MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Sun 12 April 2026
Bivalvia | Unionida | Etherioidea
family ETHERIIDAE Deshayes, 1832: cladomics
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 4 (4 total)
number of subtaxa (rank: genus) = 4
analyzed in 38 cladograms in 25 publications
maximum number of terminals in an analysis = 3 (mean = 1.37, median = 1)
maximum number of subtaxa (genus) = 3 (mean = 1.37, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 8
maximum number of included terminals = 3, (mean = 2.75 median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.75, median = 3)
proportion of cladograms that support basic monophyly = 0.25
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 8
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 3 (mean = 2.75, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.75, median = 3)
proportion of cladograms that support rigorous monophyly = 0.25
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 630 | 708 | 3 | 629 | 18 | 2 | 0 |
+----------------------------- Iridininae
|
+-----------------------------------------------------------------+ +-------------------------- Aspathariinae
| | |
| +--+ +----------------------- Etheria **
| | |
| +--+ +-------------------- Anodontites_trapesialis
| | |
| +--+ +----------------- Mulleria **
| | |
| +--+ +-------------- Lamproscapha
| | |
| +--+ +-- Monocondylaea
| | +--------+
| | | +-- Tamsiella
| +--+
| | +-------- Mycetopodinae
| | |
| +--+ +-- Anodontites_guanarensis
| | +--+
| | | +-- Fossula
| +--+
| | +-- Anodontites_trigona
| +--+
| +-- Anodontites_elongata
|
| +-------------------------------------------------------------------------------------------- Hyriidae
| |
| | +-------------------------------- Lamellidens
| | |
| | +-----+ +-- Balwantia
| | | | +--------------------------+
--+ | | | | +-- Trapezidens
| | | +--+
| | | | +-------------------------- Leoparreysiini
| | | | |
| | | +--+ +----------------------- Pseudomulleria **
| | | | |
| | | | | +-- Parreysiini
| | | +--+ +-----------------+
| | | | | +-- Coelaturini
| | | | |
| | | +--+ +-- Pseudoscabies
| | | | +--------------+
| | | | | +-- Indonaia
| | | +--+
| | | | +-------------- Indochinella
| | | | |
| | | +--+ +----------- Radiatula
| | +--------------------------------------------------+ | |
| | | | +--+ +-- Songkhlanaia_songkhramensis__2
| | | | | +-----+
| | | | | | +-- Unionetta_fabagina__2
| | | | +--+
| | | | | +----- Harmandia
| | | | +--+
| | | | | +-- Unionetta_fabagina__1
| | | | +--+
| | | | +-- Scabies
| | | |
| | | | +----- Amuranodonta
| | | | +-----------------------------+
| | | | | | +-- Anemina
+--+ | | | +--+
| | | | +-- Buldowskia
| | | |
| | | | +-- Beringiana
| | | | +-----+
| | | | | +-- Pletholophus
| | +--+ +-----------------------+
| | | | | +----- Mimo
| | | | +--+
| | | | | +-- Acudonta
| | | | +--+
| | | | +-- Sinanodonta
| | | |
| | | | +----------------------------- Cristaria
| | +--+ |
| | | | +----------------- Lepidodesmini
| | | | |
| | | | | +----------- Lanceolariini
| | | | | +--+
| | | | +--------+ | | +-------- Pseudoacuticosta_retiaria__2
| | | | | | | +--+
| | | | | | | | +----- Acuticosta_chinensis__2
| | +--+ | | | +--+
| | | | +--+ | +-- Acuticosta_chinensis__1
| | | | | +--+
| | | | | +-- Pseudoacuticosta_retiaria__1
| | | | |
| | | | | +----------- Globunio
| | | | +--+
| | | | | +-------- Aculamprotulina
| | | | +--+
| | +--+ | +----- Middendorffinaiina
| | | +--+
| | | | +-- Unionina
| | | +--+
| | | +-- Oxynaiina
| | |
| | | +-------- Pseudodontoideus
+--+ | +--------------+
| | | | +----- Lasmigona
| | | +--+
| | | | +-- Pterosyna
| | | +--+
| +--+ +-- Arcidens
| |
| | +-------------------- Anodontina
| | |
| | | +-------------- Utterbackia
| | | +--+
| +--+ | | +----------- Pyganodon_grandis
| | | +--+
| | | | +-------- Pyganodon_lacustris
| | | +--+
| | | | +----- Pyganodon_cataracta__2
| +--+ +--+
| | | +-- Pyganodon_cataracta__1
| | +--+
| | +-- Potamilus_fragilis__3
| |
| | +-------------- Utterbackiana
| | |
| +--+ +----------- Simpsonaias
| | |
| | | +-- Alasminota
| +--+ +-----+
| | | +-- Alasmidonta
| | |
| +--+ +-- Prolasmidonta
| | +--+
| | | +-- Platynaias
| +--+
| | +-- Anodontoides
| +--+
| +-- Strophitus
|
| +-------------------------------------------------------------------------------------- Rectidentini
| |
| | +----------------------------------------------------------------------------------- Margaritiferidae
| | |
| | | +-------------------- Chamberlainiini
| | | |
| | | +--+ +----------------- Parvasolenaia
| | | | | |
| | | | +--+ +-------------- Pseudopostolata
| | | | | |
| | | | +--+ +----------- Koreosolenaia
| | | | | |
| | | | +--+ +-- Cosmopseudodon
| | | | | +-----+
| | | | | | +-- Guiunio
+--+ | | +--+
| | +--------+ | +----- Ptychorhynchus
| | | | +--+
| | | | | +-- Obovalis
| | | | +--+
| | | | +-- Postolata
| | | |
| | | | +-- Caudiculatus
| | | | +--------------+
| | | | | +-- Lamprotulini
| | | | |
| | | +-----+ +-------------- Sinosolenaia_carinata
| | | | |
| | | | | +----- Gonidea
| | | +--+ +-----+
+--+ +-----------------------------------------------+ | | | +-- Microcondylaea
| | | | | +--+
| | | +--+ +-- Leguminaia
| | | |
| | | | +-------- Inversidens
| | | +--+
| | | | +----- Sinosolenaia_recognita
| | | +--+
| | | | +-- Sinosolenaia_iridinea
| | | +--+
| | | +-- Sinosolenaia_oleivora
| | |
| | | +----------------------------- Indopseudodontina
| | | |
| | | | +-- Lannanaia
| | +--+ +-----------------------+
| | | | +-- Pilsbryoconcha
| | | |
| | | | +----- Songkhlanaia_tamodienica
| | +--+ +-----------------+
| | | | | +-- Songkhlanaia_moreleti
| | | | +--+
| | | | +-- Songkhlanaia_songkhramensis__1
| | | |
| | +--+ +-- Nyeinchanconcha
| | | +-----------------+
| | | | +-- Sundadontina
| | | |
+--+ | | +-- Isannaia
| +--+ +--+
| | | +-- Namkongnaia
| | +-----------+
| | | | +-- Thaiconcha
| | | +--+
| +--+ +-- Bineurus
| |
| | +-------------- Pseudodon_walpolei
| | |
| +--+ +----------- Pseudodon_vagulus
| | |
| | | +----- Pseudodon_vondembuschianus
| +--+ +--+
| | | | +-- Pseudodon_mekongi
| | | +--+
| +--+ +-- Schepmaniini
| |
| | +----- Pseudodon_lenyanensis__2
| +--+
| | +-- Pseudodon_inoscularis
| +--+
| +-- Pseudodon_lenyanensis__1
|
| +-- Quadrulini
| +--------------------------------------------------------------------------+
| | +-- Pleurobemini
| |
| | +----- Amblema
+--+ +--------------------------------------------------------------------+
| | | +-- Nephronaias
| | +--+
| | +-- Psoronaias
| |
+--+ +----------------------------------------------------------------------- Reginaia
| |
| | +----- Popenaias
| | +--------------------------------------------------------------+
| | | | +-- Cyrtonaias
+--+ | +--+
| | +-- Glebula
| |
| | +----------------------------------------------------------------- Paetulunio
| | |
+--+ | +----------------- Toxolasma_parvum
| | |
| | | +----- Toxolasma_lividum__2
| | | +--------+
| | +--------------------------------------------+ | | +-- Toxolasma_cylindrellus
| | | | | +--+
| | | | | +-- Toxolasma_lividum__1
| | | | |
+--+ | +--+ +-------- Toxolasma_texasiense__2
| | | +--+
| | | | | +----- Toxolasma_glans
| | | | +--+
| | | | | +-- Toxolasma_paulum
| | +--+ +--+
| | | +-- Toxolasma_texasiense__1
| | |
| | | +-------- Toxolasma__4
| | +--+
+--+ | +----- Toxolasma_granulatum
| +--+
| | +-- Toxolasma__2
| +--+
| +-- Toxolasma__3
|
| +----- Toxolasma_pullus
| +-----------------------------------------------------+
| | | +-- Toxolasma_germanum
| | +--+
| | +-- Toxolasma__1
| |
| | +-- Obliquaria
| | +-----------------------------------------------------+
| | | +-- Plectomerus
+--+ |
| | +-------- Potamilus_inflatus
| | +--------+
| | | | +----- Potamilus_ohiensis
| | | +--+
| | | | +-- Potamilus_amphichaenus
| | | +--+
| | +-----------------------------------+ +-- Potamilus_streckersoni
| | | |
+--+ | | +----- Potamilus_fragilis__2
| | | +--------+
| | | | | +-- Potamilus_leptodon
| | | | +--+
| | +--+ +-- Potamilus_fragilis__1
| | |
| | | +----------- Potamilus_metnecktayi
| | | |
| | +--+ +-------- Potamilus_alatus__3
| | | |
| | +--+ +----- Potamilus_alatus__2
+--+ | |
| +--+----- Potamilus_purpuratus__2
| |
| | +-- Potamilus_alatus__1
| +--+
| +-- Potamilus_purpuratus__1
|
| +-- Villosa_villosa
| +-----------------------------------------------+
| | +-- Atlanticoncha
| |
| | +-- Cambarunio_nebulosus
| | +--------------------------------------------+
+--+ | +-- Ligumia
| |
| | +-------- Leaunio_umbrans
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus
+--+ | +--+
| | | +-- Cambarunio_iris
| | +--+
| | +-- Cambarunio_simus
| |
| | +----------------------------------------- Hamiota
| | |
+--+ | +----------------------- Pachynaias
| | |
| | +--------------+ +-------------------- Sagittunio
| | | | |
| | | | | +----- Ellipsaria
| | | +--+ +-----------+
| | | | | | +-- Potamilus_capax
| | | | | +--+
| | | | | +-- Truncilla
+--+ | +--+
| | | +----- Medionidus_acutissimus
| | | +--------+
| | | | | +-- Lemiox
| | | | +--+
| | +--+ +-- Medionidus_penicillatus
| | |
| | | +----------- Ptychobranchus
| | | |
| | +--+ +-- Cyprogenia_aberti__2
| | | +-----+
+--+ | | +-- Dromus
| +--+
| | +----- Cyprogenia_stegaria__2
| +--+
| | +-- Cyprogenia_aberti__1
| +--+
| +-- Cyprogenia_stegaria__1
|
| +-- Villosa_delumbis
| +--------------------------------+
| | +-- Villosa_vibex
| |
| | +-------------- Obovaria_olivaria
| | |
| | +-----------------+ +-- Venustaconcha
| | | | +--------+
| | | | | +-- Epioblasma
| | | +--+
| | | | +-------- Obovaria_retusa
+--+ | +--+
| | | +----- Obovaria_choctawensis
| | +--+
| | | +-- Obovaria_unicolor
| | +--+
| | +-- Obovaria_subrotunda
| |
| | +-- Mesonaias
| | +-----------------------+
| | | +-- Obovaria_arkansasensis
| | |
+--+ +--+ +-- Lampsilis_splendida
| | | +--------------------+
| | | | +-- Lampsilis_bracteata
| | | |
| | +--+ +-- Lampsilis_fasciola
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----------------- Lampsilis_ornata
| | | |
| | +--+ +-------------- Lampsilis_ovata__2
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis
| | | |
| | | | +----- Lampsilis_cardium__2
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1
+--+ | | +--+
| +--+ +-- Lampsilis_satura
| |
| | +----- Lampsilis_cariosa__2
| +--+
| | +-- Lampsilis_ovata__1
| +--+
| +-- Lampsilis_cariosa__1
|
| +----------- Ortmanniana_ligamentina__2
| +--------------+
| | | +-------- Lampsilis_abrupta
| | +--+
| | | +----- Lampsilis_higginsii__2
| | +--+
| | | +-- Ortmanniana_ligamentina__1
| | +--+
+--+ +-- Lampsilis_higginsii__1
|
| +----- Medionidus_conradicus
| +-----------------+
| | | +-- Leaunio_lienosus
| | +--+
| | +-- Leaunio_vanuxemensis
| |
+--+ +-- Lampsilis_floridensis
| +-----------------+
| | +-- Lampsilis_teres
| |
| | +-- Lampsilis_siliquoidea__1
+--+ |
| +-- Lampsilis_siliquoidea__2
| +--------------+
| | +-- Lampsilis_powellii__1
| | |
+--+ +-- Lampsilis_powellii__2
|
| +-------------- Lampsilis_sietmani
| |
+--+ +-- Lampsilis_straminea__2
| +--------+
| | +-- Lampsilis_virescens
+--+
| +-------- Lampsilis_hydiana__2
+--+
| +----- Lampsilis_hydiana__1
+--+
| +-- Lampsilis_bergmanni
+--+
+-- Lampsilis_straminea__1
Hoeh et al., 2009 [Publ 2716], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 10 | 8 | 3 | 43 | 7 | 2 | 742 |
+----------- Pseudomulleria ** --+ | +-------- Mulleria ** +--+ | +----- Etheria ** +--+ | +-- Mycetopodidae +--+ +-- Iridinidae
Graf & Cummings, 2006 [Publ 1968], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 44 | 7 | 2 | 745 |
+-- Mycetopodidae
+-----+
| +-- Iridinidae
--+
| +----- Etheria **
+--+
| +-- Mulleria **
+--+
+-- Pseudomulleria **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 28 | 21 | 3 | 31 | 5 | 2 | 731 |
+-------- Iridinidae
+--------+
| | +----- Etheria **
| +--+
| | +-- Mulleria **
--+ +--+
| +-- Mycetopodidae
|
| +-------------- Parreysiinae
| |
+--+ +----------- Pseudomulleria **
| |
+--+ +-------- Anodontini
| |
+--+ +----- Gonideinae
| |
+--+----- Unionini
|
| +-- Margaritiferidae
+--+
+-- Ambleminae
Hoeh et al., 2009 [Publ 2716], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 37 | 34 | 3 | 44 | 7 | 2 | 737 |
+-------- Iridinidae
+--------------+
| | +----- Etheria **
| +--+
| | +-- Mulleria **
--+ +--+
| +-- Mycetopodidae
|
| +-------------------- Margaritiferidae
+--+
| +----------------- Parreysiinae
+--+
| +-------------- Pseudomulleria **
+--+
| +----------- Unioninae
+--+
| +-------- Rectidentini
+--+
| +----- Ambleminae
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Hoeh et al., 2009 [Publ 2716], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 36 | 33 | 3 | 43 | 7 | 2 | 745 |
+-- Etheria **
+--+
| +-- Mulleria **
+-----------------+
| | +-- Mycetopodidae
| +--+
--+ +-- Iridinidae
|
| +-------------------- Parreysiinae
+--+
| +----------------- Pseudomulleria **
+--+
| +-------------- Margaritiferidae
+--+
| +----------- Pseudodontini
+--+
| +-------- Rectidentini
+--+
| +----- Unioninae
+--+
| +-- Gonideini
+--+
+-- Ambleminae
Walker et al., 2006 [Publ 2666], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 7 | 0 | 2 | 34 | 7 | 2 | 741 |
+----------- Iridininae --+ | +-------- Etheria ** +--+ | +----- Aspathariinae +--+ | +-- Mulleria ** +--+ +-- Mycetopodidae
Graf, 2000 [Publ 2647], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 15 | 3 | 2 | -- |
+-------- Iridinidae | --+ +----- Anodontitinae | | +--+----- Mycetopodinae | | +-- Etheria ** +--+ +-- Mulleria **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 121 | 4 | 2 | -- |
+----- Mycetopodidae --+ | +-- Etheria ** +--+ +-- Iridinidae
Bolotov et al., 2016 [Publ 3743], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 29 | 3 | 2 | -- |
+----- Mycetopodidae --+ | +-- Etheria ** +--+ +-- Iridinidae
Combosch et al., 2017 [Publ 3806], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 14 | 5 | 2 | -- |
+-------- Aspathariinae --+ | +----- Etheria ** +--+ | +-- Iridininae +--+ +-- Mycetopodidae
Graf et al., 2015 [Publ 3619], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 19 | 2 | 2 | -- |
+----- Iridinidae --+ | +-- Mycetopodidae +--+ +-- Etheria **
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 27 | 5 | 2 | -- |
+-------- Iridininae --+ | +----- Aspathariinae +--+ | +-- Etheria ** +--+ +-- Mycetopodidae
Huang et al., 2018 [Publ 3828], fig. 3_5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 28 | 5 | 2 | -- |
+-------- Iridininae --+ | +----- Aspathariinae +--+ | +-- Etheria ** +--+ +-- Mycetopodidae
Bolotov et al., 2017 [Publ 3761], fig. S2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 137 | 3 | 2 | -- |
+----- Iridinidae --+ | +-- Mycetopodidae +--+ +-- Etheria **
Wu et al., 2020 [Publ 4121], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 33 | 5 | 2 | -- |
+-------- Iridininae --+ | +----- Aspathariinae +--+ | +-- Etheria ** +--+ +-- Mycetopodidae
Bolotov et al., 2017 [Publ 3761], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 137 | 3 | 2 | -- |
+----- Mycetopodidae --+ | +-- Etheria ** +--+ +-- Iridinidae
Combosch et al., 2017 [Publ 3806], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 14 | 5 | 2 | -- |
+-------- Aspathariinae --+ | +----- Etheria ** +--+ | +-- Iridininae +--+ +-- Mycetopodidae
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 56 | 2 | 2 | -- |
+----- Mycetopodidae --+ | +-- Etheria ** +--+ +-- Iridinidae
Chen et al., 2025 [Publ 4455], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 25 | 4 | 2 | -- |
+----- Iridinidae --+ | +-- Etheria ** +--+ +-- Mycetopodidae
Huang et al., 2019 [Publ 3843], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 199 | 9 | 2 | -- |
+-- Anodontites
+--------+
| +-- Iridininae
|
--+ +-- Etheria **
| +-----+
| | +-- Aspathariinae
+--+
| +----- Mycetopodinae
+--+
| +-- Monocondylaeinae
+--+
+-- Lamproscapha
Sano et al., 2020 [Publ 4138], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 67 | 5 | 2 | -- |
+-------- Iridininae --+ | +----- Aspathariinae +--+ | +-- Etheria ** +--+ +-- Mycetopodidae
Elderkin et al., 2016 [Publ 3693], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 2 | 1 | 1 | -- |
-- Etheria_elliptica **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 56 | 2 | 2 | -- |
+----- Mycetopodidae --+ | +-- Etheria ** +--+ +-- Iridinidae
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 79 | 5 | 2 | -- |
+-------- Iridininae --+ | +----- Aspathariinae +--+ | +-- Etheria ** +--+ +-- Mycetopodidae
Wu et al., 2021 [Publ 4366], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 90 | 4 | 2 | -- |
+----- Iridinidae --+ | +-- Etheria ** +--+ +-- Mycetopodidae
Konopleva et al., 2017 [Publ 3807], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 2 | 2 | -- |
+----- Mycetopodidae --+ | +-- Etheria ** +--+ +-- Iridinidae
Wu et al., 2020 [Publ 4121], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 33 | 5 | 2 | -- |
+-------- Iridininae --+ | +----- Aspathariinae +--+ | +-- Etheria ** +--+ +-- Mycetopodidae
Huang et al., 2019 [Publ 3843], fig. S3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 197 | 9 | 2 | -- |
+-- Anodontites
+--------+
| +-- Iridininae
|
--+ +-- Etheria **
| +-----+
| | +-- Aspathariinae
+--+
| +----- Mycetopodinae
+--+
| +-- Monocondylaeinae
+--+
+-- Lamproscapha
Sano et al., 2020 [Publ 4138], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 67 | 5 | 2 | -- |
+-------- Iridininae --+ | +----- Aspathariinae +--+ | +-- Etheria ** +--+ +-- Mycetopodidae
Sano et al., 2020 [Publ 4138], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 67 | 5 | 2 | -- |
+-------- Iridininae --+ | +----- Aspathariinae +--+ | +-- Etheria ** +--+ +-- Mycetopodidae
Smith, 2021 [Publ 4331], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- |
+-- Etheria ** --+ +-- Iridinidae
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 27 | 5 | 2 | -- |
+-------- Iridininae --+ | +----- Aspathariinae +--+ | +-- Etheria ** +--+ +-- Mycetopodidae
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 39 | 2 | 2 | -- |
+----- Etheria ** --+ | +-- Mycetopodidae +--+ +-- Iridinidae
Huang et al., 2019 [Publ 3843], fig. S4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 199 | 9 | 2 | -- |
+-------- Etheria ** | +-------- Aspathariinae | --+ +-- Anodontites +-----+ | +-- Iridininae | | +----- Mycetopodinae +--+ | +-- Monocondylaeinae +--+ +-- Lamproscapha
Bolotov et al., 2016 [Publ 3743], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 29 | 3 | 2 | -- |
+----- Mycetopodidae --+ | +-- Etheria ** +--+ +-- Iridinidae
Lemer et al., 2019 [Publ 4093], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- |
+-- Etheria ** --+ +-- Iridinidae
Wu et al., 2021 [Publ 4366], fig. 2_I
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 90 | 4 | 2 | -- |
+----- Iridinidae --+ | +-- Etheria ** +--+ +-- Mycetopodidae
