Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae

tribe INDOCHINELLINI Bolotov, Pfeiffer, Vikhrev, & Konopleva in Bolotov et al., 2018: cladomics

9 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 34 (34 total)
number of subtaxa (rank: genus) = 9
analyzed in 69 cladograms in 43 publications
maximum number of terminals in an analysis = 31 (mean = 9.35, median = 4)
maximum number of subtaxa (genus) = 8 (mean = 3.43, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 42
maximum number of included terminals = 31, (mean = 14.71 median = 12)
maximum number of subtaxa (genus) = 8 (mean = 5, median = 4)
proportion of cladograms that support basic monophyly = 0.95

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 38
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 31 (mean = 15.47, median = 12)
maximum number of subtaxa (genus) = 8 (mean = 5.18, median = 4)
proportion of cladograms that support rigorous monophyly = 0.95

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Indochinellini cladogram distances
PCoA Ordination

NJ Tree of Indochinellini cladogram distances
NJ Tree

Cladograms

Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.957 29 29 27 8 33 33 4 **
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI PARREYSIINI COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2024 [Publ 4369], fig. S1

cst iT tT bi riS oT roT roS iD
0.913 29 29 27 8 34 33 4 16.0
Cladomics cladogram Parreysia LEOPARREYSIINI LAMELLIDENTINI Assamnaia COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.741 29 29 28 8 604 56 4 21.0
Cladomics cladogram PARREYSIINI COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Unionetta fabagina 2 Harmandia Unionetta fabagina 1 Scabies

Konopleva et al., 2026 [Publ 4847], fig. 2

cst iT tT bi riS oT roT roS iD
0.957 31 31 28 8 35 35 4 19.0
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI PARREYSIINI COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Scabies Unionetta Harmandia

Konopleva et al., 2026 [Publ 4847], fig. S1

cst iT tT bi riS oT roT roS iD
0.957 31 31 29 8 35 35 4 8.0
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI PARREYSIINI COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Konopleva et al., 2026 [Publ 4847], fig. S2

cst iT tT bi riS oT roT roS iD
0.913 31 31 29 8 36 35 4 16.0
Cladomics cladogram Parreysia LEOPARREYSIINI LAMELLIDENTINI Assamnaia COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Konopleva et al., 2026 [Publ 4847], fig. S3

cst iT tT bi riS oT roT roS iD
0.913 31 31 26 8 36 35 4 15.0
Cladomics cladogram Parreysia Assamnaia LEOPARREYSIINI LAMELLIDENTINI COELATURINI Pseudoscabies Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Pfeiffer et al., 2018 [Publ 3827], fig. 2

cst iT tT bi riS oT roT roS iD
0.950 17 17 15 7 11 10 4 16.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Unionetta fabagina 2 Harmandia Unionetta fabagina 1 Scabies

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 18 18 16 7 5 5 4 21.0
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI PARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2020 [Publ 4100], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 21 21 19 7 96 20 4 18.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Scabies Unionetta Harmandia

Bolotov et al., 2020 [Publ 4100], fig. S1

cst iT tT bi riS oT roT roS iD
0.947 20 20 17 7 91 20 4 16.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Radiatula crispisulcata Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4239], fig. 2

cst iT tT bi riS oT roT roS iD
0.950 26 26 24 7 32 28 4 17.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4239], fig. S1

cst iT tT bi riS oT roT roS iD
0.950 26 26 23 7 32 28 4 4.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4239], fig. S2

cst iT tT bi riS oT roT roS iD
0.950 26 26 24 7 32 28 4 21.0
Cladomics cladogram PARREYSIINI LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4252], fig. 2

cst iT tT bi riS oT roT roS iD
0.950 27 27 25 7 34 30 4 2.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Bolotov et al., 2022 [Publ 4252], fig. S1

cst iT tT bi riS oT roT roS iD
0.950 27 27 25 7 34 30 4 22.0
Cladomics cladogram PARREYSIINI LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabiellus Harmandia Unionetta Scabies

Chen et al., 2026 [Publ 4833], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 14 15 14 7 13 10 4 29.0
Cladomics cladogram Lamellidens Trapezidens PARREYSIINI LEOPARREYSIINI COELATURINI Indochinella Indonaia Radiatula PSEUDODONTINI Harmandia Chenunionetta Unionetta fabagina 2 Unionetta fabagina 1 Scabies

Konopleva et al., 2023 [Publ 4262], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 14 14 10 7 68 0 0 23.0
Cladomics cladogram PSEUDODONTINI Rectidentini GONIDEINI LAMPROTULINI Indonaia Indochinella Radiatula Scabiellus Scabies Unionetta Harmandia

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.882 11 11 9 4 128 18 4 28.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies crispata Radiatula Scabies mandarinus Scabies humilis Scabies phaselus

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.882 11 11 9 4 128 18 4 28.0
Cladomics cladogram PARREYSIINI LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies crispata Radiatula Scabies mandarinus Scabies humilis Scabies phaselus

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.647 12 13 9 4 113 26 4 24.0
Cladomics cladogram Trapezidens Lamellidens lamellatus Lamellidens jenkinsianus Lamellidens generosus Lamellidens mainwaringi Lamellidens ferrugineus Lamellidens marginalis Lamellidens savadiensis Lamellidens corrianus 1 PARREYSIINI LEOPARREYSIINI COELATURINI Indochinella Scabies Radiatula Indonaia shurtleffiana Indonaia rugosa Indonaia andersoniana Indonaia caerulea Indonaia bonneaudii Lamellidens corrianus 2

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 4 71 13 4 24.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula Scabies

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.941 12 12 9 4 189 22 4 26.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies Radiatula

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.941 12 12 9 4 187 22 4 24.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies Radiatula

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.941 12 12 9 4 189 22 4 24.0
Cladomics cladogram PARREYSIINI LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies Radiatula

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 53 6 3 25.0
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies Harmandia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 53 6 3 25.0
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Indochinella Scabies Harmandia

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

cst iT tT bi riS oT roT roS iD
0.812 8 8 3 4 28 28 2 24.0
Cladomics cladogram LEOPARREYSIINI COELATURINI Indonaia Indochinella Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Scabies

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

cst iT tT bi riS oT roT roS iD
0.812 8 8 1 4 28 28 2 26.0
Cladomics cladogram LEOPARREYSIINI COELATURINI Radiatula mouhoti Radiatula myitkyinae Radiatula chaudhurii Indochinella Indonaia Scabies

Konopleva et al., 2019 [Publ 3900], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 12 12 9 4 39 0 0 22.0
Cladomics cladogram Rectidentini Indonaia Indochinella Scabies Radiatula

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 37 5 3 26.0
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indochinella Radiatula Scabies

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 28 8 3 26.0
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indochinella Radiatula Scabies

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 3 86 8 3 26.0
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Scabies Radiatula

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.917 5 5 3 3 86 8 3 28.0
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indonaia Radiatula myitkyinae Scabies Radiatula mouhoti Radiatula chaudhurii

Bolotov et al., 2022 [Publ 4250], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 3 40 15 3 22.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI Indonaia Indochinella Radiatula

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 53 4 3 26.0
Cladomics cladogram LAMELLIDENTINI PARREYSIINI LEOPARREYSIINI Indonaia Indochinella Radiatula

Bolotov et al., 2022 [Publ 4250], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 12 12 10 3 3 3 2 21.0
Cladomics cladogram PARREYSIINI LEOPARREYSIINI Indonaia Indochinella Radiatula

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
0.923 2 2 0 2 34 7 3 --
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI COELATURINI Radiatula Indochinella

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
0.923 2 2 0 2 34 7 3 --
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI COELATURINI Radiatula Indochinella

Bolotov et al., 2019 [Publ 3974], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 2 12 7 2 23.0
Cladomics cladogram LEOPARREYSIINI COELATURINI Indochinella Radiatula

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 3 3 --
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI COELATURINI Indochinella

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 4 3 --
Cladomics cladogram LAMELLIDENTINI LEOPARREYSIINI Unionetta COELATURINI

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 121 2 2 --
Cladomics cladogram COELATURINI LEOPARREYSIINI Radiatula

Choi et al., 2020 [Publ 4169], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 2 2 --
Cladomics cladogram Radiatula LAMELLIDENTINI LEOPARREYSIINI

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 2 2 --
Cladomics cladogram COELATURINI Indonaia PARREYSIINI

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 2 2 --
Cladomics cladogram PARREYSIINI COELATURINI Indonaia

Imsong & Murali, 2023 [Publ 4370], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 2 2 --
Cladomics cladogram LAMELLIDENTINI Indonaia PARREYSIINI

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 2 2 --
Cladomics cladogram PARREYSIINI COELATURINI Indonaia

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 2 2 --
Cladomics cladogram COELATURINI Indonaia PARREYSIINI

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 2 2 --
Cladomics cladogram PARREYSIINI COELATURINI Indonaia

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 2 2 --
Cladomics cladogram PARREYSIINI COELATURINI Indonaia

Dai et al., 2024 [Publ 4385], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram LAMELLIDENTINI Scabies

Dai et al., 2024 [Publ 4385], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram LAMELLIDENTINI Scabies

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 20 1 1 --
Cladomics cladogram LAMELLIDENTINI Scabies

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 1 20 1 1 --
Cladomics cladogram LAMELLIDENTINI Scabies

Dai et al., 2026 [Publ 4757], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 1 1 --
Cladomics cladogram Scabies LAMELLIDENTINI

Combosch et al., 2017 [Publ 3806], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 212 0 0 --
Cladomics cladogram LAMPSILINI Scabies

Combosch et al., 2017 [Publ 3806], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 213 0 0 --
Cladomics cladogram Scabies LAMPSILINI

Huang et al., 2018 [Publ 3828], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 28 0 0 --
Cladomics cladogram LAMPSILINI Scabies

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram PSEUDODONTINI Scabies

Wu et al., 2020 [Publ 4121], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram LAMPSILINI Scabies

Wu et al., 2020 [Publ 4121], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram LAMPSILINI Scabies

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Scabies LAMPSILINI CHAMBERLAINIINI Inversidens LAMPROTULINI Ptychorhynchus Obovalis Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI UNIONINI

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Scabies CHAMBERLAINIINI LAMPSILINI LAMPROTULINI GONIDEINI Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LANCEOLARIINI LEPIDODESMINI UNIONINI

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 68 0 0 --
Cladomics cladogram Scabies LAMPSILINI CHAMBERLAINIINI Inversidens LAMPROTULINI Obovalis Ptychorhynchus Anemina Buldowskia Sinanodonta Pletholophus Beringiana Cristaria ANODONTINA ALASMIDONTINA LEPIDODESMINI LANCEOLARIINI UNIONINI

Wu et al., 2021 [Publ 4215], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 23 0 0 --
Cladomics cladogram Indonaia ACUTICOSTINI UNIONINI CHAMBERLAINIINI PSEUDODONTINI Sinosolenaia LAMPROTULINI Ptychorhynchus Parvasolenaia Inversidens Gonidea Microcondylaea Leguminaia

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Scabies ANODONTINI

Dai et al., 2024 [Publ 4386], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Scabies UNIONINA MIDDENDORFFINAIINA OXYNAIINA ACULAMPROTULINA ANODONTINI LEPIDODESMINI

Xiang et al., 2025 [Publ 4433], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Scabies ANODONTINI