MUSSELpdb | tribe Contradentini Modell, 1942 cladomics

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	all genera Unionidae Margaritiferidae Hyriidae Etheriidae Mycetopodidae Iridinidae Species & Genus Richness by Taxonomy


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Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae

tribe CONTRADENTINI Modell, 1942: cladomics
6 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 35 (35 total)
number of subtaxa (rank: genus) = 6
analyzed in 71 cladograms in 45 publications
maximum number of terminals in an analysis = 23 (mean = 6.35, median = 4)
maximum number of subtaxa (genus) = 5 (mean = 2.66, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 62
maximum number of included terminals = 23, (mean = 7.13 median = 5)
maximum number of subtaxa (genus) = 5 (mean = 2.9, median = 3)
proportion of cladograms that support basic monophyly = 0.97

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 56
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 23 (mean = 6.82, median = 4)
maximum number of subtaxa (genus) = 5 (mean = 2.98, median = 3)
proportion of cladograms that support rigorous monophyly = 0.96

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	52	12	5	0

  +----------------- Chamberlainiini
  |                                 
  |           +----- Lamprotulini   
--+  +--------+                     
  |  |        |  +-- Gonideini      
  |  |        +--+                  
  +--+           +-- Pseudodontini  
     |                              
     |  +----------- Rectidentini   
     |  |                           
     +--+        +-- Trapezoideus **   
        |  +-----+                  
        |  |     +-- Physunio **       
        +--+                        
           |  +----- Yaukthwa **       
           +--+                     
              |  +-- Lens **           
              +--+                  
                 +-- Solenaia **

Zieritz et al., 2024 [Publ 4381], fig. 2

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
8	8	6	4	38	27	8	5

              +-- Schepmaniini 
        +-----+                
        |     +-- Pseudodontini
  +-----+                      
  |     |  +----- Caudiculatus 
  |     +--+                   
  |        |  +-- Lamprotulini 
--+        +--+                
  |           +-- Gonideini    
  |                            
  |  +----------- Ctenodesmini 
  |  |                         
  +--+  +-------- Rectidentini 
     |  |                      
     +--+     +-- Physunio **     
        |  +--+                
        |  |  +-- Trapezoideus ** 
        +--+                   
           |  +-- Yaukthwa **     
           +--+                
              +-- Lens **

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
23	23	21	4	89	49	5	22

        +-------- Pseudodontini  
  +-----+                        
  |     |  +----- Gonideini      
  |     +--+                     
  |        |  +-- Chamberlainiini
--+        +--+                  
  |           +-- Lamprotulini   
  |                              
  |  +----------- Rectidentini   
  +--+                           
     |  +-------- Physunio **       
     +--+                        
        |  +----- Trapezoideus **   
        +--+                     
           |  +-- Lens **           
           +--+                  
              +-- Yaukthwa **

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
23	23	21	4	95	49	5	20

     +-------- Rectidentini             
     |                                  
  +--+     +-- Trapezoideus **             
  |  |  +--+                            
  |  |  |  +-- Physunio **                 
  |  +--+                               
  |     |  +-- Yaukthwa **                 
  |     +--+                            
  |        +-- Lens **                     
  |                                     
--+     +----- Pseudodontini            
  |     |                               
  |     |  +-- Sinosolenaia_iridinea    
  |     |  |                            
  |     |  +-- Sinosolenaia_recognita__1
  |     |  |                            
  |     |  +-- Sinosolenaia_carinata    
  |     |  |                            
  |     |  +-- Leguminaia               
  +-----+  |                            
        |  +-- Microcondylaea           
        |  |                            
        |  +-- Gonidea                  
        |  |                            
        |  +-- Ptychorhynchus           
        |  |                            
        |  +-- Parvasolenaia            
        |  |                            
        +--+-- Lamprotula_leaii         
           |                            
           +-- Lamprotula               
           |                            
           +-- Lamprotula_caveata       
           |                            
           +-- Potomida                 
           |                            
           +-- Pronodularia             
           |                            
           +-- Sinohyriopsis            
           |                            
           +-- Chamberlainia            
           |                            
           +-- Sinosolenaia_recognita__2
           |                            
           +-- Sinosolenaia_oleivora

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
19	19	17	4	63	49	4	17

  +-------------- Pseudodontini
  |                            
--+           +-- Gonideini    
  |  +--------+                
  |  |        +-- Lamprotulini 
  +--+                         
     |  +-------- Rectidentini 
     |  |                      
     +--+     +-- Physunio **     
        |  +--+                
        |  |  +-- Trapezoideus ** 
        +--+                   
           |  +-- Yaukthwa **     
           +--+                
              +-- Lens **

Konopleva et al., 2019 [Publ 4041], fig. 3

tT	bi	riS	oT	roT	roS	iD
19	19	17	4	15	15	2	18

  +-------------- Pseudodontini
--+                            
  |  +----------- Rectidentini 
  +--+                         
     |  +-------- Yaukthwa **     
     +--+                      
        |  +----- Lens **         
        +--+                   
           |  +-- Trapezoideus ** 
           +--+                
              +-- Physunio **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
16	16	14	4	36	22	2	17

  +----------- Rectidentini
--+                        
  |  +-------- Yaukthwa **    
  +--+                     
     |  +----- Physunio **    
     +--+                  
        |  +-- Trapezoideus **
        +--+               
           +-- Lens **

Bolotov et al., 2022 [Publ 4250], fig. S5

tT	bi	riS	oT	roT	roS	iD
15	15	12	3	0	0	0	14

  +----- Yaukthwa **    
--+                  
  |  +-- Physunio **    
  +--+               
     +-- Trapezoideus **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

tT	bi	riS	oT	roT	roS	iD
15	15	13	5	14	14	2	10

  +-------------- Lamprotulini
  |                           
--+  +----------- Rectidentini
  |  |                        
  +--+        +-- Trapezoideus **
     |  +-----+               
     |  |     +-- Physunio **    
     +--+                     
        |  +----- Yaukthwa **    
        +--+                  
           |  +-- Solenaia **    
           +--+               
              +-- Lens **

Zieritz et al., 2020 [Publ 4102], fig. 4

tT	bi	riS	oT	roT	roS	iD
14	14	12	4	17	17	6	15

  +----------------------- Chamberlainiini
  |                                       
--+  +-------------------- Lamprotulini   
  |  |                                    
  +--+  +----------------- Gonideini      
     |  |                                 
     +--+  +-------------- Pseudodontini  
        |  |                              
        +--+           +-- Ctenodesmini   
           |  +--------+                  
           |  |        +-- Rectidentini   
           +--+                           
              |  +-------- Physunio **       
              +--+                        
                 |  +----- Yaukthwa **       
                 +--+                     
                    |  +-- Trapezoideus **   
                    +--+                  
                       +-- Lens **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	113	39	5	12

  +-------- Rectidentini
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	187	44	5	12

  +-------- Rectidentini
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
12	12	7	3	187	44	5	13

  +-------------- Rectidentini     
  |                                
  |  +----------- Physunio_modelli ** 
--+  |                             
  |  +----------- Lens **             
  |  |                             
  |  +----------- Physunio_superbus **
  +--+                             
     +----------- Yaukthwa **         
     |                             
     |        +-- Lens_comptus **     
     |  +-----+                    
     |  |     +-- Lens_novoselovi **  
     +--+                          
        |  +----- Lens_eximius **     
        +--+                       
           |  +-- Lens_contradens **  
           +--+                    
              +-- Lens_micropterus **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	127	37	5	12

  +-------- Rectidentini
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Zieritz et al., 2024 [Publ 4381], fig. 7A

tT	bi	riS	oT	roT	roS	iD
12	12	10	2	0	0	0	13

  +-- Solenaia **
--+           
  +-- Lens **

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
12	12	10	2	37	11	3	13

  +----- Pseudodontini
--+                   
  |  +-- Trapezoideus ** 
  +--+                
     +-- Yaukthwa **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	185	43	5	12

  +-------- Rectidentini
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
11	11	9	3	128	38	5	13

  +-------- Rectidentini
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Physunio **    
     +--+               
        +-- Yaukthwa **

Pfeiffer et al., 2021 [Publ 4126], fig. 12

tT	bi	riS	oT	roT	roS	iD
11	11	8	2	0	0	0	12

  +-- Solenaia **
--+           
  +-- Lens **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
10	10	0	3	73	25	4	11

  +----- Rectidentini       
  |                         
  |  +-- Lens_eximius **       
  |  |                      
  |  +-- Physunio_modelli__1 **
  |  |                      
--+  +-- Yaukthwa_nesemanni ** 
  |  |                      
  |  +-- Yaukthwa_panhai **    
  |  |                      
  |  +-- Lens_comptus **       
  |  |                      
  +--+-- Lens_novoselovi **    
     |                      
     +-- Physunio_modelli__2 **
     |                      
     +-- Physunio **           
     |                      
     +-- Yaukthwa_inlenensis **
     |                      
     +-- Lens_contradens **    
     |                      
     +-- Lens_micropterus **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

tT	bi	riS	oT	roT	roS	iD
10	10	8	4	11	11	1	8

  +-------- Rectidentini
  |                     
--+     +-- Trapezoideus **
  |  +--+               
  |  |  +-- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Pfeiffer et al., 2021 [Publ 4126], fig. 25

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	0	0	0	10

  +-- Solenaia **
--+           
  +-- Yaukthwa **

Pfeiffer et al., 2021 [Publ 4126], fig. 18

tT	bi	riS	oT	roT	roS	iD
8	8	7	3	0	0	0	8

  +----- Yaukthwa **    
--+                  
  |  +-- Trapezoideus **
  +--+               
     +-- Physunio **

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	30	28	7	4

  +----------------- Chamberlainiini
  |                                 
  |              +-- Schepmaniini   
  |           +--+                  
--+           |  +-- Pseudodontini  
  |  +--------+                     
  |  |        |  +-- Lamprotulini   
  |  |        +--+                  
  +--+           +-- Gonideini      
     |                              
     |  +----------- Ctenodesmini   
     |  |                           
     +--+  +-------- Rectidentini   
        |  |                        
        +--+     +-- Physunio **       
           |  +--+                  
           |  |  +-- Trapezoideus **   
           +--+                     
              |  +-- Lens **           
              +--+                  
                 +-- Yaukthwa **

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
7	7	1	4	30	28	7	6

  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
  |                                                        
  |  +-------------------------- Pseudodontini             
  |  |                                                     
--+  |                    +----- Ensidens                  
  |  |                    |                                
  |  |                    +----- Hyriopsis_desowitzi       
  |  |                    |                                
  |  |                    +----- Rectidens                 
  |  |                    |                                
  |  |                    +----- Hyriopsis_bialata         
  +--+  +-----------------+                                
     |  |                 +----- Ctenodesmini              
     |  |                 |                                
     |  |                 |  +-- Physunio **                  
     |  |                 |  |                             
     |  |                 |  +-- Yaukthwa_paiensis **         
     |  |                 |  |                             
     |  |                 |  +-- Yaukthwa_inlenensis **       
     |  |                 +--+                             
     +--+                    +-- Trapezoideus **              
        |                    |                             
        |                    +-- Yaukthwa_elongatula **       
        |                    |                             
        |                    +-- Lens **                      
        |                                                  
        |  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	31	29	7	4

  +----------------- Chamberlainiini
  |                                 
  |        +-------- Lamprotulini   
--+  +-----+                        
  |  |     |  +----- Gonideini      
  |  |     +--+                     
  |  |        |  +-- Schepmaniini   
  +--+        +--+                  
     |           +-- Pseudodontini  
     |                              
     |  +----------- Ctenodesmini   
     |  |                           
     +--+  +-------- Rectidentini   
        |  |                        
        +--+     +-- Physunio **       
           |  +--+                  
           |  |  +-- Trapezoideus **   
           +--+                     
              |  +-- Lens **           
              +--+                  
                 +-- Yaukthwa **

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	25	21	7	6

  +----------------------- Chamberlainiini
  |                                       
--+  +-------------------- Lamprotulini   
  |  |                                    
  |  |              +----- Gonideini      
  +--+  +-----------+                     
     |  |           |  +-- Schepmaniini   
     |  |           +--+                  
     +--+              +-- Pseudodontini  
        |                                 
        |  +-------------- Ctenodesmini   
        +--+                              
           |  +----------- Rectidentini   
           +--+                           
              |  +-------- Physunio **       
              +--+                        
                 |  +----- Trapezoideus **   
                 +--+                     
                    |  +-- Lens **           
                    +--+                  
                       +-- Yaukthwa **

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	30	28	7	4

                    +-- Schepmaniini   
  +-----------------+                  
  |                 +-- Pseudodontini  
--+                                    
  |  +----------------- Gonideini      
  |  |                                 
  +--+              +-- Chamberlainiini
     |  +-----------+                  
     |  |           +-- Lamprotulini   
     +--+                              
        |  +----------- Ctenodesmini   
        |  |                           
        +--+  +-------- Rectidentini   
           |  |                        
           +--+     +-- Physunio **       
              |  +--+                  
              |  |  +-- Trapezoideus **   
              +--+                     
                 |  +-- Lens **           
                 +--+                  
                    +-- Yaukthwa **

Wu et al., 2024 [Publ 4350], fig. 5b

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	30	28	7	6

  +-------------------------------- Schepmaniini       
  |                                                    
  |  +----------------------------- Pseudodontini      
  |  |                                                 
--+  |     +----------------------- Hyriopsis_bialata  
  |  |     |                                           
  |  |  +--+                    +-- Rectidens          
  |  |  |  |  +-----------------+                      
  |  |  |  |  |                 +-- Ctenodesmini       
  +--+  |  +--+                                        
     |  |     |  +----------------- Ensidens           
     |  |     |  |                                     
     |  |     +--+  +-------------- Hyriopsis_desowitzi
     |  |        |  |                                  
     |  |        +--+  +----------- Yaukthwa_elongatula **
     |  |           |  |                               
     |  |           +--+  +-------- Trapezoideus **       
     +--+              |  |                            
        |              +--+     +-- Physunio **           
        |                 |  +--+                      
        |                 |  |  +-- Yaukthwa_inlenensis **
        |                 +--+                         
        |                    |  +-- Yaukthwa_paiensis **  
        |                    +--+                      
        |                       +-- Lens **               
        |                                              
        |     +-------------------- Inversidens        
        |     |                                        
        |     |              +----- Lamprotula         
        +-----+  +-----------+                         
              |  |           |  +-- Chamberlainiini    
              |  |           +--+                      
              +--+              +-- Potomida           
                 |                                     
                 |  +-------------- Sinosolenaia       
                 |  |                                  
                 +--+           +-- Gonidea            
                    |  +--------+                      
                    |  |        +-- Microcondylaea     
                    +--+                               
                       |  +-------- Koreosolenaia      
                       +--+                            
                          |  +----- Parvasolenaia      
                          +--+                         
                             |  +-- Postolata          
                             +--+                      
                                +-- Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	30	28	7	4

  +-------------------------- Chamberlainiini
  |                                          
  |              +----------- Ctenodesmini   
  |              |                           
  |  +-----------+  +-------- Rectidentini   
--+  |           |  |                        
  |  |           +--+     +-- Physunio **       
  |  |              |  +--+                  
  |  |              |  |  +-- Trapezoideus **   
  |  |              +--+                     
  |  |                 |  +-- Lens **           
  +--+                 +--+                  
     |                    +-- Yaukthwa **       
     |                                       
     |                    +-- Schepmaniini   
     |  +-----------------+                  
     |  |                 +-- Pseudodontini  
     |  |                                    
     +--+                 +-- Inversidens    
        |  +--------------+                  
        |  |              +-- Lamprotulini   
        +--+                                 
           |  +-------------- Sinosolenaia   
           |  |                              
           +--+           +-- Gonidea        
              |  +--------+                  
              |  |        +-- Microcondylaea 
              +--+                           
                 |  +-------- Parvasolenaia  
                 +--+                        
                    |  +----- Koreosolenaia  
                    +--+                     
                       |  +-- Postolata      
                       +--+                  
                          +-- Ptychorhynchus

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

tT	bi	riS	oT	roT	roS	iD
6	6	11	2	3	2	2	18

  +----- Rectidentini
--+                  
  |  +-- Physunio **    
  +--+               
     +-- Lens **

Bolotov et al., 2019 [Publ 3974], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	13	0	0	9

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	52	12	5	0

  +----------------- Chamberlainiini
  |                                 
  |           +----- Pseudodontini  
--+  +--------+                     
  |  |        |  +-- Gonideini      
  |  |        +--+                  
  +--+           +-- Lamprotulini   
     |                              
     |  +----------- Rectidentini   
     |  |                           
     +--+        +-- Trapezoideus **   
        |  +-----+                  
        |  |     +-- Physunio **       
        +--+                        
           |  +----- Yaukthwa **       
           +--+                     
              |  +-- Lens **           
              +--+                  
                 +-- Solenaia **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	87	25	5	6

  +-------- Rectidentini
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Lens **        
     +--+               
        +-- Yaukthwa **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	87	25	5	6

  +-------- Rectidentini
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Lens **        
     +--+               
        +-- Yaukthwa **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

tT	bi	riS	oT	roT	roS	iD
4	4	1	3	3	3	1	5

  +----- Rectidentini
  |                  
--+  +-- Physunio **    
  |  |               
  +--+-- Trapezoideus **
     |               
     +-- Lens **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	28	7	2	5.5

  +-------- Pseudodontini
--+                      
  |  +----- Rectidentini 
  +--+                   
     |  +-- Lens **         
     +--+                
        +-- Physunio **

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	42	40	5	4

  +----------------------- Pseudodontini  
  |                                       
  |                 +----- Rectidentini   
  |  +--------------+                     
--+  |              |  +-- Physunio **       
  |  |              +--+                  
  |  |                 +-- Lens **           
  |  |                                    
  +--+              +----- Inversidens    
     |  +-----------+                     
     |  |           |  +-- Chamberlainiini
     |  |           +--+                  
     |  |              +-- Lamprotulini   
     +--+                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	73	28	5	2

  +-------- Rectidentini
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Trapezoideus **
     +--+               
        +-- Physunio **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
3	4	2	2	77	24	5	6

  +-------- Rectidentini
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Physunio **    
     +--+               
        +-- unionoid

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	55	55	7	4

           +-------- Lamprotulini   
  +--------+                        
  |        |  +----- Gonideini      
  |        +--+                     
  |           |  +-- Schepmaniini   
--+           +--+                  
  |              +-- Pseudodontini  
  |                                 
  |  +-------------- Chamberlainiini
  +--+                              
     |  +----------- Ctenodesmini   
     +--+                           
        |  +-------- Rectidentini   
        +--+                        
           |  +----- Physunio **       
           +--+                     
              |  +-- Lens **           
              +--+                  
                 +-- Trapezoideus **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	33	33	5	2

     +-------- Rectidentini   
  +--+                        
  |  |  +----- Lens **           
  |  +--+                     
  |     |  +-- Trapezoideus **   
--+     +--+                  
  |        +-- Physunio **       
  |                           
  |  +-------- Pseudodontini  
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Chamberlainiini
        +--+                  
           +-- Lamprotulini

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	37	11	4	5.5

     +----- Rectidentini 
  +--+                   
  |  |  +-- Lens **         
  |  +--+                
--+     +-- Physunio **     
  |                      
  |  +----- Pseudodontini
  +--+                   
     |  +-- Gonideini    
     +--+                
        +-- Lamprotulini

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	42	40	5	4

  +----------------------- Pseudodontini  
  |                                       
  |                 +----- Rectidentini   
--+  +--------------+                     
  |  |              |  +-- Physunio **       
  |  |              +--+                  
  |  |                 +-- Lens **           
  +--+                                    
     |  +----------------- Inversidens    
     |  |                                 
     |  |              +-- Chamberlainiini
     |  +--------------+                  
     +--+              +-- Lamprotulini   
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	33	33	5	2

     +-------- Rectidentini   
  +--+                        
  |  |  +----- Lens **           
  |  +--+                     
  |     |  +-- Trapezoideus **   
--+     +--+                  
  |        +-- Physunio **       
  |                           
  |  +-------- Pseudodontini  
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Chamberlainiini
        +--+                  
           +-- Lamprotulini

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
3	5	0	3	118	30	5	7

  +-- Lens **     
  |            
  +-- Ensidens 
  |            
--+-- Physunio ** 
  |            
  +-- Rectidens
  |            
  +-- Yaukthwa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	73	28	5	2

  +-------- Rectidentini
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Trapezoideus **
     +--+               
        +-- Physunio **

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	44	39	7	4

        +-------- Ctenodesmini   
  +-----+                        
  |     |  +----- Rectidentini   
  |     +--+                     
  |        |  +-- Physunio **       
--+        +--+                  
  |           +-- Lens **           
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	45	40	7	4

        +-------- Ctenodesmini   
  +-----+                        
  |     |  +----- Rectidentini   
  |     +--+                     
  |        |  +-- Physunio **       
--+        +--+                  
  |           +-- Lens **           
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Zhang et al., 2024 [Publ 4397], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	26	24	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	83	16	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Matos et al., 2024 [Publ 4384], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	18	16	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Physunio **       
  |  |  +--+                  
  +--+     +-- Lens **           
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	21	10	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Pseudodontini  
     +--+                     
        |  +-- Lamprotulini   
        +--+                  
           +-- Gonideini

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	52	21	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Matos et al., 2024 [Publ 4383], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	18	16	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Physunio **       
  |  |  +--+                  
  +--+     +-- Lens **           
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	93	18	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	19	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	86	17	5	--

  +----- Rectidentini
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	93	18	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	15	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	51	17	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	19	5	--

  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Smith, 2021 [Publ 4331], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	2	2	--

  +----- Chamberlainiini
--+                     
  |  +-- Lens **           
  +--+                  
     +-- Lamprotulini

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	10	4	--

  +-------- Chamberlainiini
  |                        
--+     +-- Gonideini      
  |  +--+                  
  |  |  +-- Pseudodontini  
  +--+                     
     |  +-- Physunio **       
     +--+                  
        +-- Lamprotulini

Graf & Cummings, 2006 [Publ 1968], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	3	2	--

-- Lens_contradens **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	5	3	--

-- Lens_contradens **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	5	3	--

  +-- Pseudodontini
--+                
  +-- Lens **

Hoeh et al., 2009 [Publ 2716], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	3	2	--

-- Lens_contradens **

Hoeh et al., 2009 [Publ 2716], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	3	2	--

-- Lens_contradens **

Graf, 2002 [Publ 2651], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	3	2	--

-- Lens_contradens **

Hoeh et al., 2009 [Publ 2716], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	3	2	--

-- Lens_contradens **

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