Bivalvia | Unionida | Hyrioidea | Hyriidae | Hyriinae

tribe HYRIINI Swainson, 1840: cladomics

2 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (5 total)
number of subtaxa (rank: genus) = 2
analyzed in 36 cladograms in 20 publications
maximum number of terminals in an analysis = 3 (mean = 1.22, median = 1)
maximum number of subtaxa (genus) = 2 (mean = 1.17, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 6
maximum number of included terminals = 3, (mean = 2.33 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support basic monophyly = 1

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 6
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 3 (mean = 2.33, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 1

Cladograms

Prié et al., 2026 [Publ 4844], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
0.812 2 2 0 2 21 13 4 ** 
  +-------------- Diplodon   
--+                          
  |  +----------- Guyanhyrida
  +--+                       
     |  +-------- Castaliini 
     +--+                    
        |  +----- Iridea     
        +--+                 
           |  +-- Prisodon **   
           +--+              
              +-- Triplodon **

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

cstiTtTbiriSoTroTroSiD
0.667 2 2 0 2 19 12 4 0.0 
  +-------- Rhipidodontini
--+                       
  |  +----- Castaliini    
  +--+                    
     |  +-- Triplodon **     
     +--+                 
        +-- Prisodon **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

cstiTtTbiriSoTroTroSiD
0.667 2 2 0 2 19 12 4 0.0 
  +-------- Rhipidodontini
--+                       
  |  +----- Castaliini    
  +--+                    
     |  +-- Triplodon **     
     +--+                 
        +-- Prisodon **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cstiTtTbiriSoTroTroSiD
0.667 3 3 1 2 630 16 4 0.0 
  +-------- Rhipidodontini
--+                       
  |  +----- Castaliini    
  +--+                    
     |  +-- Triplodon **     
     +--+                 
        +-- Prisodon **

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

cstiTtTbiriSoTroTroSiD
0.900 3 3 1 2 4 4 2 0.0 
  +-------- Rhipidodontini
--+                       
  |  +----- Castaliini    
  +--+                    
     |  +-- Triplodon **     
     +--+                 
        +-- Prisodon **

Santos-Neto et al., 2018 [Publ 3888], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 3 3 1 0.0 
  +----- Castaliini
--+                
  |  +-- Triplodon ** 
  +--+             
     +-- Prisodon **

Huang et al., 2019 [Publ 3843], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 200 6 4 0.0 
  +----------- Echyridella   
--+                          
  |  +-------- Hyridellini   
  +--+                       
     |  +----- Rhipidodontini
     +--+                    
        |  +-- Castaliini    
        +--+                 
           +-- Triplodon **

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 198 6 4 0.0 
  +----------- Echyridella   
--+                          
  |  +-------- Hyridellini   
  +--+                       
     |  +----- Rhipidodontini
     +--+                    
        |  +-- Castaliini    
        +--+                 
           +-- Triplodon **

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.900 1 1 0 1 200 6 4 0.0 
  +----- Rhipidodontini
--+                    
  |  +-- Castaliini    
  +--+                 
     +-- Triplodon **

Graf et al., 2015 [Publ 3619], fig. 1

cstiTtTbiriSoTroTroSiD
0.900 1 1 0 1 19 6 3 0.0 
  +-------- Hyridellini   
--+                       
  |  +----- Rhipidodontini
  +--+                    
     |  +-- Triplodon **     
     +--+                 
        +-- Castaliini

Guerra et al., 2019 [Publ 4168], fig. 1

cstiTtTbiriSoTroTroSiD
0.875 1 1 0 1 40 3 3 0.0 
  +----- Rhipidodontini
--+                    
  |  +-- Castaliini    
  +--+                 
     +-- Prisodon **

Guerra et al., 2019 [Publ 4168], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 3 3 0.0 
  +-------- Echyridella   
--+                       
  |  +----- Rhipidodontini
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Prisodon **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 139 3 3 0.0 
  +-------- Echyridella   
--+                       
  |  +----- Rhipidodontini
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Prisodon **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 2 2 0.0 
  +----- Hyridellini
--+                 
  |  +-- Castaliini 
  +--+              
     +-- Triplodon **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 2 2 0.0 
  +----- Hyridellini
--+                 
  |  +-- Castaliini 
  +--+              
     +-- Triplodon **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 39 2 2 0.0 
  +----- Hyridellini
--+                 
  |  +-- Triplodon **  
  +--+              
     +-- Castaliini

Bolotov et al., 2016 [Publ 3743], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 29 2 2 0.0 
  +----- Rhipidodontini
--+                    
  |  +-- Triplodon **     
  +--+                 
     +-- Castaliini

Bolotov et al., 2016 [Publ 3743], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 29 2 2 0.0 
  +----- Rhipidodontini
--+                    
  |  +-- Triplodon **     
  +--+                 
     +-- Castaliini

Bolotov et al., 2017 [Publ 3761], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 138 2 2 0.0 
  +----- Rhipidodontini
--+                    
  |  +-- Castaliini    
  +--+                 
     +-- Triplodon **

Bolotov et al., 2017 [Publ 3761], fig. S2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 138 2 2 0.0 
  +----- Rhipidodontini
--+                    
  |  +-- Castaliini    
  +--+                 
     +-- Triplodon **

Konopleva et al., 2017 [Publ 3807], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 2 2 0.0 
  +----- Rhipidodontini
--+                    
  |  +-- Triplodon **     
  +--+                 
     +-- Castaliini

Lawrie et al., 2026 [Publ 4843], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 17 5 2 0.0 
  +-- Castaliini
--+             
  +-- Triplodon **

Combosch et al., 2017 [Publ 3806], fig. 1

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 14 1 1 0.0 
  +----- Triplodon **  
--+                 
  |  +-- Velesunio  
  +--+              
     +-- Hyridellini

Combosch et al., 2017 [Publ 3806], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 1 1 0.0 
  +-- Triplodon **  
--+              
  +-- Hyridellini

Huang et al., 2018 [Publ 3828], fig. 3_5

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 28 1 1 0.0 
  +----- Triplodon **  
--+                 
  |  +-- Velesunio  
  +--+              
     +-- Hyridellini

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 27 1 1 0.0 
  +----- Triplodon **  
--+                 
  |  +-- Velesunio  
  +--+              
     +-- Hyridellini

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 27 1 1 0.0 
  +----- Triplodon **  
--+                 
  |  +-- Velesunio  
  +--+              
     +-- Hyridellini

Wu et al., 2020 [Publ 4121], fig. 2

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 33 1 1 0.0 
  +----- Triplodon **  
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio

Wu et al., 2020 [Publ 4121], fig. S1

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 33 1 1 0.0 
  +----- Triplodon **  
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio

Sano et al., 2020 [Publ 4138], fig. 2

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 68 1 1 0.0 
  +----- Triplodon **  
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio

Sano et al., 2020 [Publ 4138], fig. 3

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 68 1 1 0.0 
  +----- Triplodon **  
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio

Sano et al., 2020 [Publ 4138], fig. 4

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 68 1 1 0.0 
  +----- Triplodon **  
--+                 
  |  +-- Hyridellini
  +--+              
     +-- Velesunio

Sonowal et al., 2021 [Publ 4221], fig. 3a

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 14 1 1 0.0 
  +-- Triplodon **  
  |              
--+-- Hyridellini
  |              
  +-- Velesunio

Sonowal et al., 2021 [Publ 4221], fig. 3b

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 14 1 1 0.0 
  +-- Triplodon **  
  |              
--+-- Velesunio  
  |              
  +-- Hyridellini

Sonowal et al., 2021 [Publ 4221], fig. 4

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 15 1 1 0.0 
  +----- Triplodon **  
--+                 
  |  +-- Velesunio  
  +--+              
     +-- Hyridellini

Chen et al., 2025 [Publ 4455], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 25 1 1 0.0 
  +-- Hyridellini
--+              
  +-- Triplodon **