MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Wed 04 February 2026
Bivalvia | Unionida | Unionoidea | Margaritiferidae | Margaritiferinae | Margaritifera middendorffi
nominal species Margaritifera togakushiensis Kondo & Kobayashi, 2005
Margaritifera togakushiensis Kondo & Kobayashi, 2005 = Margaritifera middendorffi
Kondo & Kobayashi, 2005. Venus 64(3-4): 136, figs. 5-8.
Type(s): HOLOTYPE (ORIG. DESIG.) NSMT-MO_73707, Sakasa River at Togakushi (136°6’ E, 36°45’ N, 1200 m altitude), Nagano Prefecture, Honshu, Japan.
Taxonomic Opinions
+ Margaritifera kurilensis (Zatravkin et Starobogatov, 1984) — Bogatov, 2022
+ Margaritifera middendorffi (Rosén, 1926) — Graf & Cummings, 2021
= Margaritifera togakushiensis — Sano et al., 2020
= Margaritifera togakushiensis Kondo & Kobayashi, 2005 — Zieritz et al., 2018
= Margaritifera togakushiensis Kondo & Kobayashi, 2005 — Sano et al., 2017
+ Margaritifera (s.s.) middendorffi — Bolotov et al., 2016
+ Margaritifera middendorffi (Rosén, 1926) — Araujo et al., 2016
= Margaritifera togakushiensis Kondo & Kobayashi, 2005 — Bolotov et al., 2015
= Margaritifera togakushiensis — Takeuchi et al., 2015
= Margaritifera togakushiensis Kondo & Kobayashi, 2005 — Bogan, 2010
= Margaritifera togakushiensis Kondo & Kobayashi, 2005 — Kondo, 2008
= Margaritifera togakushiensis Kondo & Kobayashi, 2005 — Graf, 2007
= Margaritifera togakushiensis Kondo & Kobayashi, 2005 — Graf & Cummings, 2007
= Margaritifera togakushiensis Kondo & Kobayahi, 2005 — Kondo & Kobayashi, 2005
