Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae

tribe ACUTICOSTINI Starobogatov, 1967: cladomics

1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 2 (3 total)
number of subtaxa (rank: genus) = 1
analyzed in 63 cladograms in 38 publications
maximum number of terminals in an analysis = 2 (mean = 1.11, median = 1)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 7
maximum number of included terminals = 2, (mean = 2 median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Acuticostini cladogram distances
PCoA Ordination

NJ Tree of Acuticostini cladogram distances
NJ Tree

Cladograms

Aunins et al., 2018 [Publ 3863], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 25 4 **
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
0.909 1 1 0 1 33 13 4 0.0
Cladomics cladogram ANODONTINI Acuticosta Nodularia Tchangsinaia LANCEOLARIINI Schistodesmus LEPIDODESMINI

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.818 1 1 0 1 33 13 4 0.0
Cladomics cladogram Strophitus Schistodesmus LEPIDODESMINI Acuticosta CRISTARIINA LANCEOLARIINI Nodularia Tchangsinaia Utterbackia Utterbackiana Alasmidonta Lasmigona

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 32 20 4 0.0
Cladomics cladogram ANODONTINI Tchangsinaia Nodularia Schistodesmus Acuticosta Inversiunio LEPIDODESMINI LANCEOLARIINI

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
0.931 1 1 0 1 52 32 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Huang et al., 2019 [Publ 3843], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 45 31 4 0.0
Cladomics cladogram Anemina Acuticosta Sinanodonta ACULAMPROTULINA Cristaria LANCEOLARIINI LEPIDODESMINI OXYNAIINA

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.912 1 1 0 1 200 47 4 0.0
Cladomics cladogram Acuticosta LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
0.926 1 1 0 1 55 25 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI Acuticosta ANODONTINI

Huang et al., 2019 [Publ 3843], fig. S1

cst iT tT bi riS oT roT roS iD
0.900 2 2 5 1 45 31 4 0.0
Cladomics cladogram Anemina Sinanodonta Acuticosta LEPIDODESMINI Cristaria LANCEOLARIINI UNIONINI

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.912 1 1 0 1 198 46 4 0.0
Cladomics cladogram Acuticosta UNIONINI LEPIDODESMINI LANCEOLARIINI ANODONTINI

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.765 1 1 0 1 200 47 4 0.0
Cladomics cladogram Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona LEPIDODESMINI Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus UNIONINA ACULAMPROTULINA LANCEOLARIINI Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 25 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.949 1 1 0 1 121 70 4 0.0
Cladomics cladogram Acuticosta LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 53 27 4 0.0
Cladomics cladogram LEPIDODESMINI LANCEOLARIINI UNIONINI Acuticosta ANODONTINI

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 25 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.565 1 1 0 1 33 21 4 0.0
Cladomics cladogram Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula LEPIDODESMINI ACULAMPROTULINA Acuticosta Cuneopsis Nodularia Schistodesmus Inversiunio

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
0.931 1 1 0 1 90 25 4 0.0
Cladomics cladogram Acuticosta LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.935 1 1 0 1 72 32 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.935 1 1 0 1 72 32 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.935 1 1 0 1 67 31 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.897 1 1 0 1 90 25 4 0.0
Cladomics cladogram Acuticosta UNIONINI LANCEOLARIINI LEPIDODESMINI ANODONTINI

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 37 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI Acuticosta ANODONTINI

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 34 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 36 15 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI Acuticosta ANODONTINI

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
0.960 1 1 0 1 38 16 4 0.0
Cladomics cladogram ANODONTINI Acuticosta LANCEOLARIINI LEPIDODESMINI ACULAMPROTULINA UNIONINA OXYNAIINA

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
0.966 1 1 0 1 55 32 4 0.0
Cladomics cladogram UNIONINI LEPIDODESMINI LANCEOLARIINI Acuticosta ANODONTINI

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 16 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.861 1 1 0 1 40 31 4 0.0
Cladomics cladogram Anemina Amuranodonta Pletholophus Sinanodonta Beringiana Lanceolaria gladiola Acuticosta LEPIDODESMINI Lanceolaria oxyrhyncha Cristaria UNIONINI ALASMIDONTINA ANODONTINA

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
0.972 1 1 0 1 40 31 4 0.0
Cladomics cladogram Acuticosta LEPIDODESMINI UNIONINI LANCEOLARIINI ANODONTINI

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 45 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.914 1 1 0 1 94 45 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.971 1 1 0 1 90 45 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.971 1 1 0 1 90 45 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 27 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI Acuticosta ANODONTINI

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
0.917 1 1 0 1 21 16 4 0.0
Cladomics cladogram Acuticosta CRISTARIINA ANODONTINA LANCEOLARIINI ACULAMPROTULINA LEPIDODESMINI UNIONINA MIDDENDORFFINAIINA OXYNAIINA

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
0.958 1 1 0 1 21 16 4 0.0
Cladomics cladogram ANODONTINA Acuticosta CRISTARIINA LEPIDODESMINI LANCEOLARIINI UNIONINI

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 26 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI Acuticosta ANODONTINI

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.895 1 1 0 1 43 34 4 0.0
Cladomics cladogram Pletholophus Sinanodonta Beringiana Cristaria ALASMIDONTINA ANODONTINA LANCEOLARIINI LEPIDODESMINI Acuticosta UNIONINI

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.868 1 1 0 1 43 34 4 0.0
Cladomics cladogram Anemina Amuranodonta Pletholophus Sinanodonta Beringiana ALASMIDONTINA ANODONTINA Lanceolaria gladiola Lanceolaria oxyrhyncha Cristaria Acuticosta LEPIDODESMINI UNIONINI

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 41 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 42 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI Acuticosta ANODONTINI

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
0.973 1 1 0 1 40 31 4 0.0
Cladomics cladogram GLOBUNIONINA ACULAMPROTULINA UNIONINA OXYNAIINA Acuticosta LEPIDODESMINI LANCEOLARIINI ANODONTINI

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
0.946 1 1 0 1 40 31 4 0.0
Cladomics cladogram UNIONINI Acuticosta ALASMIDONTINA ANODONTINA LEPIDODESMINI Cristaria LANCEOLARIINI Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.947 1 1 0 1 139 67 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI LANCEOLARIINI Acuticosta ANODONTINI

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
0.977 2 2 0 1 93 63 4 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.818 1 2 0 1 632 155 4 0.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Beringiana Pletholophus Mimo Acudonta Sinanodonta Cristaria LEPIDODESMINI LANCEOLARIINI Pseudoacuticosta retiaria 2 Acuticosta chinensis 2 Acuticosta chinensis 1 Pseudoacuticosta retiaria 1 Globunio ACULAMPROTULINA MIDDENDORFFINAIINA UNIONINA OXYNAIINA Pseudodontoideus Lasmigona Pterosyna Arcidens ANODONTINA Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus

Zhang et al., 2004 [Publ 2735], fig. 1

cst iT tT bi riS oT roT roS iD
0.938 1 1 0 1 13 11 3 0.0
Cladomics cladogram ANODONTINI UNIONINI Acuticosta LANCEOLARIINI

Zhou et al., 2007 [Publ 2658], fig. 1

cst iT tT bi riS oT roT roS iD
0.812 2 2 0 1 19 11 3 0.0
Cladomics cladogram ACULAMPROTULINA Tchangsinaia Nodularia Schistodesmus LANCEOLARIINI Acuticosta

Ouyang et al., 2011 [Publ 3408], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 12 8 3 0.0
Cladomics cladogram ANODONTINI OXYNAIINA Lanceolaria grayii Acuticosta Lanceolaria gladiola ACULAMPROTULINA

Ouyang et al., 2015 [Publ 4493], fig. 1

cst iT tT bi riS oT roT roS iD
0.882 2 2 0 1 29 14 3 0.0
Cladomics cladogram UNIONINI LANCEOLARIINI Acuticosta ANODONTINI

Ouyang et al., 2015 [Publ 4493], fig. 2

cst iT tT bi riS oT roT roS iD
0.882 2 2 4 1 29 14 3 0.0
Cladomics cladogram LANCEOLARIINI ACULAMPROTULINA Acuticosta ANODONTINI OXYNAIINA

Liu et al., 2017 [Publ 3939], fig. 2

cst iT tT bi riS oT roT roS iD
0.933 1 1 0 1 14 13 3 0.0
Cladomics cladogram Acuticosta ANODONTINI GONIDEINI

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 16 3 0.0
Cladomics cladogram UNIONINI ANODONTINI Acuticosta LANCEOLARIINI

Kim et al., 2021 [Publ 4197], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 17 16 3 0.0
Cladomics cladogram ACULAMPROTULINA Acuticosta LANCEOLARIINI UNIONINA OXYNAIINA

Wu et al., 2023 [Publ 4296], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 20 3 0.0
Cladomics cladogram Acuticosta LANCEOLARIINI ANODONTINI UNIONINI

Liu et al., 2023 [Publ 4310], fig. 4

cst iT tT bi riS oT roT roS iD
0.917 1 1 0 1 23 21 3 0.0
Cladomics cladogram Acuticosta Lanceolaria lanceolata Lanceolaria grayii Lanceolaria gladiola ANODONTINI UNIONINI

Liu et al., 2023 [Publ 4310], fig. 5

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 23 21 3 0.0
Cladomics cladogram Acuticosta Lanceolaria grayii Lanceolaria gladiola Lanceolaria lanceolata ANODONTINI UNIONINI

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
0.974 2 2 0 1 62 60 3 0.0
Cladomics cladogram LEPIDODESMINI UNIONINI Acuticosta ANODONTINI

Wu et al., 2026 [Publ 4811], fig. 3

cst iT tT bi riS oT roT roS iD
0.933 2 2 0 1 15 8 3 0.0
Cladomics cladogram UNIONINA LANCEOLARIINI GLOBUNIONINA Acuticosta ANODONTINI

Zhou et al., 2007 [Publ 2658], fig. 2

cst iT tT bi riS oT roT roS iD
0.692 1 1 0 1 17 7 2 0.0
Cladomics cladogram ACULAMPROTULINA Acuticosta OXYNAIINA

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 10 2 0.0
Cladomics cladogram ANODONTINI Acuticosta UNIONINI

Fukata & Iigo, 2019 [Publ 4108], fig. 1

cst iT tT bi riS oT roT roS iD
0.938 1 1 0 1 16 16 2 0.0
Cladomics cladogram UNIONINA OXYNAIINA ACULAMPROTULINA Acuticosta LANCEOLARIINI

Wu et al., 2021 [Publ 4215], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 23 3 1 0.0
Cladomics cladogram Acuticosta UNIONINI