Bivalvia | Unionida | Hyrioidea | Hyriidae | Hyriinae

tribe HYRIDELLINI McMichael, 1956 (1934): cladomics

4 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 9 (13 total)
number of subtaxa (rank: genus) = 4
analyzed in 48 cladograms in 30 publications
maximum number of terminals in an analysis = 4 (mean = 1.65, median = 1)
maximum number of subtaxa (genus) = 2 (mean = 1.15, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 18
maximum number of included terminals = 4, (mean = 2.72 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.39, median = 1)
proportion of cladograms that support basic monophyly = 0.61

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 7
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 4 (mean = 3.43, median = 4)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.57


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyridellini cladogram distances
PCoA Ordination

NJ Tree of Hyridellini cladogram distances
NJ Tree


Cladograms

Prié et al., 2026 [Publ 4844], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.812 2 2 0 2 21 13 4 **
Cladomics cladogram Echyridella Westralunio Hyridella Cucumerunio Velesunio Alathyria Lortiella Microdontia Diplodon Guyanhyrida CASTALIINI Iridea HYRIINI

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

cst iT tT bi riS oT roT roS iD
0.667 4 20 20 2 17 10 4 0.0
Cladomics cladogram Neotrigonia Guyanhyrida HYRIINI CASTALIINI Westralunio Diplodon Hyridella drapeta Hyridella depressa Hyridella australis Cucumerunio Echyridella Velesunio ambiguus Microdontia Lortiella Alathyria Velesunio angasi

Santos-Neto et al., 2016 [Publ 3739], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 4 13 11 2 17 10 4 0.0
Cladomics cladogram Guyanhyrida HYRIINI CASTALIINI Westralunio Diplodon Hyridella drapeta Hyridella depressa Echyridella Cucumerunio Hyridella australis Velesunio ambiguus Alathyria Velesunio angasi Lortiella Microdontia

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.667 4 18 20 2 629 15 4 0.0
Cladomics cladogram Guyanhyrida HYRIINI CASTALIINI Diplodon Hyridella depressa Hyridella drapeta Cucumerunio Hyridella australis Echyridella Velesunio 1 Alathyria jacksoni 1 Velesunio ambiguus Alathyria profuga Velesunio 3 Microdontia Lortiella Velesunio angasi Alathyria pertexta 2 Velesunio 2 Alathyria jacksoni 2 Alathyria pertexta 1

Graf et al., 2015 [Publ 3619], fig. 1

cst iT tT bi riS oT roT roS iD
0.900 4 4 2 2 16 3 3 0.0
Cladomics cladogram RHIPIDODONTINI HYRIINI CASTALIINI Hyridella depressa Hyridella drapeta Hyridella australis Cucumerunio

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 2 2 0 2 12 5 2 0.0
Cladomics cladogram RHIPIDODONTINI Cucumerunio Hyridella

Lawrie et al., 2026 [Publ 4843], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 4 4 2 2 14 2 2 0.0
Cladomics cladogram CASTALIINI HYRIINI Hyridella depressa Hyridella drapeta Hyridella australis Cucumerunio Westralunio Velesunio 6 Alathyria jacksoni 2 Alathyria profuga Velesunio ambiguus Velesunio 5 Alathyria pertexta 2 Velesunio 4 Alathyria jacksoni 1 Alathyria pertexta 1 Lortiella Velesunio 3 Microdontia Velesunio angasi Velesunio 1 Velesunio 2

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 198 4 4 0.0
Cladomics cladogram Echyridella Hyridella RHIPIDODONTINI CASTALIINI HYRIINI

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 196 4 4 0.0
Cladomics cladogram Echyridella Hyridella RHIPIDODONTINI CASTALIINI HYRIINI

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.900 3 3 1 1 198 4 4 0.0
Cladomics cladogram Echyridella Westralunio Hyridella RHIPIDODONTINI CASTALIINI HYRIINI Velesunio Alathyria jacksoni Alathyria profuga Alathyria pertexta Lortiella Microdontia

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
0.750 2 3 2 1 9 3 3 0.0
Cladomics cladogram RHIPIDODONTINI CASTALIINI Hyridella australis 2 Hyridella australis 1 Echyridella Hyridella depressa

Baker et al., 2003 [Publ 2690], fig. 5

cst iT tT bi riS oT roT roS iD
0.625 2 7 0 1 6 3 3 0.0
Cladomics cladogram LAMPSILINI Hyridella australis RHIPIDODONTINI Hyridella depressa Velesunio Echyridella CASTALIINI

Baker et al., 2004 [Publ 2685], fig. 2-3

cst iT tT bi riS oT roT roS iD
0.625 2 3 0 1 7 3 3 0.0
Cladomics cladogram Hyridella depressa Hyridella australis RHIPIDODONTINI Echyridella CASTALIINI Velesunio 3 Velesunio 4 Velesunio 2 Velesunio angasi Velesunio 1 Alathyria

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 45 4 3 0.0
Cladomics cladogram Echyridella Hyridella CASTALIINI RHIPIDODONTINI

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.625 2 8 0 1 34 3 3 0.0
Cladomics cladogram CASTALIINI RHIPIDODONTINI Hyridella depressa Hyridella australis Echyridella Velesunio ambiguus Velesunio 3 Velesunio 2 Alathyria Velesunio 1 Velesunio angasi Lortiella

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 45 4 3 0.0
Cladomics cladogram Echyridella Hyridella CASTALIINI RHIPIDODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 44 4 3 0.0
Cladomics cladogram Echyridella Hyridella CASTALIINI RHIPIDODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 44 4 3 0.0
Cladomics cladogram Echyridella Hyridella CASTALIINI RHIPIDODONTINI

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 16 2 2 0.0
Cladomics cladogram Hyridella Velesunio

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 2 2 0.0
Cladomics cladogram Hyridella RHIPIDODONTINI CASTALIINI

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 2 2 0.0
Cladomics cladogram Hyridella CASTALIINI HYRIINI

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 2 2 0.0
Cladomics cladogram Hyridella CASTALIINI HYRIINI

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 2 2 0.0
Cladomics cladogram Hyridella HYRIINI CASTALIINI

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 14 1 1 0.0
Cladomics cladogram Velesunio Hyridella

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 1 1 0.0
Cladomics cladogram HYRIINI Hyridella

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 28 1 1 0.0
Cladomics cladogram Velesunio Hyridella

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 27 1 1 0.0
Cladomics cladogram Velesunio Hyridella

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 27 1 1 0.0
Cladomics cladogram Velesunio Hyridella

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 33 1 1 0.0
Cladomics cladogram Hyridella Velesunio

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 33 1 1 0.0
Cladomics cladogram Hyridella Velesunio

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 68 1 1 0.0
Cladomics cladogram Hyridella Velesunio

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 68 1 1 0.0
Cladomics cladogram Hyridella Velesunio

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 68 1 1 0.0
Cladomics cladogram Hyridella Velesunio

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 14 1 1 0.0
Cladomics cladogram HYRIINI Hyridella Velesunio

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 14 1 1 0.0
Cladomics cladogram HYRIINI Velesunio Hyridella

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 15 1 1 0.0
Cladomics cladogram Velesunio Hyridella

Chen et al., 2025 [Publ 4455], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 25 1 1 0.0
Cladomics cladogram Hyridella HYRIINI

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 0.0
Cladomics cladogram Hyridella Margaritifera Cumberlandia UNIONINI ANODONTINI QUADRULINI PLEUROBEMINI AMBLEMINI LAMPSILINI

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 0.0
Cladomics cladogram Hyridella Margaritifera Cumberlandia UNIONINI ANODONTINI QUADRULINI PLEUROBEMINI AMBLEMINI LAMPSILINI

Walker et al., 2006 [Publ 2666], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 0.0
Cladomics cladogram Hyridella ANODONTINI LAMPROTULINI GONIDEINI PSEUDODONTINI LAMPSILINI PLEUROBEMINI Cumberlandia

Walker et al., 2006 [Publ 2666], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 0.0
Cladomics cladogram Hyridella Cumberlandia PSEUDODONTINI Gonidea Inversidens ANODONTINI PLEUROBEMINI LAMPSILINI

Walker et al., 2006 [Publ 2666], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 0.0
Cladomics cladogram Hyridella Cumberlandia PSEUDODONTINI Gonidea Inversidens ANODONTINI PLEUROBEMINI LAMPSILINI

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 0 0 0.0
Cladomics cladogram Hyridella Velesunio

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 0.0
Cladomics cladogram Hyridella Margaritifera QUADRULINI PLEUROBEMINI AMBLEMINI POPENAIADINI LAMPSILINI CHAMBERLAINIINI RECTIDENTINI LAMPROTULINI GONIDEINI PSEUDODONTINI LANCEOLARIINI UNIONINI ANODONTINI LAMELLIDENTINI LEOPARREYSIINI COELATURINI INDOCHINELLINI

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 0.0
Cladomics cladogram Hyridella Margaritifera

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 0 0 0.0
Cladomics cladogram Hyridella Velesunio

Klunzinger et al., 2021 [Publ 4086], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 0.0
Cladomics cladogram Velesunio Cucumerunio

Klunzinger et al., 2022 [Publ 4259], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 0.0
Cladomics cladogram Cucumerunio Velesunio Westralunio