| The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win! 
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 Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina genus Buldowskia Moskvicheva, 1973: cladomics5 Recent valid species (5 total species)
 Cladomics pages are experimental — even more so than the rest of this site! The purpose of the cladomics pages for each genus and family-group level taxon is to 
    provide statistics and references indicating the extent to which published phylogenetic 
    work supports the classification used here.  The Cladomics Summary provides an 
    overview of the number of studies that have included terminal species representing this 
    taxon as well as an indication of how frequently it has been recovered as monophyletic.  Following the summary, the available published clades representing the taxon are listed.  For more information, visit the Cladomics FARQ (frequently 
    asked rhetorical questions) page. Abbreviations for the various statistics are explained on 
    the FARQ! Cladomics Summarynumber of species = 5 (5 total)analyzed in 21 cladograms in 11 publications
 maximum number of terminals in an analysis = 
            6 (mean = 2.57, median = 2)
 Basic Monophylynumber of cladograms that test basic monophyly 
                (i.e., more than 1 included species) = 11maximum number of included terminals = 6, (mean = 
            4 median = 5)
 proportion of cladograms that support basic monophyly 
            = 1
 
 Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup 
        distances within the cladome. See the Cladmoics 
        FARQ for more information. 
 Wu et al., 2025 [Publ 4442], fig. 2 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** | iT | tT | bi | riS | oT | roT | roS | iD |  | 6 | 6 | 4 | 6 | 29 | 27 | 7 | 0 | 
   +-------------------------------- Cristaria                  
  |                                                            
  |                          +----- Beringiana                 
--+  +-----------------------+                                 
  |  |                       |  +-- Acudonta                   
  |  |                       +--+                              
  |  |                          +-- Sinanodonta                
  +--+                                                         
     |                       +----- Pletholophus_guangzhouensis
     |  +--------------------+                                 
     |  |                    |  +-- Pletholophus_tenuis        
     |  |                    +--+                              
     +--+                       +-- Pletholophus_reinianus     
        |                                                      
        |  +----------------------- Pletholophus_honglinhensis 
        +--+                                                   
           |  +-------------------- Anemina                    
           +--+                                                
              |  +----------------- Amuranodonta               
              +--+                                             
                 |  +-------------- Buldowskia_suifunica **       
                 +--+                                          
                    |  +----------- Buldowskia_shadini **         
                    +--+                                       
                       |  +-------- Buldowskia_flavotincta **     
                       +--+                                    
                          |  +----- Buldowskia_kamiyai **         
                          +--+                                 
                             |  +-- Buldowskia_iwakawai **        
                             +--+                              
                                +-- Buldowskia **                 
Dai et al., 2024 [Publ 4385], fig. 3-BI | iT | tT | bi | riS | oT | roT | roS | iD |  | 5 | 5 | 2 | 5 | 26 | 22 | 6 | 2 | 
            +-------- Cristaria             
  +--------+                               
  |        |  +----- Pletholophus          
  |        +--+                            
  |           |  +-- Beringiana            
--+           +--+                         
  |              +-- Sinanodonta           
  |                                        
  |  +-------------- Amuranodonta          
  |  |                                     
  +--+  +----------- Anemina               
     |  |                                  
     +--+  +-------- Buldowskia_suifunica **  
        |  |                               
        +--+  +----- Buldowskia_flavotincta **
           |  |                            
           +--+----- Buldowskia_shadini **    
              |                            
              |  +-- Buldowskia_iwakawai **   
              +--+                         
                 +-- Buldowskia_kamiyai **    
Lopes-Lima et al., 2020 [Publ 4080], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD |  | 5 | 5 | 3 | 5 | 6 | 4 | 3 | 5 | 
   +----------------- Sinanodonta           
  |                                        
--+  +-------------- Amuranodonta          
  |  |                                     
  +--+  +----------- Anemina               
     |  |                                  
     +--+        +-- Buldowskia_iwakawai **   
        |  +-----+                         
        |  |     +-- Buldowskia_kamiyai **    
        +--+                               
           |  +----- Buldowskia_shadini **    
           +--+                            
              |  +-- Buldowskia_suifunica **  
              +--+                         
                 +-- Buldowskia_flavotincta **
Bogan et al., 2023 [Publ 4295], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD |  | 5 | 5 | 3 | 5 | 26 | 22 | 6 | 3 | 
   +-------------------- Cristaria             
  |                                           
  |              +----- Pletholophus          
--+  +-----------+                            
  |  |           |  +-- Beringiana            
  |  |           +--+                         
  +--+              +-- Sinanodonta           
     |                                        
     |  +-------------- Amuranodonta          
     |  |                                     
     +--+  +----------- Anemina               
        |  |                                  
        +--+  +-------- Buldowskia_suifunica **  
           |  |                               
           +--+     +-- Buldowskia_flavotincta **
              |  +--+                         
              |  |  +-- Buldowskia_shadini **    
              +--+                            
                 |  +-- Buldowskia_iwakawai **   
                 +--+                         
                    +-- Buldowskia_kamiyai **    
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 | iT | tT | bi | riS | oT | roT | roS | iD |  | 5 | 5 | 3 | 5 | 116 | 19 | 6 | 3 | 
   +----------------- Cristaria             
  |                                        
  |           +----- Pletholophus          
  +-----------+                            
  |           |  +-- Beringiana            
--+           +--+                         
  |              +-- Sinanodonta           
  |                                        
  |  +-------------- Anemina               
  |  |                                     
  +--+  +----------- Amuranodonta          
     |  |                                  
     +--+  +-------- Buldowskia_suifunica **  
        |  |                               
        +--+     +-- Buldowskia_flavotincta **
           |  +--+                         
           |  |  +-- Buldowskia_shadini **    
           +--+                            
              |  +-- Buldowskia_iwakawai **   
              +--+                         
                 +-- Buldowskia_kamiyai **    
Dai et al., 2024 [Publ 4385], fig. 3-ML | iT | tT | bi | riS | oT | roT | roS | iD |  | 5 | 5 | 3 | 5 | 26 | 22 | 6 | 3 | 
            +-------- Cristaria             
  +--------+                               
  |        |  +----- Pletholophus          
  |        +--+                            
  |           |  +-- Beringiana            
--+           +--+                         
  |              +-- Sinanodonta           
  |                                        
  |  +-------------- Amuranodonta          
  |  |                                     
  +--+  +----------- Anemina               
     |  |                                  
     +--+  +-------- Buldowskia_suifunica **  
        |  |                               
        +--+     +-- Buldowskia_iwakawai **   
           |  +--+                         
           |  |  +-- Buldowskia_kamiyai **    
           +--+                            
              |  +-- Buldowskia_flavotincta **
              +--+                         
                 +-- Buldowskia_shadini **    
Xiang et al., 2025 [Publ 4433], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD |  | 5 | 5 | 3 | 5 | 23 | 22 | 7 | 3 | 
   +----------------------- Pletholophus          
  |                                              
  |                 +----- Beringiana            
--+  +--------------+                            
  |  |              |  +-- Mimo                  
  |  |              +--+                         
  +--+                 +-- Sinanodonta           
     |                                           
     |  +----------------- Cristaria             
     |  |                                        
     +--+  +-------------- Anemina               
        |  |                                     
        +--+  +----------- Amuranodonta          
           |  |                                  
           +--+  +-------- Buldowskia_suifunica **  
              |  |                               
              +--+     +-- Buldowskia_flavotincta **
                 |  +--+                         
                 |  |  +-- Buldowskia_shadini **    
                 +--+                            
                    |  +-- Buldowskia_iwakawai **   
                    +--+                         
                       +-- Buldowskia_kamiyai **    
Bolotov et al., 2020 [Publ 4084], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD |  | 2 | 2 | 0 | 2 | 34 | 7 | 5 | -- | 
         +----- Cristaria           
  +-----+                          
  |     |  +-- Beringiana          
  |     +--+                       
--+        +-- Sinanodonta         
  |                                
  |  +-------- Amuranodonta        
  +--+                             
     |  +----- Anemina             
     +--+                          
        |  +-- Buldowskia_suifunica **
        +--+                       
           +-- Buldowskia_shadini **  
Konopleva et al., 2023 [Publ 4269], fig. S1_4 | iT | tT | bi | riS | oT | roT | roS | iD |  | 2 | 2 | 0 | 2 | 54 | 11 | 5 | -- | 
         +----- Cristaria           
  +-----+                          
  |     |  +-- Beringiana          
  |     +--+                       
--+        +-- Sinanodonta         
  |                                
  |  +-------- Anemina             
  +--+                             
     |  +----- Amuranodonta        
     +--+                          
        |  +-- Buldowskia_suifunica **
        +--+                       
           +-- Buldowskia_shadini **  
Sano et al., 2022 [Publ 4311], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD |  | 2 | 2 | 0 | 2 | 16 | 16 | 4 | -- | 
            +-- Beringiana         
  +--------+                      
  |        +-- Sinanodonta        
--+                               
  |  +-------- Pletholophus       
  +--+                            
     |  +----- Cristaria          
     +--+                         
        |  +-- Buldowskia_kamiyai ** 
        +--+                      
           +-- Buldowskia_iwakawai **
Sano et al., 2022 [Publ 4311], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD |  | 2 | 2 | 0 | 2 | 10 | 10 | 4 | -- | 
            +-- Beringiana         
  +--------+                      
  |        +-- Sinanodonta        
--+                               
  |  +-------- Pletholophus       
  +--+                            
     |  +----- Cristaria          
     +--+                         
        |  +-- Buldowskia_iwakawai **
        +--+                      
           +-- Buldowskia_kamiyai ** 
Soroka, 2010 [Publ 3404], fig. 9a | iT | tT | bi | riS | oT | roT | roS | iD |  | 1 | 1 | 0 | 1 | 4 | 4 | 2 | -- | 
   +----- Buldowskia_flavotincta **
--+                            
  |  +-- Beringiana            
  +--+                         
     +-- Sinanodonta           
Sano et al., 2020 [Publ 4138], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD |  | 1 | 1 | 0 | 1 | 67 | 10 | 5 | -- | 
   +-- Anemina            
--+                      
  +-- Buldowskia_iwakawai **
Sano et al., 2017 [Publ 3799], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD |  | 1 | 1 | 0 | 1 | 32 | 8 | 4 | -- | 
         +-- Cristaria          
  +-----+                      
  |     +-- Buldowskia_iwakawai **
--+                            
  |  +----- Beringiana         
  +--+                         
     |  +-- Pletholophus       
     +--+                      
        +-- Sinanodonta        
Soroka, 2010 [Publ 3404], fig. 9b | iT | tT | bi | riS | oT | roT | roS | iD |  | 1 | 1 | 0 | 1 | 3 | 3 | 1 | -- | 
   +-- Buldowskia_flavotincta **
--+                         
  +-- Sinanodonta           
Sano et al., 2020 [Publ 4138], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD |  | 1 | 22 | 0 | 2 | 33 | 9 | 5 | 37 | 
   +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 ** 
  |                             
  +----- Buldowskia_iwakawai__2 ** 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio            
Sano et al., 2017 [Publ 3799], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD |  | 1 | 1 | 0 | 1 | 20 | 7 | 4 | -- | 
         +-- Cristaria          
  +-----+                      
  |     +-- Buldowskia_iwakawai **
--+                            
  |  +----- Pletholophus       
  +--+                         
     |  +-- Sinanodonta        
     +--+                      
        +-- Beringiana         
Sano et al., 2020 [Publ 4138], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD |  | 1 | 1 | 0 | 1 | 67 | 10 | 5 | -- | 
   +-- Anemina            
--+                      
  +-- Buldowskia_iwakawai **
Wu et al., 2025 [Publ 4442], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD |  | 1 | 1 | 0 | 1 | 16 | 14 | 7 | -- | 
   +----------- Cristaria         
  |                              
  |     +----- Amuranodonta      
--+  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  |  |     +-- Buldowskia_shadini **
  +--+                           
     |     +-- Beringiana        
     |  +--+                     
     |  |  +-- Pletholophus      
     +--+                        
        |  +-- Acudonta          
        +--+                     
           +-- Sinanodonta       
Sano et al., 2020 [Publ 4138], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD |  | 1 | 1 | 0 | 1 | 67 | 10 | 5 | -- | 
   +-- Anemina            
--+                      
  +-- Buldowskia_iwakawai **
Soroka, 2010 [Publ 3404], fig. 9c | iT | tT | bi | riS | oT | roT | roS | iD |  | 1 | 1 | 0 | 1 | 3 | 3 | 1 | -- | 
   +-- Buldowskia_flavotincta **
--+                         
  +-- Sinanodonta           
 |