria pertexta Lortiella Microdontia

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
0.000 2 3 2 2 9 0 0 4.0
Cladomics cladogram Diplodon Castalia Hyridella australis 2 Hyridella australis 1 Echyridella Hyridella depressa

Baker et al., 2003 [Publ 2690], fig. 5

cst iT tT bi riS oT roT roS iD
0.000 2 7 0 2 6 0 0 7.0
Cladomics cladogram Epioblasma Hyridella australis Diplodon Hyridella depressa Velesunio Echyridella Castalia

Baker et al., 2004 [Publ 2685], fig. 2-3

cst iT tT bi riS oT roT roS iD
0.000 2 3 0 2 7 0 0 3.0
Cladomics cladogram Hyridella depressa Hyridella australis Diplodon Echyridella Castalia Velesunio 3 Velesunio 4 Velesunio 2 Velesunio angasi Velesunio 1 Alathyria

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 45 0 0 --
Cladomics cladogram Hyridella depressa Hyridella australis Castalia Diplodon

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.000 2 8 0 2 34 0 0 10.0
Cladomics cladogram Castalia Diplodon Hyridella depressa Hyridella australis Echyridella Velesunio ambiguus Velesunio 3 Velesunio 2 Alathyria Velesunio 1 Velesunio angasi Lortiella

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 45 0 0 --
Cladomics cladogram Hyridella depressa Hyridella australis Castalia Diplodon

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 44 0 0 --
Cladomics cladogram Hyridella depressa Hyridella australis Castalia Diplodon

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 44 0 0 --
Cladomics cladogram Hyridella depressa Hyridella australis Castalia Diplodon

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 1 1 --
Cladomics cladogram Cucumerunio Hyridella australis

Prié et al., 2026 [Publ 4844], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 1 1 --
Cladomics cladogram Hyridella australis Cucumerunio

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 0 0 --
Cladomics cladogram Hyridella Velesunio

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Hyridella depressa Margaritifera Cumberlandia Unio Anodonta Alasmidonta Lasmigona Strophitus Pyganodon Quadrula Eurynia Pleurobema Fusconaia Amblema Ligumia Truncilla Ptychobranchus Sagittunio Lampsilis siliquoidea Epioblasma Ortmanniana Cambarunio Leaunio Lampsilis cardium Lampsilis fasciola

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Hyridella depressa Margaritifera Cumberlandia Unio Anodonta Alasmidonta Lasmigona Strophitus Pyganodon Quadrula Eurynia Pleurobema Fusconaia Amblema Truncilla Ligumia Ptychobranchus Sagittunio Lampsilis siliquoidea Ortmanniana Lampsilis cardium Lampsilis fasciola Epioblasma Cambarunio Leaunio

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Hyridella depressa Diplodon Castalia

Walker et al., 2006 [Publ 2666], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Hyridella Anodonta Pyganodon Pronodularia Gonidea Pseudodon Fusconaia Cumberlandia Mesonaias Potamilus Ligumia Ortmanniana Lampsilis

Walker et al., 2006 [Publ 2666], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Hyridella Cumberlandia Pseudodon Gonidea Inversidens Anodonta Pyganodon Fusconaia Potamilus Mesonaias Ligumia Ortmanniana Lampsilis

Walker et al., 2006 [Publ 2666], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Hyridella Cumberlandia Pseudodon Gonidea Inversidens Anodonta Pyganodon Fusconaia Mesonaias Potamilus Ligumia Ortmanniana Lampsilis

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Hyridella australis Castalia Triplodon

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Hyridella australis Castalia Triplodon

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 0 0 --
Cladomics cladogram Hyridella australis Triplodon Castalia

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 0 0 --
Cladomics cladogram Hyridella depressa Velesunio

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 212 0 0 --
Cladomics cladogram Velesunio Hyridella australis

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 213 0 0 --
Cladomics cladogram Triplodon Hyridella australis

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 28 0 0 --
Cladomics cladogram Velesunio Hyridella australis

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Velesunio Hyridella australis

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Velesunio Hyridella australis

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram Hyridella australis Margaritifera Quadrula Uniomerus Elliptoideus Elliptio Pleurobema Reginaia Plectomerus Amblema Psoronaias Popenaias Obliquaria Cyrtonaias Leaunio Lampsilis Pachynaias Chamberlainia Sinohyriopsis Lamprotula Gonidea Indopseudodon Bineurus Pilsbryoconcha Pseudodon Sundadontina Ensidens Hyriopsis Rectidens Trapezoideus Physunio Yaukthwa Lens Solenaia Lanceolaria Unio Nodularia Tchangsinaia Cristaria Sinanodonta Anodonta Pyganodon Trapezidens Lamellidens Leoparreysia Coelatura Prisodontopsis Indonaia Indochinella Scabies Harmandia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram Hyridella australis Margaritifera

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 0 0 --
Cladomics cladogram Hyridella depressa Velesunio

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Hyridella australis Velesunio

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Hyridella australis Velesunio

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Hyridella australis Velesunio

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Hyridella australis Velesunio

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Hyridella australis Velesunio

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Triplodon Hyridella australis Velesunio

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Triplodon Velesunio Hyridella australis

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Velesunio Hyridella australis

Chen et al., 2025 [Publ 4455], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 25 0 0 --
Cladomics cladogram Hyridella australis Triplodon