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Taxonomy

Species & Genus Richness by
Taxonomy

Geography

Species Richness & Endemism by
Geography

Data last updated
Mon 1 Dec 2025

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae

tribe INDOCHINELLINI Bolotov, Pfeiffer, Vikhrev, & Konopleva in Bolotov et al., 2018: cladomics

8 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 32 (32 total)
number of subtaxa (rank: genus) = 8
analyzed in 60 cladograms in 38 publications
maximum number of terminals in an analysis = 29 (mean = 7.92, median = 3)
maximum number of subtaxa (genus) = 8 (mean = 3.12, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 36
maximum number of included terminals = 29, (mean = 12.53 median = 11.5)
maximum number of subtaxa (genus) = 8 (mean = 4.53, median = 4)
proportion of cladograms that support basic monophyly = 0.97

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 32
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 29 (mean = 13.16, median = 11.5)
maximum number of subtaxa (genus) = 8 (mean = 4.69, median = 4)
proportion of cladograms that support rigorous monophyly = 0.97

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Indochinellini cladogram distances
PCoA Ordination

NJ Tree of Indochinellini cladogram distances
NJ Tree

Cladograms

Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
29	29	27	8	33	33	4	0

  +-------------------------- Lamellidentini
  |                                         
--+                       +-- Leoparreysiini
  |  +--------------------+                 
  |  |                    +-- Parreysiini   
  +--+                                      
     |  +-------------------- Coelaturini   
     |  |                                   
     +--+                 +-- Pseudoscabies ** 
        |  +--------------+                 
        |  |              +-- Indonaia **      
        +--+                                
           |  +-------------- Indochinella **  
           +--+                             
              |  +----------- Radiatula **     
              +--+                          
                 |  +-------- Scabiellus **    
                 +--+                       
                    |  +----- Harmandia **     
                    +--+                    
                       |  +-- Unionetta **     
                       +--+                 
                          +-- Scabies **

Bolotov et al., 2024 [Publ 4369], fig. S1

tT	bi	riS	oT	roT	roS	iD
29	29	27	8	34	33	4	16

  +----------------------------- Parreysia     
  |                                            
--+                          +-- Leoparreysiini
  |  +-----------------------+                 
  |  |                       +-- Lamellidentini
  +--+                                         
     |  +----------------------- Assamnaia     
     |  |                                      
     +--+  +-------------------- Coelaturini   
        |  |                                   
        +--+                 +-- Pseudoscabies ** 
           |  +--------------+                 
           |  |              +-- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Bolotov et al., 2022 [Publ 4252], fig. S1

tT	bi	riS	oT	roT	roS	iD
27	27	25	7	34	30	4	22

  +----------------------------- Parreysiini   
--+                                            
  |  +-------------------------- Lamellidentini
  +--+                                         
     |  +----------------------- Leoparreysiini
     +--+                                      
        |  +-------------------- Coelaturini   
        +--+                                   
           |  +----------------- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Bolotov et al., 2022 [Publ 4252], fig. 2

tT	bi	riS	oT	roT	roS	iD
27	27	25	7	34	30	4	2

  +----------------------------- Lamellidentini
--+                                            
  |  +-------------------------- Parreysiini   
  +--+                                         
     |  +----------------------- Leoparreysiini
     +--+                                      
        |  +-------------------- Coelaturini   
        +--+                                   
           |  +----------------- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Bolotov et al., 2022 [Publ 4239], fig. 2

tT	bi	riS	oT	roT	roS	iD
26	26	24	7	32	28	4	17

  +-------------------------- Lamellidentini
  |                                         
--+                       +-- Parreysiini   
  |  +--------------------+                 
  |  |                    +-- Leoparreysiini
  +--+                                      
     |  +-------------------- Coelaturini   
     +--+                                   
        |  +----------------- Indonaia **      
        +--+                                
           |  +-------------- Indochinella **  
           +--+                             
              |  +----------- Radiatula **     
              +--+                          
                 |  +-------- Scabiellus **    
                 +--+                       
                    |  +----- Harmandia **     
                    +--+                    
                       |  +-- Unionetta **     
                       +--+                 
                          +-- Scabies **

Bolotov et al., 2022 [Publ 4239], fig. S1

tT	bi	riS	oT	roT	roS	iD
26	26	23	7	32	28	4	4

  +----------------------------- Lamellidentini
--+                                            
  |  +-------------------------- Parreysiini   
  +--+                                         
     |  +----------------------- Leoparreysiini
     +--+                                      
        |  +-------------------- Coelaturini   
        +--+                                   
           |  +----------------- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Bolotov et al., 2022 [Publ 4239], fig. S2

tT	bi	riS	oT	roT	roS	iD
26	26	24	7	32	28	4	21

  +----------------------------- Parreysiini   
--+                                            
  |  +-------------------------- Lamellidentini
  +--+                                         
     |  +----------------------- Leoparreysiini
     +--+                                      
        |  +-------------------- Coelaturini   
        +--+                                   
           |  +----------------- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
21	21	19	7	97	20	4	18

  +-------------------------- Lamellidentini
  |                                         
--+                       +-- Parreysiini   
  |  +--------------------+                 
  |  |                    +-- Leoparreysiini
  +--+                                      
     |  +-------------------- Coelaturini   
     +--+                                   
        |  +----------------- Indonaia **      
        +--+                                
           |  +-------------- Indochinella **  
           +--+                             
              |  +----------- Radiatula **     
              +--+                          
                 |  +-------- Scabiellus **    
                 +--+                       
                    |  +----- Scabies **       
                    +--+                    
                       |  +-- Unionetta **     
                       +--+                 
                          +-- Harmandia **

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
20	20	17	7	92	20	4	16

  +----------------------------- Lamellidentini         
  |                                                     
--+                          +-- Parreysiini            
  |  +-----------------------+                          
  |  |                       +-- Leoparreysiini         
  +--+                                                  
     |  +----------------------- Coelaturini            
     |  |                                               
     +--+  +-------------------- Indonaia **               
        |  |                                            
        +--+  +----------------- Indochinella **           
           |  |                                         
           |  |           +----- Radiatula_mouhoti **      
           +--+  +--------+                             
              |  |        |  +-- Radiatula_myitkyinae **   
              |  |        +--+                          
              +--+           +-- Radiatula_chaudhurii **   
                 |                                      
                 |  +----------- Radiatula_crispisulcata **
                 +--+                                   
                    |  +-------- Scabiellus **             
                    +--+                                
                       |  +----- Harmandia **              
                       +--+                             
                          |  +-- Unionetta **              
                          +--+                          
                             +-- Scabies **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

tT	bi	riS	oT	roT	roS	iD
18	18	16	7	5	5	4	21

  +----------------------------- Lamellidentini
--+                                            
  |  +-------------------------- Leoparreysiini
  +--+                                         
     |  +----------------------- Parreysiini   
     +--+                                      
        |  +-------------------- Coelaturini   
        +--+                                   
           |  +----------------- Indonaia **      
           +--+                                
              |  +-------------- Indochinella **  
              +--+                             
                 |  +----------- Radiatula **     
                 +--+                          
                    |  +-------- Scabiellus **    
                    +--+                       
                       |  +----- Harmandia **     
                       +--+                    
                          |  +-- Unionetta **     
                          +--+                 
                             +-- Scabies **

Pfeiffer et al., 2018 [Publ 3827], fig. 2

tT	bi	riS	oT	roT	roS	iD
17	17	15	7	11	10	4	16

  +-------------------------------- Lamellidentini       
--+                                                      
  |  +----------------------------- Parreysiini          
  +--+                                                   
     |  +-------------------------- Leoparreysiini       
     +--+                                                
        |  +----------------------- Coelaturini          
        +--+                                             
           |  +-------------------- Indonaia **             
           +--+                                          
              |  +----------------- Indochinella **         
              +--+                                       
                 |  +-------------- Radiatula **            
                 +--+                                    
                    |  +----------- Scabiellus **           
                    +--+                                 
                       |  +-------- Unionetta_fabagina__2 **
                       +--+                              
                          |  +----- Harmandia **            
                          +--+                           
                             |  +-- Unionetta_fabagina__1 **
                             +--+                        
                                +-- Scabies **

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
14	14	10	7	68	0	0	23

  +----------------- Indonaia **    
--+                              
  |  +-------------- Indochinella **
  +--+                           
     |  +----------- Radiatula **   
     +--+                        
        |  +-------- Scabiellus **  
        +--+                     
           |  +----- Scabies **     
           +--+                  
              |  +-- Unionetta **   
              +--+               
                 +-- Harmandia **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
12	12	9	4	187	22	4	24

  +-------------------- Lamellidentini
--+                                   
  |  +----------------- Parreysiini   
  +--+                                
     |  +-------------- Leoparreysiini
     +--+                             
        |  +----------- Coelaturini   
        +--+                          
           |  +-------- Indonaia **      
           +--+                       
              |  +----- Indochinella **  
              +--+                    
                 |  +-- Scabies **       
                 +--+                 
                    +-- Radiatula **

Bolotov et al., 2022 [Publ 4250], fig. S2

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	3	3	2	21

        +-- Parreysiini   
  +-----+                 
  |     +-- Leoparreysiini
--+                       
  |  +----- Indonaia **      
  +--+                    
     |  +-- Indochinella **  
     +--+                 
        +-- Radiatula **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
12	12	9	4	40	0	0	22

  +-------- Indonaia **    
--+                     
  |  +----- Indochinella **
  +--+                  
     |  +-- Scabies **     
     +--+               
        +-- Radiatula **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
12	12	9	4	189	22	4	26

  +-------------------- Lamellidentini
--+                                   
  |  +----------------- Parreysiini   
  +--+                                
     |  +-------------- Leoparreysiini
     +--+                             
        |  +----------- Coelaturini   
        +--+                          
           |  +-------- Indonaia **      
           +--+                       
              |  +----- Indochinella **  
              +--+                    
                 |  +-- Scabies **       
                 +--+                 
                    +-- Radiatula **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
12	12	9	4	189	22	4	24

  +----------------- Parreysiini   
  |                                
--+----------------- Lamellidentini
  |                                
  |  +-------------- Leoparreysiini
  +--+                             
     |  +----------- Coelaturini   
     +--+                          
        |  +-------- Indonaia **      
        +--+                       
           |  +----- Indochinella **  
           +--+                    
              |  +-- Scabies **       
              +--+                 
                 +-- Radiatula **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
12	13	9	4	113	26	4	24

           +-------------- Trapezidens             
           |                                       
  +--------+  +----------- Lamellidens_lamellatus  
  |        |  |                                    
  |        +--+  +-------- Lamellidens_jenkinsianus
  |           |  |                                 
  |           |  |  +----- Lamellidens_generosus   
  |           +--+  |                              
  |              |  +----- Lamellidens_mainwaringi 
  |              |  |                              
  |              +--+  +-- Lamellidens_ferrugineus 
--+                 +--+                           
  |                 |  +-- Lamellidens_marginalis  
  |                 |                              
  |                 |  +-- Lamellidens_savadiensis 
  |                 +--+                           
  |                    +-- Lamellidens_corrianus__1
  |                                                
  |  +-------------------- Parreysiini             
  |  |                                             
  +--+  +----------------- Leoparreysiini          
     |  |                                          
     +--+  +-------------- Coelaturini             
        |  |                                       
        |  |        +----- Indochinella **            
        +--+  +-----+                              
           |  |     |  +-- Scabies **                 
           |  |     +--+                           
           +--+        +-- Radiatula **               
              |                                    
              |  +-------- Indonaia_shurtleffiana **  
              |  |                                 
              +--+  +----- Indonaia_rugosa **         
                 |  |                              
                 |  |  +-- Indonaia_andersoniana **   
                 +--+--+                           
                    |  +-- Indonaia_caerulea **       
                    |                              
                    |  +-- Indonaia_bonneaudii **     
                    +--+                           
                       +-- Lamellidens_corrianus__2

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
11	11	9	4	128	18	4	28

  +-------------------------- Lamellidentini    
  |                                             
--+  +----------------------- Parreysiini       
  |  |                                          
  +--+  +-------------------- Leoparreysiini    
     |  |                                       
     +--+  +----------------- Coelaturini       
        |  |                                    
        +--+  +-------------- Indonaia **          
           |  |                                 
           +--+           +-- Indochinella **      
              |  +--------+                     
              |  |        +-- Scabies_crispata **  
              +--+                              
                 |  +-------- Radiatula **         
                 +--+                           
                    |  +----- Scabies_mandarinus **
                    +--+                        
                       |  +-- Scabies_humilis **   
                       +--+                     
                          +-- Scabies_phaselus **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
11	11	9	4	128	18	4	28

                       +-- Parreysiini       
  +--------------------+                     
  |                    +-- Lamellidentini    
--+                                          
  |  +-------------------- Leoparreysiini    
  |  |                                       
  +--+  +----------------- Coelaturini       
     |  |                                    
     +--+  +-------------- Indonaia **          
        |  |                                 
        +--+           +-- Indochinella **      
           |  +--------+                     
           |  |        +-- Scabies_crispata **  
           +--+                              
              |  +-------- Radiatula **         
              +--+                           
                 |  +----- Scabies_mandarinus **
                 +--+                        
                    |  +-- Scabies_humilis **   
                    +--+                     
                       +-- Scabies_phaselus **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
11	11	9	4	72	13	4	24

  +-------------------- Lamellidentini
--+                                   
  |  +----------------- Parreysiini   
  +--+                                
     |  +-------------- Leoparreysiini
     +--+                             
        |  +----------- Coelaturini   
        +--+                          
           |  +-------- Indonaia **      
           +--+                       
              |  +----- Indochinella **  
              +--+                    
                 |  +-- Radiatula **     
                 +--+                 
                    +-- Scabies **

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
9	9	7	3	40	15	3	22

  +-------------- Lamellidentini
--+                             
  |  +----------- Parreysiini   
  +--+                          
     |  +-------- Leoparreysiini
     +--+                       
        |  +----- Indonaia **      
        +--+                    
           |  +-- Indochinella **  
           +--+                 
              +-- Radiatula **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

tT	bi	riS	oT	roT	roS	iD
8	8	3	4	28	28	2	24

  +-------------- Leoparreysiini      
  |                                   
--+  +----------- Coelaturini         
  |  |                                
  +--+  +-------- Indonaia **            
     |  |                             
     +--+  +----- Indochinella **        
        |  |                          
        +--+  +-- Radiatula_mouhoti **   
           |  |                       
           |  +-- Radiatula_myitkyinae **
           +--+                       
              +-- Radiatula_chaudhurii **
              |                       
              +-- Scabies **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

tT	bi	riS	oT	roT	roS	iD
8	8	1	4	28	28	2	26

  +-------- Leoparreysiini      
  |                             
--+  +----- Coelaturini         
  |  |                          
  |  |  +-- Radiatula_mouhoti **   
  +--+  |                       
     |  +-- Radiatula_myitkyinae **
     |  |                       
     |  +-- Radiatula_chaudhurii **
     +--+                       
        +-- Indochinella **        
        |                       
        +-- Indonaia **            
        |                       
        +-- Scabies **

Bolotov et al., 2019 [Publ 3974], fig. 1

tT	bi	riS	oT	roT	roS	iD
6	6	4	2	12	7	2	23

  +-------- Leoparreysiini
--+                       
  |  +----- Coelaturini   
  +--+                    
     |  +-- Indochinella **  
     +--+                 
        +-- Radiatula **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
5	5	3	3	86	8	3	26

  +-------------- Lamellidentini
--+                             
  |  +----------- Leoparreysiini
  +--+                          
     |  +-------- Coelaturini   
     +--+                       
        |  +----- Indonaia **      
        +--+                    
           |  +-- Scabies **       
           +--+                 
              +-- Radiatula **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
5	5	3	3	86	8	3	28

  +-------------------- Lamellidentini      
--+                                         
  |  +----------------- Leoparreysiini      
  +--+                                      
     |  +-------------- Coelaturini         
     +--+                                   
        |  +----------- Indonaia **            
        +--+                                
           |  +-------- Radiatula_myitkyinae **
           +--+                             
              |  +----- Scabies **             
              +--+                          
                 |  +-- Radiatula_mouhoti **   
                 +--+                       
                    +-- Radiatula_chaudhurii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	53	6	3	25

  +----------------- Lamellidentini
--+                                
  |  +-------------- Leoparreysiini
  +--+                             
     |  +----------- Coelaturini   
     +--+                          
        |  +-------- Indonaia **      
        +--+                       
           |  +----- Indochinella **  
           +--+                    
              |  +-- Scabies **       
              +--+                 
                 +-- Harmandia **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	53	6	3	25

  +----------------- Lamellidentini
--+                                
  |  +-------------- Leoparreysiini
  +--+                             
     |  +----------- Coelaturini   
     +--+                          
        |  +-------- Indonaia **      
        +--+                       
           |  +----- Indochinella **  
           +--+                    
              |  +-- Scabies **       
              +--+                 
                 +-- Harmandia **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	28	8	3	26

  +-------------- Lamellidentini
--+                             
  |  +----------- Leoparreysiini
  +--+                          
     |  +-------- Coelaturini   
     +--+                       
        |  +----- Indochinella **  
        +--+                    
           |  +-- Radiatula **     
           +--+                 
              +-- Scabies **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	37	5	3	26

  +-------------- Lamellidentini
--+                             
  |  +----------- Leoparreysiini
  +--+                          
     |  +-------- Coelaturini   
     +--+                       
        |  +----- Indochinella **  
        +--+                    
           |  +-- Radiatula **     
           +--+                 
              +-- Scabies **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	53	4	3	26

  +-------------- Lamellidentini
--+                             
  |  +----------- Parreysiini   
  +--+                          
     |  +-------- Leoparreysiini
     +--+                       
        |  +----- Indonaia **      
        +--+                    
           |  +-- Indochinella **  
           +--+                 
              +-- Radiatula **

Dai et al., 2024 [Publ 4386], fig. 5-BI

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	19	1	1	--

  +-- Lamellidentini
--+                 
  +-- Scabies **

Dai et al., 2024 [Publ 4386], fig. 5-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	19	1	1	--

  +-- Lamellidentini
--+                 
  +-- Scabies **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	7	3	--

  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Coelaturini   
     +--+                    
        |  +-- Radiatula **     
        +--+                 
           +-- Indochinella **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	7	3	--

  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Coelaturini   
     +--+                    
        |  +-- Radiatula **     
        +--+                 
           +-- Indochinella **

Combosch et al., 2017 [Publ 3806], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Scabies_crispata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	121	2	2	--

  +-- Coelaturini   
  |                 
--+-- Leoparreysiini
  |                 
  +-- Radiatula **

Dai et al., 2024 [Publ 4386], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	12	0	0	--

-- Scabies_scobinatus **

Dai et al., 2024 [Publ 4385], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Lamellidentini
--+                 
  +-- Scabies **

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	2	2	--

  +----- Parreysiini
--+                 
  |  +-- Coelaturini
  +--+              
     +-- Indonaia **

Choi et al., 2020 [Publ 4169], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	2	2	--

  +----- Radiatula **     
--+                    
  |  +-- Lamellidentini
  +--+                 
     +-- Leoparreysiini

Imsong & Murali, 2023 [Publ 4370], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	2	2	--

  +----- Lamellidentini
--+                    
  |  +-- Indonaia **      
  +--+                 
     +-- Parreysiini

Sano et al., 2020 [Publ 4138], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	0	0	--

-- Scabies_crispata **

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Scabies_crispata **

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	4	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Unionetta **     
     +--+                 
        +-- Coelaturini

Chen et al., 2023 [Publ 4302], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	2	2	--

  +-- Coelaturini
  |              
--+-- Indonaia **   
  |              
  +-- Parreysiini

Wu et al., 2020 [Publ 4121], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	0	0	--

-- Scabies_crispata **

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	2	2	--

  +-- Coelaturini
  |              
--+-- Indonaia **   
  |              
  +-- Parreysiini

Xiang et al., 2025 [Publ 4433], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	0	0	--

-- Scabies_crispata **

Combosch et al., 2017 [Publ 3806], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Scabies_crispata **

Smith, 2021 [Publ 4331], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	0	0	--

-- Scabies_phaselus **

Huang et al., 2018 [Publ 3828], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	28	0	0	--

-- Scabies_crispata **

Wu et al., 2020 [Publ 4121], fig. S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	0	0	--

-- Scabies_crispata **

Dai et al., 2024 [Publ 4385], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	1	1	--

  +-- Lamellidentini
--+                 
  +-- Scabies **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	79	3	3	--

  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinella **

Sano et al., 2020 [Publ 4138], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	0	0	--

-- Scabies_crispata **

Wu et al., 2021 [Publ 4215], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	23	0	0	--

-- Indonaia_andersoniana **

Sano et al., 2020 [Publ 4138], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	0	0	--

-- Scabies_crispata **

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	2	2	--

  +----- Parreysiini
--+                 
  |  +-- Coelaturini
  +--+              
     +-- Indonaia **

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