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Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Buldowskia Moskvicheva, 1973: cladomics
     5 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (5 total)
analyzed in 21 cladograms in 11 publications
maximum number of terminals in an analysis = 6 (mean = 2.57, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 11
maximum number of included terminals = 6, (mean = 4 median = 5)
proportion of cladograms that support basic monophyly = 1


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Wu et al., 2025 [Publ 4442], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
6646292770
  +-------------------------------- Cristaria                  
  |                                                            
  |                          +----- Beringiana                 
--+  +-----------------------+                                 
  |  |                       |  +-- Acudonta                   
  |  |                       +--+                              
  |  |                          +-- Sinanodonta                
  +--+                                                         
     |                       +----- Pletholophus_guangzhouensis
     |  +--------------------+                                 
     |  |                    |  +-- Pletholophus_tenuis        
     |  |                    +--+                              
     +--+                       +-- Pletholophus_reinianus     
        |                                                      
        |  +----------------------- Pletholophus_honglinhensis 
        +--+                                                   
           |  +-------------------- Anemina                    
           +--+                                                
              |  +----------------- Amuranodonta               
              +--+                                             
                 |  +-------------- Buldowskia_suifunica **       
                 +--+                                          
                    |  +----------- Buldowskia_shadini **         
                    +--+                                       
                       |  +-------- Buldowskia_flavotincta **     
                       +--+                                    
                          |  +----- Buldowskia_kamiyai **         
                          +--+                                 
                             |  +-- Buldowskia_iwakawai **        
                             +--+                              
                                +-- Buldowskia **                 

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
5525262262
           +-------- Cristaria             
  +--------+                               
  |        |  +----- Pletholophus          
  |        +--+                            
  |           |  +-- Beringiana            
--+           +--+                         
  |              +-- Sinanodonta           
  |                                        
  |  +-------------- Amuranodonta          
  |  |                                     
  +--+  +----------- Anemina               
     |  |                                  
     +--+  +-------- Buldowskia_suifunica **  
        |  |                               
        +--+  +----- Buldowskia_flavotincta **
           |  |                            
           +--+----- Buldowskia_shadini **    
              |                            
              |  +-- Buldowskia_iwakawai **   
              +--+                         
                 +-- Buldowskia_kamiyai **    

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

iTtTbiriSoTroTroSiD
55356435
  +----------------- Sinanodonta           
  |                                        
--+  +-------------- Amuranodonta          
  |  |                                     
  +--+  +----------- Anemina               
     |  |                                  
     +--+        +-- Buldowskia_iwakawai **   
        |  +-----+                         
        |  |     +-- Buldowskia_kamiyai **    
        +--+                               
           |  +----- Buldowskia_shadini **    
           +--+                            
              |  +-- Buldowskia_suifunica **  
              +--+                         
                 +-- Buldowskia_flavotincta **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
5535262263
  +-------------------- Cristaria             
  |                                           
  |              +----- Pletholophus          
--+  +-----------+                            
  |  |           |  +-- Beringiana            
  |  |           +--+                         
  +--+              +-- Sinanodonta           
     |                                        
     |  +-------------- Amuranodonta          
     |  |                                     
     +--+  +----------- Anemina               
        |  |                                  
        +--+  +-------- Buldowskia_suifunica **  
           |  |                               
           +--+     +-- Buldowskia_flavotincta **
              |  +--+                         
              |  |  +-- Buldowskia_shadini **    
              +--+                            
                 |  +-- Buldowskia_iwakawai **   
                 +--+                         
                    +-- Buldowskia_kamiyai **    

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
55351161963
  +----------------- Cristaria             
  |                                        
  |           +----- Pletholophus          
  +-----------+                            
  |           |  +-- Beringiana            
--+           +--+                         
  |              +-- Sinanodonta           
  |                                        
  |  +-------------- Anemina               
  |  |                                     
  +--+  +----------- Amuranodonta          
     |  |                                  
     +--+  +-------- Buldowskia_suifunica **  
        |  |                               
        +--+     +-- Buldowskia_flavotincta **
           |  +--+                         
           |  |  +-- Buldowskia_shadini **    
           +--+                            
              |  +-- Buldowskia_iwakawai **   
              +--+                         
                 +-- Buldowskia_kamiyai **    

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
5535262263
           +-------- Cristaria             
  +--------+                               
  |        |  +----- Pletholophus          
  |        +--+                            
  |           |  +-- Beringiana            
--+           +--+                         
  |              +-- Sinanodonta           
  |                                        
  |  +-------------- Amuranodonta          
  |  |                                     
  +--+  +----------- Anemina               
     |  |                                  
     +--+  +-------- Buldowskia_suifunica **  
        |  |                               
        +--+     +-- Buldowskia_iwakawai **   
           |  +--+                         
           |  |  +-- Buldowskia_kamiyai **    
           +--+                            
              |  +-- Buldowskia_flavotincta **
              +--+                         
                 +-- Buldowskia_shadini **    

Xiang et al., 2025 [Publ 4433], fig. 6

iTtTbiriSoTroTroSiD
5535232273
  +----------------------- Pletholophus          
  |                                              
  |                 +----- Beringiana            
--+  +--------------+                            
  |  |              |  +-- Mimo                  
  |  |              +--+                         
  +--+                 +-- Sinanodonta           
     |                                           
     |  +----------------- Cristaria             
     |  |                                        
     +--+  +-------------- Anemina               
        |  |                                     
        +--+  +----------- Amuranodonta          
           |  |                                  
           +--+  +-------- Buldowskia_suifunica **  
              |  |                               
              +--+     +-- Buldowskia_flavotincta **
                 |  +--+                         
                 |  |  +-- Buldowskia_shadini **    
                 +--+                            
                    |  +-- Buldowskia_iwakawai **   
                    +--+                         
                       +-- Buldowskia_kamiyai **    

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
22023475--
        +----- Cristaria           
  +-----+                          
  |     |  +-- Beringiana          
  |     +--+                       
--+        +-- Sinanodonta         
  |                                
  |  +-------- Amuranodonta        
  +--+                             
     |  +----- Anemina             
     +--+                          
        |  +-- Buldowskia_suifunica **
        +--+                       
           +-- Buldowskia_shadini **  

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
220254115--
        +----- Cristaria           
  +-----+                          
  |     |  +-- Beringiana          
  |     +--+                       
--+        +-- Sinanodonta         
  |                                
  |  +-------- Anemina             
  +--+                             
     |  +----- Amuranodonta        
     +--+                          
        |  +-- Buldowskia_suifunica **
        +--+                       
           +-- Buldowskia_shadini **  

Sano et al., 2022 [Publ 4311], fig. 3

iTtTbiriSoTroTroSiD
220216164--
           +-- Beringiana         
  +--------+                      
  |        +-- Sinanodonta        
--+                               
  |  +-------- Pletholophus       
  +--+                            
     |  +----- Cristaria          
     +--+                         
        |  +-- Buldowskia_kamiyai ** 
        +--+                      
           +-- Buldowskia_iwakawai **

Sano et al., 2022 [Publ 4311], fig. 2

iTtTbiriSoTroTroSiD
220210104--
           +-- Beringiana         
  +--------+                      
  |        +-- Sinanodonta        
--+                               
  |  +-------- Pletholophus       
  +--+                            
     |  +----- Cristaria          
     +--+                         
        |  +-- Buldowskia_iwakawai **
        +--+                      
           +-- Buldowskia_kamiyai ** 

Soroka, 2010 [Publ 3404], fig. 9a

iTtTbiriSoTroTroSiD
1101442--
  +----- Buldowskia_flavotincta **
--+                            
  |  +-- Beringiana            
  +--+                         
     +-- Sinanodonta           

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
110167105--
  +-- Anemina            
--+                      
  +-- Buldowskia_iwakawai **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
11013284--
        +-- Cristaria          
  +-----+                      
  |     +-- Buldowskia_iwakawai **
--+                            
  |  +----- Beringiana         
  +--+                         
     |  +-- Pletholophus       
     +--+                      
        +-- Sinanodonta        

Soroka, 2010 [Publ 3404], fig. 9b

iTtTbiriSoTroTroSiD
1101331--
  +-- Buldowskia_flavotincta **
--+                         
  +-- Sinanodonta           

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
12202339537
  +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 ** 
  |                             
  +----- Buldowskia_iwakawai__2 ** 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio            

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
11012074--
        +-- Cristaria          
  +-----+                      
  |     +-- Buldowskia_iwakawai **
--+                            
  |  +----- Pletholophus       
  +--+                         
     |  +-- Sinanodonta        
     +--+                      
        +-- Beringiana         

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
110167105--
  +-- Anemina            
--+                      
  +-- Buldowskia_iwakawai **

Wu et al., 2025 [Publ 4442], fig. 6

iTtTbiriSoTroTroSiD
110116147--
  +----------- Cristaria         
  |                              
  |     +----- Amuranodonta      
--+  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  |  |     +-- Buldowskia_shadini **
  +--+                           
     |     +-- Beringiana        
     |  +--+                     
     |  |  +-- Pletholophus      
     +--+                        
        |  +-- Acudonta          
        +--+                     
           +-- Sinanodonta       

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
110167105--
  +-- Anemina            
--+                      
  +-- Buldowskia_iwakawai **

Soroka, 2010 [Publ 3404], fig. 9c

iTtTbiriSoTroTroSiD
1101331--
  +-- Buldowskia_flavotincta **
--+                         
  +-- Sinanodonta           
 
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