Bivalvia | Unionida

superfamily HYRIOIDEA Swainson, 1840: cladomics

1 family

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 94 (127 total)
number of subtaxa (rank: family) = 1
analyzed in 115 cladograms in 58 publications
maximum number of terminals in an analysis = 29 (mean = 4.22, median = 3)
maximum number of subtaxa (family) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 66
maximum number of included terminals = 29, (mean = 6.61 median = 5)
maximum number of subtaxa (family) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Hyrioidea cladogram distances
PCoA Ordination

NJ Tree of Hyrioidea cladogram distances
NJ Tree

Cladograms

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 36 35 2 **
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Graf, 2000 [Publ 2647], fig. 1

cst iT tT bi riS oT roT roS iD
0.848 4 4 2 1 13 12 2 5.5
Cladomics cladogram MARGARITIFERIDAE UNIONINI PLEUROBEMINI ANODONTINI LAMPSILINI PARREYSIINI COELATURINI HYRIIDAE ETHERIOIDEA

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
0.958 1 1 0 1 29 28 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
0.958 1 1 0 1 29 28 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
0.891 5 5 3 1 29 28 2 7.0
Cladomics cladogram HYRIIDAE IRIDINIDAE Etheria Mulleria MYCETOPODIDAE PARREYSIINAE Pseudomulleria ANODONTINI GONIDEINAE UNIONINI MARGARITIFERIDAE AMBLEMINAE

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
0.817 5 5 1 1 26 25 2 7.0
Cladomics cladogram MARGARITIFERIDAE AMBLEMINI PARREYSIINAE Eurynia Fusconaia GONIDEINAE Pleurobema QUADRULINI Ortmanniana Glebula Potamilus Ligumia Toxolasma Cyrtonaias UNIONINI ANODONTINI HYRIIDAE ETHERIOIDEA

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
0.917 5 5 3 1 26 25 2 7.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
0.900 5 5 3 1 26 25 2 7.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
0.900 5 5 2 1 26 25 2 7.0
Cladomics cladogram UNIONOIDEA ETHERIOIDEA HYRIIDAE

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
0.967 5 5 3 1 26 25 2 7.0
Cladomics cladogram MARGARITIFERIDAE UNIONIDAE ETHERIOIDEA HYRIIDAE

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
0.974 9 9 7 1 38 37 2 7.5
Cladomics cladogram UNIONOIDEA HYRIIDAE ETHERIOIDEA

Walker et al., 2006 [Publ 2666], fig. 6

cst iT tT bi riS oT roT roS iD
0.857 10 10 7 1 26 25 2 13.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
0.897 9 9 7 1 38 37 2 7.5
Cladomics cladogram HYRIIDAE IRIDINIDAE Etheria Mulleria MYCETOPODIDAE MARGARITIFERIDAE PARREYSIINAE Pseudomulleria UNIONINAE Rectidentini AMBLEMINAE GONIDEINI PSEUDODONTINI

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
0.908 9 9 7 1 37 36 2 7.5
Cladomics cladogram HYRIIDAE Etheria Mulleria MYCETOPODIDAE IRIDINIDAE PARREYSIINAE Pseudomulleria MARGARITIFERIDAE PSEUDODONTINI Rectidentini UNIONINAE GONIDEINI AMBLEMINAE

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
0.922 9 9 7 1 37 36 2 7.5
Cladomics cladogram PARREYSIINAE MARGARITIFERIDAE HYRIIDAE ETHERIOIDEA PSEUDODONTINI Rectidentini UNIONINAE GONIDEINI AMBLEMINAE

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
0.946 4 4 2 1 32 31 2 0.0
Cladomics cladogram HYRIIDAE MARGARITIFERIDAE ETHERIOIDEA UNIONIDAE

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
0.946 4 4 2 1 32 31 2 0.0
Cladomics cladogram ETHERIOIDEA UNIONIDAE MARGARITIFERIDAE HYRIIDAE

Bieler et al., 2014 [Publ 3622], fig. 26

cst iT tT bi riS oT roT roS iD
0.944 1 1 0 1 97 3 2 --
Cladomics cladogram HYRIIDAE UNIONIDAE ETHERIOIDEA MARGARITIFERIDAE

Bieler et al., 2014 [Publ 3622], fig. 27

cst iT tT bi riS oT roT roS iD
0.944 1 1 0 1 97 3 2 --
Cladomics cladogram HYRIIDAE UNIONIDAE ETHERIOIDEA MARGARITIFERIDAE

Bieler et al., 2014 [Publ 3622], fig. 28

cst iT tT bi riS oT roT roS iD
0.944 1 1 0 1 97 3 2 --
Cladomics cladogram ETHERIOIDEA MARGARITIFERIDAE UNIONIDAE HYRIIDAE

Bieler et al., 2014 [Publ 3622], fig. 29

cst iT tT bi riS oT roT roS iD
0.944 1 1 0 1 97 3 2 --
Cladomics cladogram HYRIIDAE UNIONIDAE ETHERIOIDEA MARGARITIFERIDAE

Bieler et al., 2014 [Publ 3622], fig. 30

cst iT tT bi riS oT roT roS iD
0.944 1 1 0 1 97 3 2 --
Cladomics cladogram HYRIIDAE UNIONIDAE ETHERIOIDEA MARGARITIFERIDAE

Bieler et al., 2014 [Publ 3622], fig. 31

cst iT tT bi riS oT roT roS iD
0.944 1 1 0 1 97 3 2 --
Cladomics cladogram HYRIIDAE UNIONIDAE MARGARITIFERIDAE ETHERIOIDEA

Graf et al., 2015 [Publ 3619], fig. 1

cst iT tT bi riS oT roT roS iD
0.946 14 14 12 1 6 5 2 10.0
Cladomics cladogram HYRIIDAE UNIONOIDEA ETHERIOIDEA

González & Giribet, 2015 [Publ 3632], fig. 2

cst iT tT bi riS oT roT roS iD
0.944 1 1 0 1 5 3 2 --
Cladomics cladogram ETHERIOIDEA MARGARITIFERIDAE UNIONIDAE HYRIIDAE

González & Giribet, 2015 [Publ 3632], fig. 3

cst iT tT bi riS oT roT roS iD
0.944 1 1 0 1 5 3 2 --
Cladomics cladogram UNIONIDAE HYRIIDAE ETHERIOIDEA MARGARITIFERIDAE

González & Giribet, 2015 [Publ 3632], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 3 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

González & Giribet, 2015 [Publ 3632], fig. 6

cst iT tT bi riS oT roT roS iD
0.944 1 1 0 1 5 3 2 --
Cladomics cladogram HYRIIDAE UNIONIDAE ETHERIOIDEA MARGARITIFERIDAE

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
0.955 9 9 7 1 21 18 2 7.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
0.955 9 9 7 1 21 18 2 7.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.944 2 2 0 1 79 78 2 --
Cladomics cladogram HYRIIDAE MARGARITIFERIDAE ETHERIOIDEA UNIONIDAE

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.920 9 9 7 1 130 128 2 7.0
Cladomics cladogram MARGARITIFERIDAE UNIONIDAE ETHERIOIDEA HYRIIDAE

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.856 9 9 7 1 130 128 2 7.0
Cladomics cladogram Chamberlainia MARGARITIFERIDAE ETHERIOIDEA HYRIIDAE PARREYSIINAE UNIONINAE Rectidentini AMBLEMINAE PSEUDODONTINI GONIDEINI Sinohyriopsis LAMPROTULINI

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
0.980 1 1 0 1 24 23 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
0.980 1 1 0 1 24 23 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
0.923 3 3 1 1 210 10 2 3.0
Cladomics cladogram HYRIIDAE UNIONIDAE MARGARITIFERIDAE ETHERIOIDEA

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
0.974 3 3 1 1 211 10 2 1.0
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 28 26 2 3.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.959 3 3 1 1 26 24 2 3.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 40 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

cst iT tT bi riS oT roT roS iD
0.953 3 3 1 1 25 23 2 3.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

cst iT tT bi riS oT roT roS iD
0.953 3 3 1 1 25 23 2 3.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.899 14 14 12 1 187 185 2 10.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
0.972 1 1 0 1 55 54 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.893 14 14 12 1 185 183 2 10.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.824 14 14 9 1 187 185 2 11.0
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
0.985 1 1 0 1 45 45 2 --
Cladomics cladogram UNIONIDAE MARGARITIFERIDAE HYRIIDAE ETHERIOIDEA

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 55 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
0.990 1 1 0 1 56 55 2 --
Cladomics cladogram ETHERIOIDEA UNIONIDAE HYRIIDAE MARGARITIFERIDAE

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
0.928 5 5 3 1 36 35 2 5.0
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
0.928 5 5 3 1 36 35 2 5.0
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.948 2 2 0 1 120 118 2 --
Cladomics cladogram HYRIIDAE MARGARITIFERIDAE ETHERIOIDEA UNIONIDAE

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
0.941 3 3 1 1 31 29 2 3.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 46 2 --
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
0.922 3 3 1 1 31 29 2 3.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.926 3 3 1 1 66 64 2 3.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.926 3 3 1 1 66 64 2 3.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.926 3 3 1 1 66 64 2 3.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
0.965 3 3 1 1 88 86 2 4.0
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.966 1 1 0 1 72 71 2 --
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.966 1 1 0 1 72 71 2 --
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.964 1 1 0 1 67 66 2 --
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.947 3 3 1 1 88 86 2 4.0
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
0.988 1 1 0 1 72 72 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
0.990 1 1 0 1 94 93 2 --
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.960 1 1 0 1 94 93 2 --
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 19 19 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 20 20 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.980 1 1 0 1 90 89 2 --
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.980 1 1 0 1 90 89 2 --
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
0.990 1 1 0 1 86 86 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
0.978 1 1 0 1 84 84 2 --
Cladomics cladogram ETHERIOIDEA UNIONIDAE HYRIIDAE MARGARITIFERIDAE

Chen et al., 2025 [Publ 4455], fig. 1

cst iT tT bi riS oT roT roS iD
0.943 2 2 0 1 24 22 2 --
Cladomics cladogram HYRIIDAE ETHERIOIDEA UNIONOIDEA

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.943 5 5 3 1 135 134 2 5.0
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.737 29 29 34 1 604 602 2 34.0
Cladomics cladogram IRIDININAE ASPATHARIINAE Etheria Anodontites trapesialis Mulleria Lamproscapha Monocondylaea Tamsiella MYCETOPODINAE Anodontites guanarensis Fossula Anodontites trigona Anodontites elongata HYRIIDAE UNIONINAE Lamellidens Balwantia Trapezidens LEOPARREYSIINI Pseudomulleria INDOCHINELLINI PARREYSIINI COELATURINI Rectidentini MARGARITIFERIDAE AMBLEMINAE INDOPSEUDODONTINA Lannanaia Pilsbryoconcha Songkhlanaia Nyeinchanconcha Sundadontina Isannaia Namkongnaia Thaiconcha Bineurus Pseudodon walpolei Pseudodon vagulus Pseudodon vondembuschianus Pseudodon mekongi SCHEPMANIINI Pseudodon lenyanensis 2 Pseudodon inoscularis Pseudodon lenyanensis 1 CHAMBERLAINIINI Parvasolenaia Pseudopostolata Koreosolenaia Cosmopseudodon Guiunio Ptychorhynchus Obovalis Postolata Caudiculatus LAMPROTULINI Sinosolenaia carinata Gonidea Microcondylaea Leguminaia Inversidens Sinosolenaia recognita Sinosolenaia iridinea Sinosolenaia oleivora

Prié et al., 2026 [Publ 4844], fig. 2

cst iT tT bi riS oT roT roS iD
0.886 20 20 15 1 3 3 2 17.0
Cladomics cladogram ETHERIOIDEA UNIONOIDEA HYRIIDAE

Prié et al., 2026 [Publ 4844], fig. 3

cst iT tT bi riS oT roT roS iD
0.870 1 1 0 1 14 14 2 --
Cladomics cladogram ETHERIOIDEA HYRIIDAE UNIONOIDEA

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

cst iT tT bi riS oT roT roS iD
0.900 6 6 5 1 5 5 1 5.5
Cladomics cladogram MARGARITIFERIDAE UNIONIDAE HYRIIDAE

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
0.977 4 4 2 1 14 13 1 3.5
Cladomics cladogram HYRIIDAE UNIONOIDEA

Hoeh et al., 2002 [Publ 2673], fig. 1-F

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 8 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Hoeh et al., 2002 [Publ 2673], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 8 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Baker et al., 2003 [Publ 2690], fig. 5

cst iT tT bi riS oT roT roS iD
0.833 7 7 5 1 1 1 1 8.0
Cladomics cladogram UNIONOIDEA HYRIIDAE

Baker et al., 2004 [Publ 2685], fig. 2-3

cst iT tT bi riS oT roT roS iD
0.789 8 8 9 1 1 1 1 12.5
Cladomics cladogram UNIONOIDEA HYRIIDAE

Walker et al., 2006 [Publ 2666], fig. 2

cst iT tT bi riS oT roT roS iD
0.912 1 1 0 1 14 13 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Walker et al., 2006 [Publ 2666], fig. 3

cst iT tT bi riS oT roT roS iD
0.939 1 1 0 1 14 13 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Walker et al., 2006 [Publ 2666], fig. 4

cst iT tT bi riS oT roT roS iD
0.939 1 1 0 1 14 13 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Walker et al., 2006 [Publ 2666], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 13 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
0.938 1 1 0 1 37 37 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 17 17 1 --
Cladomics cladogram CRISTARIINA LANCEOLARIINI Anodonta UNIONINI ALASMIDONTINA Pseudanodonta AMBLEMINAE HYRIIDAE LAMPROTULINI CHAMBERLAINIINI

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
0.983 1 1 0 1 30 30 1 --
Cladomics cladogram MARGARITIFERIDAE HYRIIDAE UNIONIDAE

Marshall et al., 2014 [Publ 3593], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 10 10 8 1 4 4 1 10.0
Cladomics cladogram MARGARITIFERIDAE UNIONINAE AMBLEMINAE HYRIIDAE

Elderkin et al., 2016 [Publ 3693], fig. 2

cst iT tT bi riS oT roT roS iD
0.909 1 1 0 1 2 2 1 --
Cladomics cladogram ETHERIIDAE IRIDINIDAE HYRIIDAE

Guerra et al., 2017 [Publ 3772], fig. 2a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 7 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Guerra et al., 2017 [Publ 3772], fig. 2b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 7 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 16 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
0.936 1 1 0 1 38 38 1 --
Cladomics cladogram MARGARITIFERIDAE UNIONINAE LAMPSILINI HYRIIDAE AMBLEMINI PLEUROBEMINI QUADRULINI

Chase et al., 2018 [Publ 3847], fig. 5

cst iT tT bi riS oT roT roS iD
0.962 1 1 0 1 9 9 1 --
Cladomics cladogram UNIONIDAE MARGARITIFERIDAE HYRIIDAE

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 10 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
0.966 1 1 0 1 10 10 1 --
Cladomics cladogram UNIONIDAE MARGARITIFERIDAE HYRIIDAE

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 22 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 20 20 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 20 20 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Klunzinger et al., 2021 [Publ 4086], fig. 2

cst iT tT bi riS oT roT roS iD
0.944 4 4 2 1 2 2 1 5.0
Cladomics cladogram UNIONOIDEA HYRIIDAE

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
0.758 3 3 0 1 12 9 1 2.0
Cladomics cladogram HYRIIDAE TRIGONIOIDEA

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
0.758 3 3 0 1 12 9 1 2.0
Cladomics cladogram HYRIIDAE UNIONOIDEA

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 3 3 1 1 13 10 1 3.0
Cladomics cladogram HYRIIDAE UNIONOIDEA

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 45 44 1 --
Cladomics cladogram HYRIIDAE UNIONOIDEA

Lawrie et al., 2026 [Publ 4843], fig. 1

cst iT tT bi riS oT roT roS iD
0.826 17 17 21 1 1 1 1 21.0
Cladomics cladogram UNIONOIDEA HYRIIDAE

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

cst iT tT bi riS oT roT roS iD
0.708 20 20 20 1 1 0 0 21.0
Cladomics cladogram TRIGONIOIDEA HYRIIDAE

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.923 7 7 6 1 0 0 0 8.0
Cladomics cladogram HYRIINAE

Santos-Neto et al., 2016 [Publ 3739], fig. 4

cst iT tT bi riS oT roT roS iD
0.708 20 20 21 1 1 0 0 21.0
Cladomics cladogram TRIGONIOIDEA HYRIIDAE

Santos-Neto et al., 2018 [Publ 3888], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 1 0 0 0 5.0
Cladomics cladogram HYRIINAE

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 1 0 0 0 5.0
Cladomics cladogram CASTALIINI

Klunzinger et al., 2022 [Publ 4259], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 1 0 0 0 5.0
Cladomics cladogram HYRIIDAE

Lawrie et al., 2026 [Publ 4843], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 9 9 13 1 0 0 0 22.0
Cladomics cladogram VELESUNIONINAE