MUSSELp

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Unionidae

Margaritiferidae

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Iridinidae


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Data last updated
Mon 1 Dec 2025

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae

tribe COELATURINI Modell, 1942: cladomics

10 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 37 (41 total)
number of subtaxa (rank: genus) = 10
analyzed in 51 cladograms in 33 publications
maximum number of terminals in an analysis = 27 (mean = 3.59, median = 2)
maximum number of subtaxa (genus) = 7 (mean = 2.06, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 27
maximum number of included terminals = 27, (mean = 5.89 median = 4)
maximum number of subtaxa (genus) = 7 (mean = 3, median = 3)
proportion of cladograms that support basic monophyly = 1

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 26
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 27 (mean = 5.69, median = 4)
maximum number of subtaxa (genus) = 7 (mean = 3.04, median = 3)
proportion of cladograms that support rigorous monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Coelaturini cladogram distances
PCoA Ordination

NJ Tree of Coelaturini cladogram distances
NJ Tree

Cladograms

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
27 27 16 7 9 9 2 0 
  +-------------------------------------- Leoparreysiini               
  |                                                                    
--+  +----------------------------------- Indochinellini               
  |  |                                                                 
  +--+  +-------------------------------- Grandidieria **                 
     |  |                                                              
     |  |                             +-- Pseudospatha **                 
     +--+  +--------------------------+                                
        |  |                          +-- Moncetia **                     
        |  |                                                           
        |  |  +-------------------------- Nitia **                        
        +--+  |                                                        
           |  |                       +-- Coelatura_aegyptiaca **         
           |  |                       |                                
           |  |                       +-- Coelatura_alluaudi **           
           +--+                       |                                
              |  +--------------------+-- Coelatura_bakeri **             
              |  |                    |                                
              |  |                    +-- Coelatura_hauttecoeuri **       
              |  |                    |                                
              |  |                    +-- Coelatura_stuhlmanni **         
              +--+                                                     
                 |                    +-- Coelatura_leopoldvillensis **   
                 |  +-----------------+                                
                 |  |                 +-- Coelatura_mesafricana **        
                 |  |                                                  
                 |  |                 +-- Coelatura_horei **              
                 +--+                 |                                
                    |  +--------------+-- Coelatura_ratidota **           
                    |  |              |                                
                    |  |              +-- Coelatura **                    
                    +--+                                               
                       |  +-------------- Coelatura_gabonensis **         
                       |  |                                            
                       |  +-------------- Coelatura_stanleyvillensis **   
                       +--+                                            
                          |  +----------- Coelatura_kunenensis **         
                          |  |                                         
                          |  |     +----- Prisodontopsis_aviculaeformis **
                          +--+  +--+                                   
                             |  |  |  +-- Coelatura_briarti **            
                             |  |  +--+                                
                             |  |     +-- Prisodontopsis_mweruensis **    
                             +--+                                      
                                |     +-- Coelatura_choziensis **         
                                |     |                                
                                |     +-- Coelatura_hypsiprymna **        
                                |     |                                
                                +-----+-- Coelatura_luapulaensis **       
                                      |                                
                                      +-- Coelatura_mossambicensis **     
                                      |                                
                                      +-- Nyassunio **

Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 4 2 3 58 58 4 23 
  +----------------- Lamellidentini      
  |                                      
--+              +-- Leoparreysiini      
  |  +-----------+                       
  |  |           +-- Parreysiini         
  +--+                                   
     |  +----------- Indochinellini      
     +--+                                
        |  +-------- Nitia **               
        +--+                             
           |  +----- Coelatura_aegyptiaca **
           +--+                          
              |  +-- Coelatura_kunenensis **
              +--+                       
                 +-- Nyassunio **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iTtTbiriSoTroTroSiD
27 27 16 7 9 9 2 5 
  +-------------------------------- Leoparreysiini               
  |                                                              
  |  +----------------------------- Indochinellini               
--+  |                                                           
  |  |                       +----- Grandidieria **                 
  |  |  +--------------------+                                   
  +--+  |                    |  +-- Pseudospatha **                 
     |  |                    +--+                                
     |  |                       +-- Moncetia **                     
     |  |                                                        
     +--+  +----------------------- Nitia **                        
        |  |                                                     
        |  |                    +-- Coelatura_aegyptiaca **         
        |  |                    |                                
        |  |                    +-- Coelatura_alluaudi **           
        +--+                    |                                
           |  +-----------------+-- Coelatura_bakeri **             
           |  |                 |                                
           |  |                 +-- Coelatura_hauttecoeuri **       
           |  |                 |                                
           |  |                 +-- Coelatura_stuhlmanni **         
           +--+                                                  
              |                 +-- Coelatura_leopoldvillensis **   
              |  +--------------+                                
              |  |              +-- Coelatura_mesafricana **        
              |  |                                               
              |  |              +-- Coelatura_horei **              
              +--+              |                                
                 |  +-----------+-- Coelatura_ratidota **           
                 |  |           |                                
                 |  |           +-- Coelatura **                    
                 |  |                                            
                 +--+           +-- Coelatura_gabonensis **         
                    |  +--------+                                
                    |  |        +-- Coelatura_stanleyvillensis **   
                    |  |                                         
                    +--+  +-------- Coelatura_kunenensis **         
                       |  |                                      
                       |  |     +-- Coelatura_briarti **            
                       |  |     |                                
                       +--+  +--+-- Prisodontopsis_mweruensis **    
                          |  |  |                                
                          |  |  +-- Prisodontopsis_aviculaeformis **
                          |  |                                   
                          +--+  +-- Coelatura_choziensis **         
                             |  |                                
                             |  +-- Coelatura_hypsiprymna **        
                             |  |                                
                             +--+-- Coelatura_luapulaensis **       
                                |                                
                                +-- Coelatura_mossambicensis **     
                                |                                
                                +-- Nyassunio **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
11 11 12 2 17 0 0 28 
  +----------------- Coelatura_horei **          
  |                                           
  |              +-- Coelatura_gabonensis **     
  |     +--------+                            
  |     |        +-- Coelatura_aegyptiaca__4 **  
  |     |                                     
  |  +--+  +-------- Coelatura_kunenensis **     
--+  |  |  |                                  
  |  |  |  |  +----- Coelatura_luapulaensis__1 **
  |  |  +--+  |                               
  |  |     |  +----- Coelatura_luapulaensis__3 **
  |  |     |  |                               
  |  |     +--+----- Coelatura_luapulaensis__4 **
  |  |        |                               
  |  |        +----- Coelatura_choziensis__2 **  
  +--+        |                               
     |        |  +-- Coelatura_luapulaensis__2 **
     |        +--+                            
     |           +-- Coelatura_choziensis__1 **  
     |                                        
     |  +----------- Coelatura_alluaudi__2 **    
     |  |                                     
     |  +----------- Nitia **                    
     |  |                                     
     |  |  +-------- Coelatura_aegyptiaca__1 **  
     +--+  |                                  
        |  +-------- Coelatura_bakeri__2 **      
        |  |                                  
        |  |     +-- Coelatura_aegyptiaca__2 **  
        |  +-----+                            
        +--+     +-- Coelatura_stuhlmanni__1 **  
           |                                  
           |     +-- Coelatura_stuhlmanni__2 **  
           +-----+                            
           |     +-- Coelatura_bakeri__1 **      
           |                                  
           |  +----- Coelatura_aegyptiaca__3 **  
           +--+                               
              |  +-- Coelatura_alluaudi__1 **    
              +--+                            
                 +-- Coelatura_hauttecoeuri **

Graf et al., 2014 [Publ 3591], fig. 3

iTtTbiriSoTroTroSiD
8 8 11 3 3 1 1 24 
  +----------------------- Leoparreysiini           
  |                                                 
--+  +-------------------- Nitia **                    
  |  |                                              
  +--+  +----------------- Coelatura_aegyptiaca__2 **  
     |  |                                           
     +--+  +-------------- Coelatura_aegyptiaca__1 **  
        |  |                                        
        +--+  +----------- Coelatura_gabonensis **     
           |  |                                     
           +--+  +-------- Coelatura_kunenensis **     
              |  |                                  
              |  |  +----- Prisodontopsis **           
              +--+  |                               
                 |  |  +-- Coelatura_choziensis__1 **  
                 |  |  |                            
                 +--+  +-- Coelatura_choziensis__2 **  
                    |  |                            
                    |  +-- Coelatura_luapulaensis__1 **
                    |  |                            
                    +--+-- Coelatura_luapulaensis__2 **
                       |                            
                       +-- Coelatura_luapulaensis__3 **
                       |                            
                       +-- Coelatura_luapulaensis__4 **
                       |                            
                       +-- Coelatura_luapulaensis__5 **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
8 8 0 3 117 30 4 26 
           +-------------- Trapezidens                  
           |                                            
  +--------+  +----------- Lamellidens_lamellatus       
  |        |  |                                         
  |        +--+  +-------- Lamellidens_jenkinsianus     
  |           |  |                                      
  |           |  |  +----- Lamellidens_generosus        
  |           +--+  |                                   
  |              |  +----- Lamellidens_mainwaringi      
  |              |  |                                   
  |              +--+  +-- Lamellidens_ferrugineus      
  |                 +--+                                
--+                 |  +-- Lamellidens_marginalis       
  |                 |                                   
  |                 |  +-- Lamellidens_savadiensis      
  |                 +--+                                
  |                    +-- Lamellidens_corrianus__1     
  |                                                     
  |  +-------------------- Parreysiini                  
  |  |                                                  
  |  |  +----------------- Leoparreysiini               
  |  |  |                                               
  |  |  |              +-- Nitia **                        
  +--+  |              |                                
     |  |              +-- Coelatura_gabonensis **         
     |  |              |                                
     |  |              +-- Prisodontopsis_mweruensis **    
     |  |              |                                
     +--+              +-- Coelatura_kunenensis **         
        |              |                                
        |  +-----------+-- Coelatura_aegyptiaca__1 **      
        |  |           |                                
        |  |           +-- Coelatura_aegyptiaca__2 **      
        |  |           |                                
        |  |           +-- Prisodontopsis_aviculaeformis **
        |  |           |                                
        |  |           +-- Coelatura_luapulaensis **       
        +--+           |                                
           |           +-- Coelatura_choziensis **         
           |                                            
           |        +----- Indochinella                 
           |  +-----+                                   
           |  |     |  +-- Scabies                      
           |  |     +--+                                
           +--+        +-- Radiatula                    
              |                                         
              |  +-------- Indonaia_shurtleffiana       
              |  |                                      
              +--+  +----- Indonaia_rugosa              
                 |  |                                   
                 |  |  +-- Indonaia_andersoniana        
                 +--+--+                                
                    |  +-- Indonaia_caerulea            
                    |                                   
                    |  +-- Indonaia_bonneaudii          
                    +--+                                
                       +-- Lamellidens_corrianus__2

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 4 3 194 29 4 27 
  +----------------------- Lamellidentini         
  |                                               
--+  +-------------------- Parreysiini            
  |  |                                            
  +--+  +----------------- Leoparreysiini         
     |  |                                         
     +--+  +-------------- Indochinellini         
        |  |                                      
        +--+  +----------- Coelatura_aegyptiaca__2 **
           |  |                                   
           +--+        +-- Coelatura_aegyptiaca__1 **
              |  +-----+                          
              |  |     +-- Nitia **                  
              +--+                                
                 |  +----- Coelatura_kunenensis **   
                 +--+                             
                    |  +-- Coelatura_gabonensis **   
                    +--+                          
                       +-- Prisodontopsis **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
5 5 4 3 134 24 4 25 
  +-------------------------- Lamellidentini         
--+                                                  
  |  +----------------------- Parreysiini            
  +--+                                               
     |  +-------------------- Leoparreysiini         
     +--+                                            
        |  +----------------- Indochinellini         
        +--+                                         
           |  +-------------- Nitia **                  
           +--+                                      
              |  +----------- Coelatura_aegyptiaca__2 **
              +--+                                   
                 |  +-------- Coelatura_aegyptiaca__1 **
                 +--+                                
                    |  +----- Prisodontopsis **         
                    +--+                             
                       |  +-- Coelatura_gabonensis **   
                       +--+                          
                          +-- Coelatura_kunenensis **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
5 5 4 3 134 24 4 23 
                       +-- Parreysiini            
  +--------------------+                          
  |                    +-- Lamellidentini         
--+                                               
  |  +-------------------- Leoparreysiini         
  +--+                                            
     |  +----------------- Indochinellini         
     +--+                                         
        |  +-------------- Nitia **                  
        +--+                                      
           |  +----------- Coelatura_aegyptiaca__1 **
           +--+                                   
              |  +-------- Coelatura_aegyptiaca__2 **
              +--+                                
                 |  +----- Coelatura_gabonensis **   
                 +--+                             
                    |  +-- Coelatura_kunenensis **   
                    +--+                          
                       +-- Prisodontopsis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 2 3 196 29 4 25 
  +-------------------- Lamellidentini         
  |                                            
--+  +----------------- Parreysiini            
  |  |                                         
  +--+  +-------------- Leoparreysiini         
     |  |                                      
     +--+  +----------- Indochinellini         
        |  |                                   
        +--+  +-------- Coelatura_aegyptiaca__2 **
           |  |                                
           +--+  +----- Coelatura_aegyptiaca__1 **
              |  |                             
              |  +----- Nitia **                  
              +--+                             
                 |  +-- Coelatura_gabonensis **   
                 |  |                          
                 +--+-- Coelatura_kunenensis **   
                    |                          
                    +-- Prisodontopsis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 2 3 196 29 4 24 
  +----------------- Parreysiini            
  |                                         
  +----------------- Lamellidentini         
--+                                         
  |  +-------------- Leoparreysiini         
  |  |                                      
  +--+  +----------- Indochinellini         
     |  |                                   
     +--+  +-------- Coelatura_aegyptiaca__2 **
        |  |                                
        |  +-------- Nitia **                  
        +--+                                
           |  +----- Coelatura_aegyptiaca__1 **
           |  |                             
           +--+  +-- Coelatura_gabonensis **   
              |  |                          
              +--+-- Coelatura_kunenensis **   
                 |                          
                 +-- Prisodontopsis **

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
4 4 2 3 59 58 4 23 
  +-------------------- Parreysia           
  |                                         
--+                 +-- Leoparreysiini      
  |  +--------------+                       
  |  |              +-- Lamellidentini      
  +--+                                      
     |  +-------------- Assamnaia           
     +--+                                   
        |  +----------- Indochinellini      
        +--+                                
           |  +-------- Nitia **               
           +--+                             
              |  +----- Coelatura_aegyptiaca **
              +--+                          
                 |  +-- Nyassunio **           
                 +--+                       
                    +-- Coelatura_kunenensis **

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 3 54 50 4 23 
  +----------------- Lamellidentini      
  |                                      
--+              +-- Parreysiini         
  |  +-----------+                       
  |  |           +-- Leoparreysiini      
  +--+                                   
     |  +----------- Indochinellini      
     +--+                                
        |  +-------- Nitia **               
        +--+                             
           |  +----- Coelatura_aegyptiaca **
           +--+                          
              |  +-- Coelatura_kunenensis **
              +--+                       
                 +-- Nyassunio **

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
4 4 0 3 57 53 4 25 
  +-------------- Parreysiini         
  |                                   
--+  +----------- Lamellidentini      
  |  |                                
  +--+  +-------- Leoparreysiini      
     |  |                             
     +--+  +----- Indochinellini      
        |  |                          
        +--+  +-- Coelatura_aegyptiaca **
           |  |                       
           |  +-- Coelatura_kunenensis **
           +--+                       
              +-- Nitia **               
              |                       
              +-- Nyassunio **

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
4 4 2 3 54 50 4 23 
  +-------------------- Lamellidentini      
--+                                         
  |  +----------------- Parreysiini         
  +--+                                      
     |  +-------------- Leoparreysiini      
     +--+                                   
        |  +----------- Indochinellini      
        +--+                                
           |  +-------- Nitia **               
           +--+                             
              |  +----- Coelatura_aegyptiaca **
              +--+                          
                 |  +-- Coelatura_kunenensis **
                 +--+                       
                    +-- Nyassunio **

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
4 4 2 3 54 50 4 23 
  +-------------------- Parreysiini         
--+                                         
  |  +----------------- Lamellidentini      
  +--+                                      
     |  +-------------- Leoparreysiini      
     +--+                                   
        |  +----------- Indochinellini      
        +--+                                
           |  +-------- Nitia **               
           +--+                             
              |  +----- Coelatura_aegyptiaca **
              +--+                          
                 |  +-- Coelatura_kunenensis **
                 +--+                       
                    +-- Nyassunio **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
4 4 3 3 32 5 3 24 
  +-------------------- Lamellidentini         
--+                                            
  |  +----------------- Leoparreysiini         
  +--+                                         
     |  +-------------- Indochinellini         
     +--+                                      
        |  +----------- Nitia **                  
        +--+                                   
           |  +-------- Coelatura_aegyptiaca__2 **
           +--+                                
              |  +----- Coelatura_aegyptiaca__1 **
              +--+                             
                 |  +-- Coelatura_gabonensis **   
                 +--+                          
                    +-- Prisodontopsis **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
4 4 2 3 32 5 3 24 
  +----------------- Lamellidentini         
  |                                         
--+  +-------------- Leoparreysiini         
  |  |                                      
  +--+  +----------- Indochinellini         
     |  |                                   
     +--+  +-------- Nitia **                  
        |  |                                
        +--+  +----- Coelatura_aegyptiaca__1 **
           |  |                             
           +--+----- Coelatura_aegyptiaca__2 **
              |                             
              |  +-- Coelatura_gabonensis **   
              +--+                          
                 +-- Prisodontopsis **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 3 57 53 4 23 
  +-------------------- Lamellidentini      
--+                                         
  |  +----------------- Parreysiini         
  +--+                                      
     |  +-------------- Leoparreysiini      
     +--+                                   
        |  +----------- Indochinellini      
        +--+                                
           |  +-------- Nitia **               
           +--+                             
              |  +----- Coelatura_aegyptiaca **
              +--+                          
                 |  +-- Nyassunio **           
                 +--+                       
                    +-- Coelatura_kunenensis **

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 25 24 4 24 
  +----------------- Lamellidentini
--+                                
  |  +-------------- Parreysiini   
  +--+                             
     |  +----------- Leoparreysiini
     +--+                          
        |  +-------- Indochinellini
        +--+                       
           |  +----- Nitia **         
           +--+                    
              |  +-- Coelatura **     
              +--+                 
                 +-- Prisodontopsis **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 110 38 4 -- 
  +----------- Lamellidentini
  |                          
--+        +-- Parreysiini   
  |  +-----+                 
  |  |     +-- Leoparreysiini
  +--+                       
     |  +----- Indochinellini
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Nitia **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 116 39 4 -- 
  +----------- Lamellidentini
  |                          
--+        +-- Parreysiini   
  |  +-----+                 
  |  |     +-- Leoparreysiini
  +--+                       
     |  +----- Indochinellini
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Nitia **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 81 22 4 -- 
  +-------------- Lamellidentini
--+                             
  |  +----------- Parreysiini   
  +--+                          
     |  +-------- Leoparreysiini
     +--+                       
        |  +----- Indochinellini
        +--+                    
           |  +-- Coelatura **     
           +--+                 
              +-- Nitia **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 44 3 3 -- 
  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Indochinellini
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Nitia **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
2 2 0 2 55 8 3 -- 
  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Indochinellini
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Prisodontopsis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 55 8 3 -- 
  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Indochinellini
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Prisodontopsis **

Bolotov et al., 2019 [Publ 3974], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 16 11 2 -- 
  +-------- Leoparreysiini
--+                       
  |  +----- Indochinellini
  +--+                    
     |  +-- Nitia **         
     +--+                 
        +-- Coelatura **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 10 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelatura **     
     +--+                 
        +-- Indochinellini

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 2 2 -- 
  +-- Coelatura **     
  |                 
--+-- Leoparreysiini
  |                 
  +-- Indochinellini

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 7 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelatura **     
     +--+                 
        +-- Indochinellini

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 2 2 -- 
  +----- Parreysiini   
--+                    
  |  +-- Coelatura **     
  +--+                 
     +-- Indochinellini

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Coelatura_aegyptiaca **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 12 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelatura **     
     +--+                 
        +-- Indochinellini

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Coelatura_aegyptiaca **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 22 22 4 -- 
  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Parreysiini   
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Indochinellini

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Coelatura_aegyptiaca **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Coelatura_aegyptiaca **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 2 2 -- 
  +-- Coelatura **     
  |                 
--+-- Indochinellini
  |                 
  +-- Parreysiini

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Coelatura_aegyptiaca **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Coelatura_aegyptiaca **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 2 2 -- 
  +-- Coelatura **     
  |                 
--+-- Indochinellini
  |                 
  +-- Parreysiini

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Lamellidentini
--+                 
  +-- Coelatura **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Coelatura_aegyptiaca **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 1 1 -- 
-- Grandidieria **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Coelatura_aegyptiaca **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Coelatura_aegyptiaca **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 44 2 2 -- 
  +----- Parreysiini   
--+                    
  |  +-- Coelatura **     
  +--+                 
     +-- Indochinellini

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 3 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelatura **     
     +--+                 
        +-- Indochinellini

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 1 1 -- 
  +-- Lamellidentini
--+                 
  +-- Coelatura **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Coelatura_aegyptiaca **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 12 3 -- 
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelatura **     
     +--+                 
        +-- Indochinellini
 
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