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Data last updated
Thu 18 Dec 2025

Bivalvia | Unionoida | Hyrioidea | Hyriidae | Hyriinae

tribe HYRIDELLINI McMichael, 1956 (1934): cladomics

4 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 9 (13 total)
number of subtaxa (rank: genus) = 4
analyzed in 45 cladograms in 27 publications
maximum number of terminals in an analysis = 4 (mean = 1.53, median = 1)
maximum number of subtaxa (genus) = 2 (mean = 1.09, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 15
maximum number of included terminals = 4, (mean = 2.6 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.27, median = 1)
proportion of cladograms that support basic monophyly = 0.6

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 4
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 4 (mean = 3.5, median = 4)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.5

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyridellini cladogram distances
PCoA Ordination

NJ Tree of Hyridellini cladogram distances
NJ Tree

Cladograms

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 19 20 2 16 9 4 0 
                 +----- Diplodon_suavidicus__1
  +--------------+                            
  |              |  +-- Hyriini               
  |              +--+                         
  |                 +-- Castaliini            
  |                                           
  |           +-------- Westralunio           
  |           |                               
  +-----------+     +-- Hyridella_drapeta **     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa **    
  |           +--+                            
  |              |  +-- Diplodon_gratus       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2
  |                                           
  |                 +-- Hyridella_australis **   
  |  +--------------+                         
  |  |              +-- Cucumerunio **           
  +--+                                        
     |  +-------------- Echyridella           
     +--+                                     
        |  +----------- Velesunio_ambiguus    
        +--+                                  
           |  +-------- Microdontia           
           +--+                               
              |  +----- Lortiella             
              +--+                            
                 |  +-- Alathyria             
                 +--+                         
                    +-- Velesunio_angasi

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
4 13 11 2 16 9 4 16 
                    +----- Diplodon_suavidicus__2
  +-----------------+                            
  |                 |  +-- Hyriini               
  |                 +--+                         
  |                    +-- Castaliini            
  |                                              
  |              +-------- Westralunio           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta **     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa **    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1
  +--+              +--+                         
     |                 +-- Diplodon_gratus       
     |                                           
     |  +----------------- Echyridella           
     |  |                                        
     +--+              +-- Cucumerunio **           
        |  +-----------+                         
        |  |           +-- Hyridella_australis **   
        +--+                                     
           |  +----------- Velesunio_ambiguus    
           +--+                                  
              |  +-------- Alathyria             
              +--+                               
                 |  +----- Velesunio_angasi      
                 +--+                            
                    |  +-- Lortiella             
                    +--+                         
                       +-- Microdontia

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 2 16 3 3 19 
     +----- Rhipidodontini     
  +--+                         
  |  |  +-- Hyriini            
  |  +--+                      
  |     +-- Castaliini         
--+                            
  |     +-- Hyridella_depressa ** 
  |  +--+                      
  |  |  +-- Hyridella_drapeta **  
  +--+                         
     |  +-- Hyridella_australis **
     +--+                      
        +-- Cucumerunio **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3 3 1 1 196 4 4 20 
  +----------- Echyridella   
--+                          
  |  +-------- Hyridella **     
  +--+                       
     |  +----- Rhipidodontini
     +--+                    
        |  +-- Castaliini    
        +--+                 
           +-- Hyriini

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3 3 1 1 198 4 4 20 
1

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 198 4 4 20 
  +----------- Echyridella   
--+                          
  |  +-------- Hyridella **     
  +--+                       
     |  +----- Rhipidodontini
     +--+                    
        |  +-- Castaliini    
        +--+                 
           +-- Hyriini

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
2 7 0 1 6 3 3 23 
           +-- Hyridella_australis **
  +--------+                      
  |        +-- Rhipidodontini     
--+                               
  |  +-------- Hyridella_depressa ** 
  +--+                            
     |  +----- Velesunio          
     +--+                         
        |  +-- Echyridella        
        +--+                      
           +-- Castaliini

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 1 44 4 3 -- 
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 45 4 3 -- 
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 44 4 3 -- 
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 12 5 2 -- 
  +----- Rhipidodontini
--+                    
  |  +-- Cucumerunio **   
  +--+                 
     +-- Hyridella **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
2 3 0 1 9 3 3 23 
           +-- Rhipidodontini        
  +--------+                         
  |        +-- Castaliini            
--+                                  
  |  +-------- Hyridella_australis__2 **
  +--+                               
     |  +----- Hyridella_australis__1 **
     +--+                            
        |  +-- Echyridella           
        +--+                         
           +-- Hyridella_depressa **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 45 4 3 -- 
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
2 8 0 1 34 3 3 20.5 
  +----------------- Castaliini         
  |                                     
  +----------------- Rhipidodontini     
--+                                     
  |  +-------------- Hyridella_depressa ** 
  |  |                                  
  |  +-------------- Hyridella_australis **
  +--+                                  
     |  +----------- Echyridella        
     |  |                               
     |  |        +-- Velesunio_ambiguus 
     +--+  +-----+                      
        |  |     +-- Velesunio__3       
        |  |                            
        +--+  +----- Velesunio__2       
           |  |                         
           |  |  +-- Alathyria          
           +--+--+                      
              |  +-- Velesunio__1       
              |                         
              |  +-- Velesunio_angasi   
              +--+                      
                 +-- Lortiella

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
2 3 0 1 7 3 3 22 
  +----- Hyridella_depressa ** 
--+                         
  |  +-- Hyridella_australis **
  +--+                      
     +-- Rhipidodontini

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 2 2 -- 
  +----- Hyridella **     
--+                    
  |  +-- Rhipidodontini
  +--+                 
     +-- Castaliini

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 28 1 1 -- 
-- Hyridella_australis **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 27 1 1 -- 
-- Hyridella_australis **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Hyridella_australis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Hyridella_australis **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
-- Hyridella_australis **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
  +-- Hyriini  
--+            
  +-- Hyridella **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 67 1 1 -- 
-- Hyridella_australis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 0 0 -- 
-- Hyridella_depressa **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 0 0 -- 
-- Hyridella_depressa **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 27 1 1 -- 
-- Hyridella_australis **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Hyridella_australis **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 2 2 -- 
-- Hyridella **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
-- Hyridella_australis **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 67 1 1 -- 
-- Hyridella_australis **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
-- Hyridella_australis **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 67 1 1 -- 
-- Hyridella_australis **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Hyridella_depressa **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Hyridella_australis **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Hyridella_depressa **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Hyridella_australis **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 2 2 -- 
  +----- Hyridella ** 
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
  +----- Hyridella ** 
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 25 1 1 -- 
  +-- Hyridella **
--+            
  +-- Hyriini

Klunzinger et al., 2022 [Publ 4259], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Cucumerunio_novaehollandiae **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
  +----- Hyridella ** 
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Klunzinger et al., 2021 [Publ 4086], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Cucumerunio_novaehollandiae **
 
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