Bivalvia | Unionida | Etherioidea

family IRIDINIDAE Swainson, 1840: cladomics

2 subfamilies

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 39 (52 total)
number of subtaxa (rank: subfamily) = 2
analyzed in 73 cladograms in 41 publications
maximum number of terminals in an analysis = 5 (mean = 1.97, median = 2)
maximum number of subtaxa (subfamily) = 2 (mean = 1.33, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 38
maximum number of included terminals = 5, (mean = 2.87 median = 3)
maximum number of subtaxa (subfamily) = 2 (mean = 1.63, median = 2)
proportion of cladograms that support basic monophyly = 0.5

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 23
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subfamily) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 5 (mean = 3.35, median = 3)
maximum number of subtaxa (subfamily) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.17

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Iridinidae cladogram distances
PCoA Ordination

NJ Tree of Iridinidae cladogram distances
NJ Tree

Cladograms

Walker et al., 2006 [Publ 2666], fig. 6

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
3 9 7 2 33 6 2 0 
  +----------- Iridininae **   
--+                         
  |  +-------- Etheria      
  +--+                      
     |  +----- Aspathariinae **
     +--+                   
        |  +-- Mulleria     
        +--+                
           +-- Mycetopodidae

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 10 8 2 194 5 2 16 
           +-- Anodontites     
  +--------+                   
  |        +-- Iridininae **      
  |                            
--+        +-- Etheriidae      
  |  +-----+                   
  |  |     +-- Aspathariinae **   
  +--+                         
     |  +----- Mycetopodinae   
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Lamproscapha

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 10 8 2 196 5 2 16 
           +-- Anodontites     
  +--------+                   
  |        +-- Iridininae **      
  |                            
--+        +-- Etheriidae      
  |  +-----+                   
  |  |     +-- Aspathariinae **   
  +--+                         
     |  +----- Mycetopodinae   
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Lamproscapha

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 10 6 2 196 5 2 14 
  +-------- Etheriidae      
  |                         
  +-------- Aspathariinae **   
  |                         
--+     +-- Anodontites     
  +-----+                   
  |     +-- Iridininae **      
  |                         
  |  +----- Mycetopodinae   
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Lamproscapha

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
4 4 1 2 43 6 2 8 
        +-- Chambardia **   
        |                
  +-----+-- Aspatharia **   
  |     |                
--+     +-- Iridininae **   
  |                      
  |  +----- Etheria      
  +--+                   
     |  +-- Mulleria     
     +--+                
        +-- Mycetopodidae

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 2 24 0 0 7 
  +-- Iridininae **   
--+                
  +-- Aspathariinae **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 2 43 6 2 8 
  +-------- Etheriidae   
--+                      
  |  +----- Mycetopodidae
  +--+                   
     |  +-- Aspathariinae **
     +--+                
        +-- Iridininae **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 2 42 6 2 8 
  +-------- Etheriidae   
--+                      
  |  +----- Mycetopodidae
  +--+                   
     |  +-- Iridininae **   
     +--+                
        +-- Aspathariinae **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 2 42 6 2 8 
  +-------------- Pseudomulleria
--+                             
  |  +----------- Mulleria      
  +--+                          
     |  +-------- Etheria       
     +--+                       
        |  +----- Mycetopodidae 
        +--+                    
           |  +-- Iridininae **    
           +--+                 
              +-- Aspathariinae **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
3 6 4 2 25 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
3 6 4 2 77 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
3 6 4 2 31 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 6 4 2 65 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
3 6 4 2 12 3 2 13 
  +-------- Aspathariinae **
--+                      
  |  +----- Etheriidae   
  +--+                   
     |  +-- Iridininae **   
     +--+                
        +-- Mycetopodidae

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
3 5 3 2 33 2 1 12 
  +----- Aspathariinae **
--+                   
  |  +-- Iridininae **   
  +--+                
     +-- Mycetopodidae

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 6 4 2 65 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
3 6 4 2 12 3 2 13 
  +-------- Aspathariinae **
--+                      
  |  +----- Etheriidae   
  +--+                   
     |  +-- Iridininae **   
     +--+                
        +-- Mycetopodidae

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
3 6 4 2 31 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 6 4 2 65 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
3 5 3 2 33 2 1 12 
  +----- Aspathariinae **
--+                   
  |  +-- Iridininae **   
  +--+                
     +-- Mycetopodidae

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
3 6 4 2 26 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
3 6 4 2 25 3 2 11 
  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 28 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
2 7 0 2 39 5 1 11 
  +----- Aspathariinae **
--+                   
  |  +-- Iridininae **   
  +--+                
     +-- Mycetopodidae

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
2 2 0 1 120 3 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 1 137 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
2 2 0 1 89 3 2 -- 
  +----- Aspathariinae **
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Mycetopodidae

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 29 3 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
2 7 0 2 39 5 1 9 
  +----- Iridininae **   
--+                   
  |  +-- Aspathariinae **
  +--+                
     +-- Mycetopodidae

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 1 89 3 2 -- 
  +----- Aspathariinae **
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Mycetopodidae

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
2 2 0 1 29 3 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
2 2 0 1 29 3 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
2 2 0 1 29 3 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 24 3 2 -- 
  +----- Aspathariinae **
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Mycetopodidae

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
2 2 0 1 29 3 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 1 137 2 2 -- 
  +----- Aspathariinae **
--+                   
  |  +-- Mycetopodidae
  +--+                
     +-- Etheriidae

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 32 6 2 -- 
  +-------- Iridininae **   
--+                      
  |  +----- Etheria      
  +--+                   
     |  +-- Mulleria     
     +--+                
        +-- Mycetopodidae

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 28 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Mutela_rostrata **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 1 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

González & Giribet, 2015 [Publ 3632], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Aspatharia_pfeifferiana **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Mutela_rostrata **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 2 2 -- 
  +----- Etheriidae   
--+                   
  |  +-- Mycetopodidae
  +--+                
     +-- Aspathariinae **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

González & Giribet, 2015 [Publ 3632], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Aspatharia_pfeifferiana **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Etheriidae   
--+                
  +-- Aspathariinae **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 1 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Lemer et al., 2019 [Publ 4093], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Etheriidae   
--+                
  +-- Aspathariinae **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 4 2 -- 
  +----- Iridininae **   
  |                   
--+  +-- Anodontitinae
  |  |                
  +--+-- Mycetopodinae
     |                
     +-- Etheriidae

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Aspatharia_pfeifferiana **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

González & Giribet, 2015 [Publ 3632], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Aspatharia_pfeifferiana **

González & Giribet, 2015 [Publ 3632], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Aspatharia_pfeifferiana **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Bieler et al., 2014 [Publ 3622], fig. 28

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Aspatharia_pfeifferiana **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 2 2 -- 
  +----- Aspathariinae **
--+                   
  |  +-- Mycetopodidae
  +--+                
     +-- Etheriidae

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Mutela_rostrata **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Mutela_rostrata **

Elderkin et al., 2016 [Publ 3693], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 2 1 1 -- 
-- Aspatharia_pfeifferiana **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Mutela_dubia **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Mycetopodidae
--+                
  +-- Iridininae **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 1 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Iridininae **   
--+                
  +-- Mycetopodidae