MUSSELpdb
Taxonomy
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freshwater mussels
by Taxonomy
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freshwater bivalves
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freshwater mussels
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Data last updated
Sun 12 April 2026
Publication Germain (1931): species taxonomic opinions
Full Citation
Germain, L. 1931. Mollusques terrestres et fluviatiles. Faune de France 22: 479-897. https://archive.org/details/FauneDeFrance21-22.
Taxonomic Opinions
1-71
Anodonta anatina Linné = Anodonta anatina + Anodonta exulcerata + Anodonta cygnea
+ Anodonta anatina abbreviata Brot, 1867
+ Anodonta anatina Brot, 1867
+ Anodonta acallia ‘J. Ray, in litt.’ Bourguignat, 1881
+ Anodonta coarctata Kobelt, 1879
+ Anodonta aechmopsis Bourguignat, 1881
+ Anodonta alethinia Locard, 1890
+ Anodonta alsatica Locard, 1890
+ Anodonta piscinalis var. anatinella Stabile, 1859
+ Anodonta anceyi Bourguignat in Servain, 1888
+ Anodonta antorida Bourguignat, 1881
75 more...
Anodonta anatina forme subarealis Fagot = Anodonta anatina
+ Anodonta anatina Dupuy, 1850
+ Anodonta scaldina Kobelt, 1879
Anodonta anatina var. chresimella Bourguignat = Anodonta anatina
+ Anodonta autriciaca Locard, 1890
+ Anodonta autricensis Locard, 1890
+ Anodonta marcida Locard, 1890
Anodonta ataxiaca Baichère = Anodonta anatina
Anodonta avonensis Montagu, 1803 = Anodonta cygnea + Anodonta anatina
+ Anodonta avonica Bourguignat, 1881
+ Anodonta aequorea Locard, 1890
+ Anodonta amnica Drouët, 1888
+ Anodonta aresta Locard, 1890
+ Anodonta arvernica Bourguignat, 1881
+ Anodonta calara Servain, 1885
+ Anodonta carisiana ‘Mabille, in litt.’ Bourguignat, 1881
+ Anodonta coenosella Locard, 1890
+ Anodonta coutagnei Locard, 1882
+ Anodonta doei Bourguignat, 1881
41 more...
Anodonta avonensis forme glyca Bourguignat = Anodonta anatina
Anodonta avonensis forme subponderosa Dupuy = Anodonta anatina
+ Anodonta cadomensis Locard, 1890
+ Anodonta ponderosa var. elongata Garnier, 1840
Anodonta avonensis var. carvalhoi Castro = Anodonta anatina
+ Anodonta carvalhopsis Locard, 1890
+ Anodonta bouvieri Rochebrune, 1903
Anodonta avonensis var. ponderiformis Locard = Anodonta anatina
+ Anodonta ponderosa Dupuy, 1850
Anodonta brebissoni Locard = Anodonta anatina
Anodonta cygnaea [sic] Linné = Anodonta cygnea + Anodonta anatina + Pseudanodonta complanata
+ Mytilus zellensis Gmelin, 1791
+ Anodonta cygnaea ‘Linnaeus’ of Authors
+ Mytilus radiatus Müller, 1774
+ Anodonta acyrta Bourguignat, 1881
+ Anodonta annesiaca Locard, 1890
+ Anodonta arelatensis Dupuy, 1850
+ Anodonta cariosa Küster, 1842
+ Anodonta cellensis rostrata Brot, 1867
+ Anodonta cariosula Locard, 1890
+ Anodonta catocyrta Locard, 1890
56 more...
Anodonta cygnaea [sic] forme arenaria Schröter = Anodonta cygnea
+ Anodonta cellensis Pfeiffer, 1821
+ Anodonta cellensis Drouët, 1852
Anodonta cygnaea [sic] forme fragilissima Clessin = Anodonta cygnea
+ Anodonta fragillima Bourguignat, 1881
Anodonta cygnaea [sic] forme maxima Schröter = Anodonta cygnea
+ Anodonta pammegala Bourguignat, 1881
+ Anodonta maxima Drouët, 1889
+ Anodonta cygnea Rossmässler, 1837
Anodonta cygnaea [sic] forme sturmi Bourg. = Anodonta cygnea + Anodonta anatina
+ Anodonta intermedia Pfeiffer, 1821
Anodonta cygnaea [sic] forme ventricosa C. Pfeiffer = Anodonta cygnea
+ Anodonta gallica Bourguignat, 1881
+ Mytilus anatinus Sheppard, 1821
+ Anodonta cellensis Brown, 1845
Anodonta intermedia de Lamarck = Anodonta cygnea + Anodonta anatina
+ Anodonta alsterica Servain, 1888
+ Anodonta arundinum Locard, 1884
+ Anodonta borboraeca Bourguignat in Locard, 1882
+ Anodonta blaca Locard, 1890
+ Anodonta tremula Drouët, 1888
+ Anodonta macrostena var. curta Servain, 1882
+ Anodonta cyrtoptychia Bourguignat, 1881
+ Anodonta gibba Clessin, 1873
+ Anodonta dinellina Locard, 1890
+ Anodonta divinita Locard, 1890
20 more...
Anodonta intermedia var. spondea (Bourguignat) Locard = Anodonta anatina
+ Anodonta armoricana Coutagne, 1895
Anodonta maculata Sheppard = Anodonta cygnea + Anodonta anatina
+ Anodonta arnouldi Bourguignat, 1883
+ Anodonta caletengis Locard, 1890
+ Anodonta circulus Locard, 1890
+ Anodonta gabatiformis Locard, 1890
+ Anodonta labelliformis Locard, 1890
+ Anodonta mea Locard, 1890
+ Anodonta nicolloni Locard, 1890
+ Anodonta nitefacta Locard, 1890
+ Anodonta orivalensis Locard, 1890
+ Anodonta pentagona Locard, 1890
7 more...
Anodonta ponderosa C. Pfeiffer = Anodonta anatina
Anodonta rostrata Kokeil = Anodonta anatina
+ Anodonta charpentierii Küster, 1842
+ Anodonta culoxiana Nicolas, 1890
+ Anodonta helvetica Coutagne, 1895
+ Anodonta limpida Clessin, 1874
+ Anodonta sebinensis Adami in Bourguignat, 1881
+ Anodonta idrina Clessin, 1876
+ Anodonta blanci Bourguignat, 1881
+ Anodonta marconi Locard, 1890
+ Anodonta mitula Locard, 1890
Anodonta rostrata var. campyla (Bourguignat) Locard = Anodonta anatina
Anodonta subrhombea Brown = Anodonta anatina
+ Anodonta elachista Bourguignat, 1865
+ Anodonta piscinalis Drouët, 1852
+ Anodonta avenionensis Locard, 1890
+ Anodonta dantessantyi ‘J. Ray, in litt.’ Bourguignat, 1881
+ Anodonta piscinalis var. testa Rossmässler, 1837
+ Anodonta episema Bourguignat, 1881
+ Anodonta exocha Bourguignat, 1881
+ Mytilus cygneus Schröter, 1779
+ Anodonta formosa Drouët, 1888
+ Anodonta indetrita Locard, 1890
6 more...
Margaritana auricularia (Spengler) = Pseudunio auricularius + Unio crassus
+ Unio margaritifera Draparnaud, 1801
+ Unio rugosa Poiret, 1801
+ Unio crassissima Férussac, 1827
+ Unio arcuata Bouchard-Chantereaux, 1838
+ Unio sinuata Lamarck, 1819
+ Unio araris Barbié of Authors
+ Unio garumnae de Grateloup of Authors
Margaritana margaritifera (Linné) = Margaritifera margaritifera
+ Unio margaritifer var. minor Rossmässler, 1835
+ Unio elongata Lamarck, 1819
+ Unio roissyi Michaud, 1831
+ Unio elongata Michaud, 1831
+ Unio brunneus Bonhomme, 1840
+ Margaritana michaudi Locard, 1889
+ Margaritana pyrenaica Bourguignat in Locard, 1889
Pseudanodonta complanata (Ziegler) Rossmässler
Pseudanodonta dorsuosa Drouët = Pseudanodonta complanata
+ Pseudanodonta dorsuata Locard, 1882
+ Anodonta (Pseudanodonta) arasiana Westerlund, 1890
+ Pseudanodonta locardi Bourguignat in Locard, 1882
+ Pseudanodonta ararisana Bourguignat in Locard, 1882
+ Pseudanodonta arnouldi Bourguignat in Locard, 1890
+ Pseudanodonta cazioti Bourguignat in Locard, 1890
+ Pseudanodonta euthymei Bourguignat in Locard, 1890
+ Pseudanodonta pacomei Bourguignat in Locard, 1890
+ Pseudanodonta pechaudi Bourguignat in Locard, 1890
+ Pseudanodonta rivalis Bourguignat in Locard, 1890
1 more...
Pseudanodonta elongata Holandre = Pseudanodonta complanata
+ Anodonta jobae Dupuy, 1849
+ Anodonta gratelupeana Gassies, 1849
+ Anodonta gratelupeana var. globosa Gassies, 1849
+ Anodonta normandi Dupuy, 1849
+ Pseudanodonta rayi ‘Mabille in litt.’ Bourguignat, 1880
+ Anodonta (Pseudanodonta) albica Drouët, 1893
+ Pseudanodonta aploa Bourguignat in Locard, 1890
+ Pseudanodonta imperialis Bourguignat in Locard, 1890
+ Pseudanodonta isarana Bourguignat in Locard, 1890
+ Pseudanodonta lacustris Bourguignat in Locard, 1890
10 more...
Pseudanodonta elongata var. dumasi Locard = Pseudanodonta complanata
+ Pseudanodonta berryacensis Dumas in Locard, 1894
Pseudanodonta klettii Rossmässler = Pseudanodonta complanata
+ Anodonta minima Joba, 1844
+ Anodonta elongata Joba, 1851
Psilunio littoralis Cuvier = Potomida littoralis
+ Mya crassa Vallot, 1801
+ Unio subtetragonus Michaud, 1831
+ Unio draparnaldii Deshayes, 1831
+ Unio pianensis Farines in Boubée, 1833
+ Unio barraudii Bonhomme, 1840
+ Unio sinuatus Rossmässler, 1836
+ Unio rhomboideus Moquin-Tandon, 1855
+ Unio moulinsianus Dupuy, 1851
+ Unio bigerrensis Millet, 1843
+ Unio bigorriensis Locard, 1882
9 more...
Unio batavus Maton et Rackett = Unio crassus + Unio nanus + Unio tumidus
+ Unio batavus var. courtilieri de Joannis, 1859
+ Unio andegavensis Locard, 1882
+ Unio cyprinorum Bourguignat in Locard, 1882
+ Unio ligericus Bourguignat in Locard, 1882
+ Unio batavus var. ovalis de Joannis, 1859
+ Unio batavus var. longus de Joannis, 1859
+ Unio sequanicus Bourguignat in Locard, 1882
+ Unio diptychus Bourguignat in Locard, 1889
+ Unio ingrandiensis Bourguignat in Locard, 1889
+ Unio matronicus Bourguignat in Locard, 1882
27 more...
Unio batavus var. baraceus de Joannis = Unio crassus
+ Unio batavus var. major Drouët, 1898
+ Unio batavellus Locard, 1885
+ Unio feliciani Bourguignat in Locard, 1882
+ Unio arenarum Bourguignat in Locard, 1882
+ Unio lancelevei Locard, 1893
Unio capigliolo Payraudeau = Unio mancus
+ Unio exauratus Locard, 1889
Unio consentaneus (Ziegler) Rossmässler = Unio nanus + Unio crassus
+ Unio batavus var. squamosus Charpentier, 1837
+ Unio batavus var. squamosus Brot, 1867
+ Unio ignari Bourguignat in Locard, 1889
+ Unio ignariformis Bourguignat in Locard, 1889
+ Unio ater Drouët, 1857
+ Unio danemorae Bourguignat in Locard, 1882
+ Unio lambottei Malzine, 1867
+ Unio moquinianus Drouët, 1852
+ Unio brevierei Locard, 1882
+ Unio margaritifera Draparnaud, 1806
7 more...
Unio consentaneus var. ardusianus Reyniés = Unio mancus
+ Unio turtoni Drouët, 1852
Unio consentaneus var. jacquemini Dupuy = Unio mancus
+ Unio arcuata Jacquemin, 1835
+ Unio fabaeformis Bourguignat in Locard, 1882
+ Unio arelatus Locard, 1893
Unio consentaneus var. moquini = Unio mancus + Unio nanus + Unio pictorum + Unio crassus
+ Unio moquinianus Dupuy, 1843
+ Unio scotinus Locard, 1889
+ Unio requieni var. rostralis de Joannis, 1859
+ Unio stygnus Locard, 1889
+ Unio arcuata Bouchard-Chantereaux, 1838
+ Unio bouchardi Locard, 1889
+ Unio marcellinus Bourguignat in Locard, 1882
+ Unio antimoquinianus Locard, 1889
+ Unio passavanti Bourguignat in Locard, 1889
Unio consentaneus var. oxyrhynchus Brevière = Unio crassus
Unio crassus Philippson = Unio crassus + Unio elongatulus + Unio nanus
+ Unio corrosus Villa, 1841
+ Unio crassatellus Bourguignat in Locard, 1882
+ Unio socardianus Bourguignat in Locard, 1882
+ Unio locardianus Bourguignat in Locard, 1882
+ Unio subrobustus Servain, 1885
+ Unio batavus Miller, 1873
+ Unio hamburgiensis Servain, 1888
Unio crassus var. piscinalis Ziegler = Unio nanus + Unio crassus
+ Unio fusculus Rossmässler, 1836
+ Unio courtillieri Hattemann, 1859
+ Unio pruinosus J.F. Schmidt, 1840
+ Unio reniformis Rossmässler, 1836
+ Unio carcasinus Sourbieu, 1887
+ Unio ovatus Studer, 1820
+ Unio catalaunicus Bourguignat in Locard, 1889
Unio limosus Nilsson = Unio pictorum
+ Unio jousseaumei Bourguignat in Locard, 1882
Unio mancus de Lamarck = Unio mancus + Unio crassus + Unio pictorum + Unio nanus
+ Unio manca ‘Lamarck’ of Authors
+ Unio mancus var. sabaudina Locard, 1882
+ Unio bourgeticus Bourguignat in Locard, 1882
+ Unio pilloti Bourguignat in Locard, 1882
+ Unio rayi Bourguignat in Locard, 1882
+ Unio dubisianus Locard, 1882
+ Unio zoasthenus Locard, 1889
+ Unio manculus Locard, 1889
+ Unio lagnisicus Bourguignat in Locard, 1882
+ Unio suborbicularis Drouët, 1888
Unio mancus var. amnicus Ziegler = Unio nanus + Unio crassus
+ Unio batavus var. minimus de Joannis, 1859
+ Unio subtilis Drouët, 1879
+ Unio melas Coutagne in Locard, 1882
+ Unio batavus var. nanus de Joannis, 1859
+ Unio dubisiopsis Locard, 1882
+ Unio ampulodon Locard, 1889
+ Unio carantoni Bourguignat in Locard, 1882
+ Unio minutulus Servain, 1888
+ Unio minutus Locard, 1882
+ Unio jurianus Locard, 1889
2 more...
Unio mancus var. amnicus form riparius Pfeiffer = Unio nanus + Unio crassus
+ Unio berthelini Bourguignat in Locard, 1882
+ Unio aturicus Locard, 1889
Unio mancus var. elongatulus Mühlfeld = Unio elongatulus + Unio mancus + Unio crassus + Unio nanus
+ Unio riciacensis Bourguignat in Locard, 1882
+ Unio orthus Coutagne in Locard, 1882
+ Unio orthellus Bourguignat in Locard, 1882
+ Unio lepidulus Drouët, 1898
+ Unio andeliacus Bourguignat in Locard, 1889
+ Unio valliericus Bourguignat in Locard, 1889
+ Unio nubilus Locard, 1889
+ Unio lemotheuxi Bourguignat in Locard, 1889
Unio mancus var. macrorhynchus Bourguignat = Unio nanus + Unio crassus
+ Unio ater Bourguignat, 1864
+ Unio neocomensis Drouët, 1881
+ Unio necomensis ‘Drouët’ of Authors
+ Unio ostiorum Servain, 1885
+ Unio sanderi Bourguignat, 1862
+ Unio sandriopsis Servain, 1885
Unio mancus var. nanus de Lamarck = Unio nanus + Unio crassus
+ Unio nanus Dupuy, 1852
+ Unio nanus Drouët, 1857
Unio michaudi Des Moulins = Unio pictorum
+ Unio michaudiana Des Moulins, 1833
Unio requieni Michaud = Unio mancus + Unio delphinus + Unio pictorum + Unio elongatulus
+ Unio albanorum Locard, 1889
+ Unio aramonensis Locard, 1889
+ Unio ararisianus Bourguignat in Locard, 1889
+ Unio bramicus Baichère, 1890
+ Unio caficianus Bourguignat, 1883
+ Unio campylus Bourguignat in Locard, 1889
+ Unio condatinus Bourguignat in Locard, 1882
+ Unio dolfusianus Bourguignat in Locard, 1882
+ Unio fascellinus Bourguignat in Locard, 1882
+ Unio frayssianus Bourguignat in Locard, 1889
32 more...
Unio requieni var. danielis Gassies = Unio pictorum
+ Unio corbini Bourguignat in Locard, 1882
+ Unio pictorum var. longirostris de Joannis, 1859
+ Unio requienii Dupuy, 1878
+ Unio fagoti Bourguignat in Locard, 1882
Unio requieni var. platyrhynchoideus [sic] Dupuy = Unio pictorum
+ Unio platyrhynchoideus ‘Dupuy’ of Authors
+ Unio renei Locard, 1882
Unio requieni var. pornae Bourguignat = Unio mancus + Unio pictorum
+ Unio pictorum de Joannis, 1859
+ Unio cancrorum Bourguignat in Locard, 1882
+ Unio pictorum Dupuy, 1852
+ Unio cavarellus Servain, 1887
+ Unio charpyi Drouët, 1888
+ Unio cristulatus Drouët, 1889
+ Unio falsus Bourguignat in Locard, 1882
+ Unio plebeius Drouët, 1888
+ Unio lesumicus Bourguignat in Locard, 1889
+ Unio rostratus Michaud, 1831
8 more...
Unio requieni var. rousi Dupuy = Unio mancus + Unio pictorum
+ Unio ronsii Dupuy, 1852
+ Unio requieni Gassies, 1849
Unio triffoiricus Bourguignat = Unio mancus + Unio pictorum
+ Unio gestroianus Bourguignat in Locard, 1882
+ Unio valens Drouët, 1897
Unio tumidus Philippson = Unio tumidus + Unio pictorum
+ Unio tumidus Rossmässler, 1836
+ Mya ovata Donovan, 1802
+ Unio tumidulus Locard, 1889
+ Unio aldemaricus Locard, 1889
+ Unio alpecanus Bourguignat in Locard, 1882
+ Unio bardus Bourguignat, in sched. Servain, 1881
+ Unio berilloni Locard, 1882
+ Unio catinulus Locard, 1893
+ Unio conus Spengler, 1793
+ Unio edyus Bourguignat in Locard, 1882
16 more...
Unio tumidus var. maximus Mörch = Unio tumidus + Unio mancus
+ Unio rostratus var. amplus Drouët, 1898
+ Unio malafossianus Locard, 1882
+ Unio requienii var. permaxima Dupuy, 1877
+ Unio aegericus Locard, 1889
Unio tumidus var. rostratus de Lamarck = Unio pictorum
+ Unio pictorum Rossmässler, 1837
+ Unio longirostris Rossmässler, 1836
+ Unio lugdunicus Bourguignat in Locard, 1889
+ Unio macropisthus Locard, 1889
+ Unio perroudi Locard, 1889
+ Unio proechistus Bourguignat, 1870
+ Unio rostratus var. falcatus Drouët, 1898
+ Unio rostratellus Bourguignat in Locard, 1882
+ Unio ruginosus Drouët, 1898
+ Unio siliquiformis Locard, 1889
Unio turtoni Payraudeau = Unio mancus
+ Unio turtoni Dupuy, 1850
+ Unio berenguieri Bourguignat in Locard, 1882
+ Unio forojuliensis Bourguignat in Locard, 1882
+ Unio prolatus Locard, 1893
+ Unio linguiformis Locard, 1893
Unio turtoni var. aleroni Companyo et Massot = Unio mancus
Unio turtoni var. philippei [sic] Dupuy = Unio pictorum
+ Unio philippei ‘Dupuy’ of Authors
+ Unio hauterivianus Bourguignat in Locard, 1882
+ Unio brindosopsis Locard, 1889
Unio villae Stabile (Villa) = Unio elongatulus + Unio pictorum + Unio mancus
+ Unio veillanensis Locard, 1882
+ Unio brebissoni Locard, 1889
+ Unio pictorum l’Hôpital, 1859
+ Unio hopitali Locard, 1889
+ Unio requieni var. minima l’Hôpital, 1859
+ Unio amblyus Locard, 1889
+ Unio baicheri Baichère, 1890
Unio villae var. brindosianus Folin et Bérillon = Unio mancus
+ Unio lusitanus Drouët, 1879
+ Unio bayonnensis Folin & Bérillon, 1877
+ Unio moreleti Folin & Bérillon, 1874
+ Unio mongazonae Servain, 1887
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