Bivalvia | Unionida | Unionoidea | Margaritiferidae

subfamily GIBBOSULINAE Bogan, Bolotov, Froufe & Lopes-Lima in Lopes-Lima et al., 2018: cladomics

1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (5 total)
number of subtaxa (rank: genus) = 1
analyzed in 88 cladograms in 52 publications
maximum number of terminals in an analysis = 4 (mean = 1.4, median = 1)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 25
maximum number of included terminals = 4, (mean = 2.4 median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 0.84

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Gibbosulinae cladogram distances
PCoA Ordination

NJ Tree of Gibbosulinae cladogram distances
NJ Tree

Cladograms

Bolotov et al., 2016 [Publ 3732], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 1 1 **
Cladomics cladogram Gibbosula MARGARITIFERINAE

Zhou et al., 2007 [Publ 2658], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 20 2 1 0.0
Cladomics cladogram Gibbosula GONIDEINAE

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 29 9 1 0.0
Cladomics cladogram Pseudunio Margaritifera Gibbosula Cumberlandia

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 29 9 1 0.0
Cladomics cladogram Pseudunio Margaritifera Gibbosula Cumberlandia

Araujo et al., 2017 [Publ 3746], fig. 1_2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 10 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Araujo et al., 2017 [Publ 3746], fig. 2a

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 10 10 1 0.0
Cladomics cladogram Cumberlandia Pseudunio Gibbosula Margaritifera

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 138 9 1 0.0
Cladomics cladogram Cumberlandia Margaritifera Gibbosula Pseudunio

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 138 9 1 0.0
Cladomics cladogram Cumberlandia Gibbosula Pseudunio Margaritifera

Vikhrev et al., 2017 [Publ 3810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Bespalaya et al., 2018 [Publ 3811], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Vikhrev et al., 2018 [Publ 3812], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 10 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 27 10 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 25 11 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 25 11 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Huang et al., 2019 [Publ 3843], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 1 1 0.0
Cladomics cladogram MARGARITIFERINAE Gibbosula

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.714 2 13 0 1 199 11 1 0.0
Cladomics cladogram Gibbosula rochechouartii Gibbosula laosensis MARGARITIFERINAE PARREYSIINAE UNIONINAE RECTIDENTINI AMBLEMINAE CHAMBERLAINIINI LAMPROTULINI GONIDEINI PSEUDODONTINI

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Huang et al., 2019 [Publ 3843], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.714 2 13 0 1 197 11 1 0.0
Cladomics cladogram Gibbosula rochechouartii Gibbosula laosensis MARGARITIFERINAE PARREYSIINAE PSEUDODONTINI RECTIDENTINI UNIONINAE CHAMBERLAINIINI AMBLEMINAE LAMPROTULINI GONIDEINI

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.714 2 13 0 1 199 11 1 0.0
Cladomics cladogram Gibbosula rochechouartii Gibbosula laosensis MARGARITIFERINAE PARREYSIINAE Chamberlainia Megalonaias Uniomerus QUADRULINA Sinohyriopsis LAMPROTULINI GONIDEINI PSEUDODONTINI RECTIDENTINI UNIONINAE Amblema Toxolasma PLEUROBEMINI Obliquaria Plectomerus Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 3 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Konopleva et al., 2019 [Publ 3900], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 50 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 8 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Lyubas et al., 2019 [Publ 3963], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 13 13 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 6 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 6 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Bolotov et al., 2020 [Publ 4100], fig. 1

cst iT tT bi riS oT roT roS iD
0.600 1 1 0 1 116 4 1 0.0
Cladomics cladogram Gibbosula Margaritifera dahurica Margaritifera margaritifera Margaritifera laevis Margaritifera middendorffi

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 53 2 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 1 32 10 1 0.0
Cladomics cladogram Cumberlandia Margaritifera Gibbosula Pseudunio

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 3 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
0.714 2 12 0 1 32 10 1 0.0
Cladomics cladogram Gibbosula rochechouartii Gibbosula laosensis MARGARITIFERINAE AMBLEMINAE PARREYSIINAE GONIDEINAE UNIONINAE

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 2 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 1 67 10 1 0.0
Cladomics cladogram Cumberlandia Margaritifera Gibbosula Pseudunio

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 1 67 10 1 0.0
Cladomics cladogram Cumberlandia Pseudunio Gibbosula Margaritifera

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 1 67 10 1 0.0
Cladomics cladogram Cumberlandia Pseudunio Gibbosula Margaritifera

Wu et al., 2020 [Publ 4147], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 2 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 6 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 71 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 71 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 66 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 6 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Bolotov et al., 2022 [Publ 4250], fig. 8

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 48 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE Sinosolenaia Leguminaia Gonidea Lamprotula Potomida

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 75 9 1 0.0
Cladomics cladogram Cumberlandia Gibbosula Pseudunio Margaritifera

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 75 9 1 0.0
Cladomics cladogram Cumberlandia Margaritifera Gibbosula Pseudunio

Dai et al., 2023 [Publ 4285], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2023 [Publ 4285], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Bogan et al., 2023 [Publ 4295], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 93 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 93 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 19 11 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 19 11 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2024 [Publ 4385], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2024 [Publ 4385], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 0.0
Cladomics cladogram MARGARITIFERINAE Gibbosula

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 0.0
Cladomics cladogram MARGARITIFERINAE Gibbosula

Dai et al., 2024 [Publ 4389], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 1 1 0.0
Cladomics cladogram MARGARITIFERINAE Gibbosula

Kashiri et al., 2024 [Publ 4822], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2025 [Publ 4441], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 5 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Chen et al., 2025 [Publ 4455], fig. 1

cst iT tT bi riS oT roT roS iD
0.714 4 4 2 1 22 11 1 0.0
Cladomics cladogram Pseudunio marocanus Pseudunio auricularius 1 Pseudunio auricularius 2 Pseudunio homsensis Gibbosula Cumberlandia Margaritifera

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 0.0
Cladomics cladogram MARGARITIFERINAE Gibbosula

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 0.0
Cladomics cladogram MARGARITIFERINAE Gibbosula

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 63 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Chen et al., 2025 [Publ 4663], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Dai et al., 2026 [Publ 4757], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 137 11 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.857 3 3 1 1 630 11 1 0.0
Cladomics cladogram Pseudunio Gibbosula Cumberlandia Margaritifera

Wu et al., 2026 [Publ 4811], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 1 1 0.0
Cladomics cladogram MARGARITIFERINAE Gibbosula

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 64 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Wu et al., 2026 [Publ 4852], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 1 1 0.0
Cladomics cladogram Gibbosula MARGARITIFERINAE

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 0.0
Cladomics cladogram Gibbosula AMBLEMINAE

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 0 0 0.0
Cladomics cladogram Gibbosula PARREYSIINAE UNIONINAE AMBLEMINAE GONIDEINAE

Bogan et al., 2023 [Publ 4295], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 0 0 0.0
Cladomics cladogram Gibbosula UNIONINAE

Bogan et al., 2023 [Publ 4295], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 0.0
Cladomics cladogram Gibbosula UNIONINAE

Bogan et al., 2023 [Publ 4295], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 0.0
Cladomics cladogram Gibbosula UNIONINAE

Chen et al., 2026 [Publ 4827], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 8 0 0 0.0
Cladomics cladogram Gibbosula UNIONINAE

Chen et al., 2026 [Publ 4833], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 25 0 0 0.0
Cladomics cladogram Gibbosula Lamellidens Trapezidens PARREYSIINI LEOPARREYSIINI COELATURINI Indochinella Indonaia Radiatula GONIDEINAE Harmandia Chenunionetta Unionetta fabagina 2 Unionetta fabagina 1 Scabies