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Unionidae

Margaritiferidae

Hyriidae

Etheriidae

Mycetopodidae

Iridinidae


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Data last updated
Mon 1 Dec 2025

Bivalvia | Unionoida | Etherioidea

family IRIDINIDAE Swainson, 1840: cladomics

2 subfamilies

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 39 (52 total)
number of subtaxa (rank: subfamily) = 2
analyzed in 73 cladograms in 41 publications
maximum number of terminals in an analysis = 5 (mean = 1.97, median = 2)
maximum number of subtaxa (subfamily) = 2 (mean = 1.33, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 38
maximum number of included terminals = 5, (mean = 2.87 median = 3)
maximum number of subtaxa (subfamily) = 2 (mean = 1.63, median = 2)
proportion of cladograms that support basic monophyly = 0.5

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 23
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subfamily) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 5 (mean = 3.35, median = 3)
maximum number of subtaxa (subfamily) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.17

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Iridinidae cladogram distances
PCoA Ordination

NJ Tree of Iridinidae cladogram distances
NJ Tree

Cladograms

Walker et al., 2006 [Publ 2666], fig. 6

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
3 9 7 2 33 6 2 0 
  +----------- Iridininae **   
--+                         
  |  +-------- Etheria      
  +--+                      
     |  +----- Aspathariinae **
     +--+                   
        |  +-- Mulleria     
        +--+                
           +-- Mycetopodidae

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 10 8 2 194 5 2 16 
           +-- Anodontites     
  +--------+                   
  |        +-- Iridininae **      
  |                            
--+        +-- Etheriidae      
  |  +-----+                   
  |  |     +-- Aspathariinae **   
  +--+                         
     |  +----- Mycetopodinae   
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Lamproscapha

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 10 8 2 196 5 2 16 
           +-- Anodontites     
  +--------+                   
  |        +-- Iridininae **      
  |                            
--+        +-- Etheriidae      
  |  +-----+                   
  |  |     +-- Aspathariinae **   
  +--+                         
     |  +----- Mycetopodinae   
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Lamproscapha

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 10 6 2 196 5 2 14 
  +-------- Etheriidae      
  |                         
  +-------- Aspathariinae **   
  |                         
--+     +-- Anodontites     
  +-----+                   
  |     +-- Iridininae **      
  |                         
  |  +----- Mycetopodinae   
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Lamproscapha

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 2 42 6 2 8 
  +-------- Etheriidae   
--+                      
  |  +----- Mycetopodidae
  +--+                   
     |  +-- Iridininae **   
     +--+                
        +-- Aspathariinae **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 2 42 6 2 8 
  +-------------- Pseudomulleria
--+                             
  |  +----------- Mulleria      
  +--+                          
     |  +-------- Etheria       
     +--+                       
        |  +----- Mycetopodidae 
        +--+                    
           |  +-- Iridininae **    
           +--+                 
              +-- Aspathariinae **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
4 4 1 2 43 6 2 8 
        +-- Chambardia **   
        |                
  +-----+-- Aspatharia **   
  |     |                
--+     +-- Iridininae **   
  |                      
  |  +----- Etheria      
  +--+                   
     |  +-- Mulleria     
     +--+                
        +-- Mycetopodidae

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 2 43 6 2 8 
  +-------- Etheriidae   
--+                      
  |  +----- Mycetopodidae
  +--+                   
     |  +-- Aspathariinae **
     +--+                
        +-- Iridininae **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 2 24 0 0 7 
  +-- Iridininae **   
--+                
  +-- Aspathariinae **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
3 6 4 2 12 3 2 13 
  +-------- Aspathariinae **
--+                      
  |  +----- Etheriidae   
  +--+                   
     |  +-- Iridininae **   
     +--+                
        +-- Mycetopodidae

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
3 6 4 2 12 3 2 13 
  +-------- Aspathariinae **
--+                      
  |  +----- Etheriidae   
  +--+                   
     |  +-- Iridininae **   
     +--+                
        +-- Mycetopodidae

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
3 6 4 2 25 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 6 4 2 65 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
3 6 4 2 31 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
3 6 4 2 25 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
3 6 4 2 26 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
3 6 4 2 31 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 6 4 2 65 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 6 4 2 65 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
3 5 3 2 33 2 1 12 
  +----- Aspathariinae **
--+                   
  |  +-- Iridininae **   
  +--+                
     +-- Mycetopodidae

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
3 5 3 2 33 2 1 12 
  +----- Aspathariinae **
--+                   
  |  +-- Iridininae **   
  +--+                
     +-- Mycetopodidae

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
3 6 4 2 77 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
2 7 0 2 39 5 1 9 
  +----- Iridininae **   
--+                   
  |  +-- Aspathariinae **
  +--+                
     +-- Mycetopodidae

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
2 2 0 1 120 3 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 24 3 2 -- 
  +----- Aspathariinae **
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Mycetopodidae

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 28 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
2 2 0 1 29 3 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 32 6 2 -- 
  +-------- Iridininae **   
--+                      
  |  +----- Etheria      
  +--+                   
     |  +-- Mulleria     
     +--+                
        +-- Mycetopodidae

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
2 2 0 1 89 3 2 -- 
  +----- Aspathariinae **
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Mycetopodidae

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 1 137 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
2 7 0 2 39 5 1 11 
  +----- Aspathariinae **
--+                   
  |  +-- Iridininae **   
  +--+                
     +-- Mycetopodidae

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 1 137 2 2 -- 
  +----- Aspathariinae **
--+                   
  |  +-- Mycetopodidae
  +--+                
     +-- Etheriidae

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 3 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
2 2 0 1 29 3 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 1 89 3 2 -- 
  +----- Aspathariinae **
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Mycetopodidae

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 28 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
2 2 0 1 29 3 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
2 2 0 1 29 3 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Mutela_rostrata **

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 2 2 -- 
  +----- Aspathariinae **
--+                   
  |  +-- Mycetopodidae
  +--+                
     +-- Etheriidae

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 2 2 -- 
  +----- Etheriidae   
--+                   
  |  +-- Mycetopodidae
  +--+                
     +-- Aspathariinae **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Mutela_rostrata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

González & Giribet, 2015 [Publ 3632], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Aspatharia_pfeifferiana **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Mutela_dubia **

Elderkin et al., 2016 [Publ 3693], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 1 1 -- 
-- Aspatharia_pfeifferiana **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Mutela_rostrata **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Mutela_rostrata **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Bieler et al., 2014 [Publ 3622], fig. 28

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Aspatharia_pfeifferiana **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 1 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

González & Giribet, 2015 [Publ 3632], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Aspatharia_pfeifferiana **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Etheriidae   
--+                
  +-- Aspathariinae **

González & Giribet, 2015 [Publ 3632], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Aspatharia_pfeifferiana **

Lemer et al., 2019 [Publ 4093], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Etheriidae   
--+                
  +-- Aspathariinae **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 1 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

González & Giribet, 2015 [Publ 3632], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Aspatharia_pfeifferiana **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 4 2 -- 
  +----- Iridininae **   
  |                   
--+  +-- Anodontitinae
  |  |                
  +--+-- Mycetopodinae
     |                
     +-- Etheriidae

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 1 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Aspatharia_pfeifferiana **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **
 
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