Bivalvia | Unionida | Hyrioidea | Hyriidae | Hyriinae

tribe HYRIDELLINI McMichael, 1956 (1934): cladomics

4 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 9 (13 total)
number of subtaxa (rank: genus) = 4
analyzed in 46 cladograms in 28 publications
maximum number of terminals in an analysis = 4 (mean = 1.59, median = 1)
maximum number of subtaxa (genus) = 2 (mean = 1.11, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 16
maximum number of included terminals = 4, (mean = 2.69 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.31, median = 1)
proportion of cladograms that support basic monophyly = 0.56

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 5
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 4 (mean = 3.6, median = 4)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.4

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyridellini cladogram distances
PCoA Ordination

NJ Tree of Hyridellini cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 18 20 2 626 14 4 0 
                             +----- Diplodon_suavidicus__2
  +--------------------------+                            
  |                          |  +-- Hyriini               
  |                          +--+                         
  |                             +-- Castaliini            
  |                                                       
  |                             +-- Hyridella_depressa **    
--+                       +-----+                         
  |                       |     +-- Hyridella_drapeta **     
  |  +--------------------+                               
  |  |                    |  +----- Diplodon_suavidicus__1
  |  |                    +--+                            
  |  |                       |  +-- Diplodon_chilensis    
  |  |                       +--+                         
  +--+                          +-- Diplodon_gratus       
     |                                                    
     |                          +-- Cucumerunio **           
     |  +-----------------------+                         
     |  |                       +-- Hyridella_australis **   
     |  |                                                 
     +--+  +----------------------- Echyridella           
        |  |                                              
        |  |              +-------- Velesunio__1          
        +--+  +-----------+                               
           |  |           |  +----- Alathyria_jacksoni__1 
           |  |           +--+                            
           |  |              |  +-- Velesunio_ambiguus    
           +--+              +--+                         
              |                 +-- Alathyria_profuga     
              |                                           
              |  +----------------- Velesunio__3          
              |  |                                        
              +--+  +-------------- Microdontia           
                 |  |                                     
                 +--+           +-- Lortiella             
                    |  +--------+                         
                    |  |        +-- Velesunio_angasi      
                    +--+                                  
                       |  +-------- Alathyria_pertexta__2 
                       +--+                               
                          |  +----- Velesunio__2          
                          +--+                            
                             |  +-- Alathyria_jacksoni__2 
                             +--+                         
                                +-- Alathyria_pertexta__1

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 2 16 3 3 18 
     +----- Rhipidodontini     
  +--+                         
  |  |  +-- Hyriini            
  |  +--+                      
  |     +-- Castaliini         
--+                            
  |     +-- Hyridella_depressa ** 
  |  +--+                      
  |  |  +-- Hyridella_drapeta **  
  +--+                         
     |  +-- Hyridella_australis **
     +--+                      
        +-- Cucumerunio **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
4 13 11 2 16 9 4 17.5 
                    +----- Diplodon_suavidicus__2
  +-----------------+                            
  |                 |  +-- Hyriini               
  |                 +--+                         
  |                    +-- Castaliini            
  |                                              
  |              +-------- Westralunio           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta **     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa **    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1
  +--+              +--+                         
     |                 +-- Diplodon_gratus       
     |                                           
     |  +----------------- Echyridella           
     |  |                                        
     +--+              +-- Cucumerunio **           
        |  +-----------+                         
        |  |           +-- Hyridella_australis **   
        +--+                                     
           |  +----------- Velesunio_ambiguus    
           +--+                                  
              |  +-------- Alathyria             
              +--+                               
                 |  +----- Velesunio_angasi      
                 +--+                            
                    |  +-- Lortiella             
                    +--+                         
                       +-- Microdontia

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iTtTbiriSoTroTroSiD
4 19 20 2 16 9 4 17.5 
                 +----- Diplodon_suavidicus__1
  +--------------+                            
  |              |  +-- Hyriini               
  |              +--+                         
  |                 +-- Castaliini            
  |                                           
  |           +-------- Westralunio           
  |           |                               
  +-----------+     +-- Hyridella_drapeta **     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa **    
  |           +--+                            
  |              |  +-- Diplodon_gratus       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2
  |                                           
  |                 +-- Hyridella_australis **   
  |  +--------------+                         
  |  |              +-- Cucumerunio **           
  +--+                                        
     |  +-------------- Echyridella           
     +--+                                     
        |  +----------- Velesunio_ambiguus    
        +--+                                  
           |  +-------- Microdontia           
           +--+                               
              |  +----- Lortiella             
              +--+                            
                 |  +-- Alathyria             
                 +--+                         
                    +-- Velesunio_angasi

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3 3 1 1 198 4 4 19 
1

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3 3 1 1 196 4 4 19 
  +----------- Echyridella   
--+                          
  |  +-------- Hyridella **     
  +--+                       
     |  +----- Rhipidodontini
     +--+                    
        |  +-- Castaliini    
        +--+                 
           +-- Hyriini

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 1 198 4 4 19 
  +----------- Echyridella   
--+                          
  |  +-------- Hyridella **     
  +--+                       
     |  +----- Rhipidodontini
     +--+                    
        |  +-- Castaliini    
        +--+                 
           +-- Hyriini

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
2 8 0 1 34 3 3 14 
  +----------------- Castaliini         
  |                                     
  +----------------- Rhipidodontini     
--+                                     
  |  +-------------- Hyridella_depressa ** 
  |  |                                  
  |  +-------------- Hyridella_australis **
  +--+                                  
     |  +----------- Echyridella        
     |  |                               
     |  |        +-- Velesunio_ambiguus 
     +--+  +-----+                      
        |  |     +-- Velesunio__3       
        |  |                            
        +--+  +----- Velesunio__2       
           |  |                         
           |  |  +-- Alathyria          
           +--+--+                      
              |  +-- Velesunio__1       
              |                         
              |  +-- Velesunio_angasi   
              +--+                      
                 +-- Lortiella

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
2 3 0 1 9 3 3 22 
           +-- Rhipidodontini        
  +--------+                         
  |        +-- Castaliini            
--+                                  
  |  +-------- Hyridella_australis__2 **
  +--+                               
     |  +----- Hyridella_australis__1 **
     +--+                            
        |  +-- Echyridella           
        +--+                         
           +-- Hyridella_depressa **

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
2 3 0 1 7 3 3 21 
  +----- Hyridella_depressa ** 
--+                         
  |  +-- Hyridella_australis **
  +--+                      
     +-- Rhipidodontini

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 45 4 3 -- 
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 12 5 2 -- 
  +----- Rhipidodontini
--+                    
  |  +-- Cucumerunio **   
  +--+                 
     +-- Hyridella **

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
2 7 0 1 6 3 3 21 
           +-- Hyridella_australis **
  +--------+                      
  |        +-- Rhipidodontini     
--+                               
  |  +-------- Hyridella_depressa ** 
  +--+                            
     |  +----- Velesunio          
     +--+                         
        |  +-- Echyridella        
        +--+                      
           +-- Castaliini

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 1 44 4 3 -- 
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 44 4 3 -- 
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 45 4 3 -- 
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
-- Hyridella_australis **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Hyridella_australis **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 25 1 1 -- 
  +-- Hyridella **
--+            
  +-- Hyriini

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
  +----- Hyridella ** 
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Klunzinger et al., 2022 [Publ 4259], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Cucumerunio_novaehollandiae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 0 0 -- 
-- Hyridella_depressa **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
-- Hyridella_australis **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Hyridella_depressa **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 2 2 -- 
  +----- Hyridella ** 
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Hyridella_depressa **

Klunzinger et al., 2021 [Publ 4086], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Cucumerunio_novaehollandiae **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 68 1 1 -- 
-- Hyridella_australis **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 2 2 -- 
  +----- Hyridella **     
--+                    
  |  +-- Rhipidodontini
  +--+                 
     +-- Castaliini

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
  +----- Hyridella ** 
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
-- Hyridella_australis **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Hyridella_australis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 27 1 1 -- 
-- Hyridella_australis **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 28 1 1 -- 
-- Hyridella_australis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Hyridella_australis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 80 0 0 -- 
-- Hyridella_depressa **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
  +-- Hyriini  
--+            
  +-- Hyridella **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 2 2 -- 
-- Hyridella **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 1 1 -- 
-- Hyridella_australis **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Hyridella **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Hyridella_australis **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 68 1 1 -- 
-- Hyridella_australis **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Hyridella_australis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 27 1 1 -- 
-- Hyridella_australis **