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Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Etherioidea | Mycetopodidae

subfamily ANODONTITINAE Modell, 1942: cladomics
     2 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 27 (27 total)
number of subtaxa (rank: genus) = 2
analyzed in 47 cladograms in 25 publications
maximum number of terminals in an analysis = 2 (mean = 1.34, median = 1)
maximum number of subtaxa (genus) = 2 (mean = 1.11, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 16
maximum number of included terminals = 2, (mean = 2 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.31, median = 1)
proportion of cladograms that support basic monophyly = 0.63

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 5
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 2 (mean = 2, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Huang et al., 2019 [Publ 3843], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
21002197220
           +-- Anodontites **     
  +--------+                   
  |        +-- Iridininae      
  |                            
--+        +-- Etheria         
  |  +-----+                   
  |  |     +-- Aspathariinae   
  +--+                         
     |  +----- Mycetopodinae   
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Lamproscapha **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
250239319
  +----------- Anodontites **  
--+                         
  |  +-------- Monocondylaea
  +--+                      
     |  +----- Lamproscapha ** 
     +--+                   
        |  +-- Tamsiella    
        +--+                
           +-- Fossula

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
22012911--
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
22014511--
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
21002197224
  +-------- Etheria         
  |                         
  +-------- Aspathariinae   
  |                         
--+     +-- Anodontites **     
  +-----+                   
  |     +-- Iridininae      
  |                         
  |  +----- Mycetopodinae   
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Lamproscapha **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
22012911--
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
22013211--
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
2301291111
  +-- Anodontites_guanarensis **
  |                          
--+-- Anodontites_trigona **    
  |                          
  +-- Monocondylaeinae

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
22013421--
  +-- Anodontites **     
--+                   
  +-- Monocondylaeinae

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
250239319
  +----------- Anodontites **  
--+                         
  |  +-------- Monocondylaea
  +--+                      
     |  +----- Lamproscapha ** 
     +--+                   
        |  +-- Tamsiella    
        +--+                
           +-- Fossula

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
22014411--
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
22012911--
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
22014411--
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
22014511--
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
22012911--
  +-- Monocondylaeinae
--+                   
  +-- Anodontites **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
21002195220
           +-- Anodontites **     
  +--------+                   
  |        +-- Iridininae      
  |                            
--+        +-- Etheria         
  |  +-----+                   
  |  |     +-- Aspathariinae   
  +--+                         
     |  +----- Mycetopodinae   
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Lamproscapha **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
11018400--
-- Anodontites_trapesialis **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
11013311--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
11012711--
  +-- Lamproscapha **    
--+                   
  +-- Monocondylaeinae

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
11011611--
-- Anodontites **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
11011411--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
11012811--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019011--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
11016711--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
11012711--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017200--
-- Anodontites_trapesialis **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012400--
-- Anodontites_trapesialis **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
11011411--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017200--
-- Anodontites_trapesialis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019400--
-- Anodontites_trapesialis **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019000--
-- Anodontites_trapesialis **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018700--
-- Anodontites_trapesialis **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
11012511--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014800--
-- Anodontites_trapesialis **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013900--
-- Lamproscapha_ensiformis **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019011--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
11016711--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
11014100--
-- Anodontites_trapesialis **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
11015500--
-- Anodontites_trapesialis **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017200--
-- Anodontites_trapesialis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112011--
  +-- Lamproscapha ** 
--+                
  +-- Mycetopodinae

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
11016711--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019400--
-- Anodontites_trapesialis **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012400--
-- Anodontites_trapesialis **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019000--
-- Anodontites_trapesialis **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
11016700--
-- Anodontites_trapesialis **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
11013311--
  +-- Monocondylaeinae
--+                   
  +-- Lamproscapha **
 
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