Bivalvia | Unionida | Etherioidea | Mycetopodidae

subfamily ANODONTITINAE Modell, 1942: cladomics

3 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 29 (29 total)
number of subtaxa (rank: genus) = 3
analyzed in 52 cladograms in 28 publications
maximum number of terminals in an analysis = 8 (mean = 1.67, median = 1)
maximum number of subtaxa (genus) = 3 (mean = 1.17, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 20
maximum number of included terminals = 8, (mean = 2.75 median = 2)
maximum number of subtaxa (genus) = 3 (mean = 1.45, median = 1)
proportion of cladograms that support basic monophyly = 0.5

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 8
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 8 (mean = 3.5, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.13, median = 2)
proportion of cladograms that support rigorous monophyly = 0

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Anodontitinae cladogram distances
PCoA Ordination

NJ Tree of Anodontitinae cladogram distances
NJ Tree

Cladograms

Prié et al., 2026 [Publ 4844], fig. 3

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
0.700 8 12 9 3 7 4 2 ** 
  +----------------- Etheria                
  |                                         
  |  +-------------- Anodontites_trapesialis **
--+  |                                      
  |  |     +-------- Anodontites_guanarensis **
  |  |     |                                
  +--+  +--+-------- Fossula                
     |  |  |                                
     |  |  |  +----- Anodontites_elongata **   
     |  |  +--+                             
     |  |     |  +-- Pseudanodontites **       
     +--+     +--+                          
        |        +-- Anodontites_trigona **    
        |                                   
        |  +-------- Mycetopodinae          
        |  |                                
        +--+     +-- Anodontites_cryptica **   
           |  +--+                          
           |  |  +-- Anodontites_crispata **   
           +--+                             
              |  +-- Lamproscapha **           
              +--+                          
                 +-- Tamsiella

Huang et al., 2019 [Publ 3843], fig. 2

cstiTtTbiriSoTroTroSiD
0.750 2 10 0 2 199 2 2 14.0 
                          +-- Anodontites **             
                 +--------+                           
                 |        +-- Iridininae              
                 |                                    
  +--------------+        +-- Etheria                 
  |              |  +-----+                           
  |              |  |     +-- Aspathariinae           
  |              +--+                                 
  |                 |  +----- Mycetopodinae           
  |                 +--+                              
  |                    |  +-- Monocondylaeinae        
--+                    +--+                           
  |                       +-- Lamproscapha **            
  |                                                   
  |                    +----- Gibbosula_rochechouartii
  |  +-----------------+                              
  |  |                 |  +-- Gibbosula_laosensis     
  |  |                 +--+                           
  +--+                    +-- Margaritiferinae        
     |                                                
     |  +-------------------- Parreysiinae            
     +--+                                             
        |  +----------------- Unioninae               
        +--+                                          
           |  +-------------- Rectidentini            
           +--+                                       
              |  +----------- Ambleminae              
              +--+                                    
                 |  +-------- Chamberlainiini         
                 +--+                                 
                    |  +----- Lamprotulini            
                    +--+                              
                       |  +-- Gonideini               
                       +--+                           
                          +-- Pseudodontini

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
0.750 2 10 0 2 197 2 2 14.0 
                          +-- Anodontites **             
                 +--------+                           
                 |        +-- Iridininae              
                 |                                    
  +--------------+        +-- Etheria                 
  |              |  +-----+                           
  |              |  |     +-- Aspathariinae           
  |              +--+                                 
  |                 |  +----- Mycetopodinae           
  |                 +--+                              
  |                    |  +-- Monocondylaeinae        
--+                    +--+                           
  |                       +-- Lamproscapha **            
  |                                                   
  |                    +----- Gibbosula_rochechouartii
  |  +-----------------+                              
  |  |                 |  +-- Gibbosula_laosensis     
  |  |                 +--+                           
  +--+                    +-- Margaritiferinae        
     |                                                
     |  +-------------------- Parreysiinae            
     +--+                                             
        |  +----------------- Pseudodontini           
        +--+                                          
           |  +-------------- Rectidentini            
           +--+                                       
              |  +----------- Unioninae               
              +--+                                    
                 |  +-------- Chamberlainiini         
                 +--+                                 
                    |  +----- Ambleminae              
                    +--+                              
                       |  +-- Lamprotulini            
                       +--+                           
                          +-- Gonideini

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.750 2 10 0 2 199 2 2 14.0 
                 +-------- Etheria                 
                 |                                 
                 +-------- Aspathariinae           
                 |                                 
  +--------------+     +-- Anodontites **             
  |              +-----+                           
  |              |     +-- Iridininae              
  |              |                                 
  |              |  +----- Mycetopodinae           
  |              +--+                              
  |                 |  +-- Monocondylaeinae        
  |                 +--+                           
  |                    +-- Lamproscapha **            
  |                                                
  |           +----------- Echyridella             
  |           |                                    
  |           +----------- Westralunio             
  |           |                                    
--+           +----------- Hyridellini             
  |           |                                    
  |           |     +----- Rhipidodontini          
  |           +-----+                              
  |  +--------+     |  +-- Castaliini              
  |  |        |     +--+                           
  |  |        |        +-- Hyriini                 
  |  |        |                                    
  |  |        |     +----- Velesunio               
  |  |        |  +--+                              
  |  |        |  |  |  +-- Alathyria_jacksoni      
  |  |        |  |  +--+                           
  |  |        +--+     +-- Alathyria_profuga       
  |  |           |                                 
  |  |           |  +----- Alathyria_pertexta      
  +--+           +--+                              
     |              |  +-- Lortiella               
     |              +--+                           
     |                 +-- Microdontia             
     |                                             
     |              +----- Gibbosula_rochechouartii
     |  +-----------+                              
     |  |           |  +-- Gibbosula_laosensis     
     |  |           +--+                           
     |  |              +-- Margaritiferinae        
     |  |                                          
     |  |  +-------------- Parreysiinae            
     +--+  |                                       
        |  |  +----------- Chamberlainia           
        |  |  |                                    
        |  |  +----------- Megalonaias             
        |  |  |                                    
        |  |  +----------- Uniomerus               
        |  |  |                                    
        +--+  +----------- Quadrulina              
           |  |                                    
           |  +----------- Sinohyriopsis           
           |  |                                    
           |  +----------- Lamprotulini            
           |  |                                    
           |  +----------- Gonideini               
           |  |                                    
           +--+----------- Pseudodontini           
              |                                    
              +----------- Rectidentini            
              |                                    
              +----------- Unioninae               
              |                                    
              |  +-------- Amblema                 
              |  |                                 
              |  +-------- Toxolasma               
              |  |                                 
              |  +-------- Pleurobemini            
              |  |                                 
              |  |     +-- Obliquaria              
              |  +-----+                           
              |  |     +-- Plectomerus             
              +--+                                 
                 |  +----- Ellipsaria              
                 |  |                              
                 |  +----- Potamilus_leptodon      
                 |  |                              
                 |  +----- Medionidus              
                 |  |                              
                 |  +----- Potamilus_alatus        
                 |  |                              
                 |  +----- Ptychobranchus          
                 +--+                              
                    +----- Truncilla               
                    |                              
                    +----- Venustaconcha           
                    |                              
                    +----- Cambarunio              
                    |                              
                    +----- Leaunio                 
                    |                              
                    |  +-- Ortmanniana             
                    +--+                           
                       +-- Lampsilis

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cstiTtTbiriSoTroTroSiD
0.600 5 12 10 2 628 6 2 11.0 
  +----------------------- Etheria                
  |                                               
--+  +-------------------- Anodontites_trapesialis **
  |  |                                            
  +--+  +----------------- Mulleria               
     |  |                                         
     +--+  +-------------- Lamproscapha **           
        |  |                                      
        +--+           +-- Monocondylaea          
           |  +--------+                          
           |  |        +-- Tamsiella              
           +--+                                   
              |  +-------- Mycetopodinae          
              |  |                                
              +--+     +-- Anodontites_guanarensis **
                 |  +--+                          
                 |  |  +-- Fossula                
                 +--+                             
                    |  +-- Anodontites_trigona **    
                    +--+                          
                       +-- Anodontites_elongata **

Guerra et al., 2019 [Publ 4168], fig. 1

cstiTtTbiriSoTroTroSiD
0.571 3 5 3 2 38 2 1 9.0 
  +-------------- Aspathariinae          
--+                                      
  |  +----------- Anodontites_trapesialis **
  +--+                                   
     |  +-------- Monocondylaea          
     +--+                                
        |  +----- Lamproscapha **           
        +--+                             
           |  +-- Anodontites_elongata **   
           +--+                          
              +-- Fossula

Guerra et al., 2019 [Publ 4168], fig. 2

cstiTtTbiriSoTroTroSiD
0.571 3 5 3 2 38 2 1 9.0 
  +-------------- Iridininae             
--+                                      
  |  +----------- Anodontites_trapesialis **
  +--+                                   
     |  +-------- Monocondylaea          
     +--+                                
        |  +----- Lamproscapha **           
        +--+                             
           |  +-- Anodontites_elongata **   
           +--+                          
              +-- Fossula

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cstiTtTbiriSoTroTroSiD
0.571 3 5 3 2 137 2 1 9.0 
  +-------------- Aspathariinae          
--+                                      
  |  +----------- Anodontites_trapesialis **
  +--+                                   
     |  +-------- Monocondylaea          
     +--+                                
        |  +----- Lamproscapha **           
        +--+                             
           |  +-- Anodontites_elongata **   
           +--+                          
              +-- Fossula

Graf, 2000 [Publ 2647], fig. 1

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 16 1 1 -- 
  +----- Anodontites **  
  |                   
--+----- Mycetopodinae
  |                   
  |  +-- Etheria      
  +--+                
     +-- Mulleria

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 32 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cstiTtTbiriSoTroTroSiD
0.600 2 3 0 1 29 1 1 11.0 
  +----- Iridininae             
  |                             
--+  +-- Anodontites_guanarensis **
  |  |                          
  +--+-- Anodontites_trigona **    
     |                          
     +-- Monocondylaeinae

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 29 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Hoeh et al., 2002 [Publ 2653], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 29 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 29 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 29 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Graf & Cummings, 2006 [Publ 1968], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 45 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Walker et al., 2006 [Publ 2666], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 34 2 1 -- 
  +-- Anodontites **     
--+                   
  +-- Monocondylaeinae

Hoeh et al., 2009 [Publ 2716], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 45 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Hoeh et al., 2009 [Publ 2716], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 44 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Hoeh et al., 2009 [Publ 2716], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 1 44 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Combosch et al., 2017 [Publ 3806], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Combosch et al., 2017 [Publ 3806], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Huang et al., 2018 [Publ 3828], fig. 3_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 28 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 27 1 1 -- 
  +-- Lamproscapha **    
--+                   
  +-- Monocondylaeinae

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 27 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 121 1 1 -- 
  +-- Lamproscapha ** 
--+                
  +-- Mycetopodinae

Wu et al., 2020 [Publ 4121], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Wu et al., 2020 [Publ 4121], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Sano et al., 2020 [Publ 4138], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 68 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Sano et al., 2020 [Publ 4138], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 68 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Sano et al., 2020 [Publ 4138], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 68 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Wu et al., 2021 [Publ 4366], fig. 2_I

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Wu et al., 2021 [Publ 4366], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Chen et al., 2025 [Publ 4455], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 25 1 1 -- 
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Martínez et al., 2026 [Publ 4820], fig. 2

cstiTtTbiriSoTroTroSiD
0.600 5 7 5 1 11 2 1 11.0 
  +-------------- Ambleminae             
  |                                      
--+  +----------- Anodontites_trapesialis **
  |  |                                   
  +--+        +-- Anodontites_trigona **    
     |  +-----+                          
     |  |     +-- Anodontites_elongata **   
     +--+                                
        |  +----- Anodontites_guanarensis **
        +--+                             
           |  +-- Anodontites_tortilis **   
           +--+                          
              +-- Mycetopodinae

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 39 0 0 -- 
  +-- Lamproscapha ** 
--+                
  +-- Aspathariinae

Guerra et al., 2017 [Publ 3772], fig. 2a

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 24 0 0 -- 
  +-- Iridininae 
--+              
  +-- Anodontites **

Guerra et al., 2017 [Publ 3772], fig. 2b

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 24 0 0 -- 
  +-- Iridininae 
--+              
  +-- Anodontites **

Huang et al., 2018 [Publ 3828], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 41 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Huang et al., 2019 [Publ 3843], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 55 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Wu et al., 2020 [Publ 4121], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Wu et al., 2021 [Publ 4366], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Wu et al., 2021 [Publ 4366], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Wu et al., 2021 [Publ 4366], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 67 0 0 -- 
  +-- Iridininae 
--+              
  +-- Anodontites **

Wu et al., 2022 [Publ 4238], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 0 0 -- 
  +-- Iridininae 
--+              
  +-- Anodontites **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Dai et al., 2025 [Publ 4440], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 86 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Chen et al., 2025 [Publ 4448], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 84 0 0 -- 
  +-- Anodontites **
--+              
  +-- Iridininae

Prié et al., 2026 [Publ 4844], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 0 0 -- 
  +-- Anodontites **
--+              
  +-- Etheria