Bivalvia | Unionida | Hyrioidea | Hyriidae | Hyriinae

tribe CASTALIINI Morretes, 1949: cladomics

3 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 17 (18 total)
number of subtaxa (rank: genus) = 3
analyzed in 42 cladograms in 27 publications
maximum number of terminals in an analysis = 5 (mean = 1.45, median = 1)
maximum number of subtaxa (genus) = 3 (mean = 1.29, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 7
maximum number of included terminals = 5, (mean = 3.71 median = 4)
maximum number of subtaxa (genus) = 3 (mean = 2.71, median = 3)
proportion of cladograms that support basic monophyly = 1

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 6
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 5 (mean = 3.83, median = 4)
maximum number of subtaxa (genus) = 3 (mean = 2.83, median = 3)
proportion of cladograms that support rigorous monophyly = 1


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Castaliini cladogram distances
PCoA Ordination

NJ Tree of Castaliini cladogram distances
NJ Tree


Cladograms

Prié et al., 2026 [Publ 4844], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.812 4 4 2 3 19 11 4 **
Cladomics cladogram Diplodon Guyanhyrida Iridea HYRIINI Castalia cordata Callonaia Castalia ambigua Castaliella

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

cst iT tT bi riS oT roT roS iD
0.667 4 4 5 3 17 10 4 6.0
Cladomics cladogram RHIPIDODONTINI HYRIINI Castalia stevensi Callonaia Castalia ambigua 1 Castalia ambigua 4 Castalia ambigua 2 Castaliella Castalia ambigua 3

Santos-Neto et al., 2016 [Publ 3739], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 4 4 5 3 17 10 4 6.0
Cladomics cladogram RHIPIDODONTINI HYRIINI Castalia ambigua 4 Callonaia Castalia ambigua 3 Castalia stevensi Castalia ambigua 2 Castaliella Castalia ambigua 1

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.667 5 5 6 3 628 14 4 7.0
Cladomics cladogram RHIPIDODONTINI HYRIINI Castalia ambigua 4 Callonaia Castalia stevensi Castalia ambigua 3 Castalia ambigua 1 Castaliella Castalia ambigua 2 Castalia

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.900 3 3 2 3 4 4 2 3.0
Cladomics cladogram RHIPIDODONTINI HYRIINI Callonaia Castalia ambigua 2 Castalia ambigua 1 Castaliella

Santos-Neto et al., 2018 [Publ 3888], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 2 2 1 3.0
Cladomics cladogram HYRIINI Callonaia Castalia

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 2 0 0 0 3.0
Cladomics cladogram Callonaia Castalia

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 200 6 4 --
Cladomics cladogram Echyridella HYRIDELLINI RHIPIDODONTINI Castalia HYRIINI

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 198 6 4 --
Cladomics cladogram Echyridella HYRIDELLINI RHIPIDODONTINI Castalia HYRIINI

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 200 6 4 --
Cladomics cladogram RHIPIDODONTINI Castalia HYRIINI

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 10 4 3 --
Cladomics cladogram RHIPIDODONTINI Castalia Hyridella australis 2 Hyridella australis 1 Echyridella Hyridella depressa

Baker et al., 2003 [Publ 2690], fig. 5

cst iT tT bi riS oT roT roS iD
0.625 1 1 0 1 7 4 3 --
Cladomics cladogram Echyridella Castalia

Baker et al., 2004 [Publ 2685], fig. 2-3

cst iT tT bi riS oT roT roS iD
0.625 1 1 0 1 8 4 3 --
Cladomics cladogram Echyridella Castalia

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 5 3 --
Cladomics cladogram Echyridella HYRIDELLINI Castalia RHIPIDODONTINI

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.625 1 1 0 1 35 4 3 --
Cladomics cladogram Castalia RHIPIDODONTINI Hyridella depressa Hyridella australis Echyridella Velesunio ambiguus Velesunio 3 Velesunio 2 Alathyria Velesunio 1 Velesunio angasi Lortiella

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 5 3 --
Cladomics cladogram Echyridella HYRIDELLINI Castalia RHIPIDODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 5 3 --
Cladomics cladogram Echyridella HYRIDELLINI Castalia RHIPIDODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 5 3 --
Cladomics cladogram Echyridella HYRIDELLINI Castalia RHIPIDODONTINI

Graf et al., 2015 [Publ 3619], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 1 1 0 1 19 6 3 --
Cladomics cladogram HYRIDELLINI RHIPIDODONTINI HYRIINI Castalia

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 40 3 3 --
Cladomics cladogram RHIPIDODONTINI Castalia HYRIINI

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 3 3 --
Cladomics cladogram Echyridella RHIPIDODONTINI Castalia HYRIINI

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 139 3 3 --
Cladomics cladogram Echyridella RHIPIDODONTINI Castalia HYRIINI

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 16 2 2 --
Cladomics cladogram Castalia RHIPIDODONTINI HYRIDELLINI Velesunio

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 33 2 2 --
Cladomics cladogram Castalia RHIPIDODONTINI

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 30 2 2 --
Cladomics cladogram Castalia Velesunio

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 30 2 2 --
Cladomics cladogram Castalia RHIPIDODONTINI

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 2 2 --
Cladomics cladogram HYRIDELLINI RHIPIDODONTINI Castalia

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 30 2 2 --
Cladomics cladogram Castalia RHIPIDODONTINI

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 30 2 2 --
Cladomics cladogram Castalia Velesunio

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 30 2 2 --
Cladomics cladogram Castalia RHIPIDODONTINI

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 2 2 --
Cladomics cladogram HYRIDELLINI Castalia HYRIINI

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 2 2 --
Cladomics cladogram HYRIDELLINI Castalia HYRIINI

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 2 2 --
Cladomics cladogram HYRIDELLINI HYRIINI Castalia

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 2 2 --
Cladomics cladogram RHIPIDODONTINI HYRIINI Castalia

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 2 2 --
Cladomics cladogram RHIPIDODONTINI HYRIINI Castalia

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 138 2 2 --
Cladomics cladogram RHIPIDODONTINI Castalia HYRIINI

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 138 2 2 --
Cladomics cladogram RHIPIDODONTINI Castalia HYRIINI

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 2 2 --
Cladomics cladogram RHIPIDODONTINI HYRIINI Castalia

Lawrie et al., 2026 [Publ 4843], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 17 5 2 --
Cladomics cladogram Castalia HYRIINI

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Castalia RHIPIDODONTINI

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram RHIPIDODONTINI Castalia

Prié et al., 2026 [Publ 4844], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Castalia Margaritifera