Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini

subtribe MIDDENDORFFINAIINA Lopes-Lima, Bolotov & Bogan in Lopes-Lima et al., 2020: cladomics

1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 1 (1 total)
number of subtaxa (rank: genus) = 1
analyzed in 19 cladograms in 12 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Cladograms

Dai et al., 2025 [Publ 4440], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
1 1 0 1 44 21 4 -- 
  +----------------------- Globunionina    
  |                                        
--+  +-------------------- Aculamprotulina 
  |  |                                     
  +--+  +----------------- Middendorffinaia **
     |  |                                  
     +--+  +-------------- Nodularia       
        |  |                               
        +--+  +----------- Cuneopsis       
           |  |                            
           |  |        +-- Diaurora        
           +--+--------+                   
              |        +-- Schistodesmus   
              |                            
              |  +-------- Pseudocuneopsis 
              +--+                         
                 |  +----- Unionina        
                 +--+                      
                    |  +-- Tchangsinaia    
                    +--+                   
                       +-- Inversiunio

Hou et al., 2025 [Publ 4432], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 10 9 1 -- 
  +-------- Tchangsinaia    
--+                         
  |  +----- Pseudocuneopsis 
  +--+                      
     |  +-- Middendorffinaia **
     +--+                   
        +-- Cuneopsis

Hou et al., 2025 [Publ 4432], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 27 25 1 -- 
  +-- Middendorffinaia **
--+                   
  +-- Oxynaiina

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 11 3 -- 
  +-------- Aculamprotulina 
--+                         
  |  +----- Unionina        
  +--+                      
     |  +-- Middendorffinaia **
     +--+                   
        +-- Oxynaiina

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 35 4 2 -- 
  +----- Middendorffinaia **
--+                      
  |  +-- Oxynaiina       
  +--+                   
     +-- Unionina

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

iTtTbiriSoTroTroSiD
1 1 0 1 9 7 1 -- 
  +-- Middendorffinaia **
--+                   
  +-- Oxynaiina

Klishko et al., 2019 [Publ 3886], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 10 7 2 -- 
  +----- Unionina        
--+                      
  |  +-- Middendorffinaia **
  +--+                   
     +-- Oxynaiina

Hou et al., 2025 [Publ 4401], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 27 25 1 -- 
  +-- Middendorffinaia **
--+                   
  +-- Oxynaiina

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 64 32 3 -- 
  +-------- Aculamprotulina 
--+                         
  |  +----- Unionina        
  +--+                      
     |  +-- Middendorffinaia **
     +--+                   
        +-- Oxynaiina

Sayenko et al., 2021 [Publ 4189], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 12 12 2 -- 
  +-------- Unionina        
--+                         
  |  +----- Inversiunio     
  +--+                      
     |  +-- Middendorffinaia **
     +--+                   
        +-- Nodularia

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 17 3 -- 
  +----------------- Aculamprotulina 
  |                                  
  |              +-- Middendorffinaia **
--+  +-----------+                   
  |  |           +-- Nodularia       
  |  |                               
  +--+  +----------- Cuneopsis       
     |  |                            
     |  |        +-- Diaurora        
     +--+--------+                   
        |        +-- Schistodesmus   
        |                            
        |  +-------- Pseudocuneopsis 
        +--+                         
           |  +----- Inversiunio     
           +--+                      
              |  +-- Tchangsinaia    
              +--+                   
                 +-- Unionina

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
1 1 0 1 55 11 3 -- 
  +-------- Middendorffinaia **
--+                         
  |  +----- Aculamprotulina 
  +--+                      
     |  +-- Unionina        
     +--+                   
        +-- Oxynaiina

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 12 1 -- 
           +----- Cuneopsis_rufescens   
  +--------+                            
  |        |  +-- Middendorffinaia **      
  |        +--+                         
--+           +-- Pseudocuneopsis       
  |                                     
  |  +----------- Tchangsinaia          
  |  |                                  
  +--+        +-- Cuneopsis_kiangsiensis
     |  +-----+                         
     |  |     +-- Cuneopsis_celtiformis 
     +--+                               
        |  +----- Cuneopsis_szechenyii  
        +--+                            
           |  +-- Cuneopsis_heudei      
           +--+                         
              +-- Cuneopsis_demangei

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 27 3 -- 
  +-------- Aculamprotulina 
--+                         
  |  +----- Middendorffinaia **
  +--+                      
     |  +-- Unionina        
     +--+                   
        +-- Oxynaiina

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 75 48 3 -- 
  +-------- Aculamprotulina 
--+                         
  |  +----- Unionina        
  +--+                      
     |  +-- Middendorffinaia **
     +--+                   
        +-- Oxynaiina

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 17 3 -- 
  +----------------- Aculamprotulina 
  |                                  
  |              +-- Middendorffinaia **
--+  +-----------+                   
  |  |           +-- Nodularia       
  |  |                               
  +--+  +----------- Cuneopsis       
     |  |                            
     |  |        +-- Diaurora        
     +--+--------+                   
        |        +-- Schistodesmus   
        |                            
        |  +-------- Pseudocuneopsis 
        +--+                         
           |  +----- Inversiunio     
           +--+                      
              |  +-- Tchangsinaia    
              +--+                   
                 +-- Unionina

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 11 3 -- 
  +----- Unionina        
--+                      
  |  +-- Middendorffinaia **
  +--+                   
     +-- Oxynaiina

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 21 4 -- 
  +-------------------- Globunionina    
  |                                     
--+  +----------------- Aculamprotulina 
  |  |                                  
  |  |              +-- Middendorffinaia **
  +--+  +-----------+                   
     |  |           +-- Nodularia       
     |  |                               
     +--+  +----------- Cuneopsis       
        |  |                            
        |  |        +-- Diaurora        
        +--+--------+                   
           |        +-- Schistodesmus   
           |                            
           |  +-------- Pseudocuneopsis 
           +--+                         
              |  +----- Inversiunio     
              +--+                      
                 |  +-- Tchangsinaia    
                 +--+                   
                    +-- Unionina

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 13 10 3 -- 
  +-- Middendorffinaia **
--+                   
  +-- Oxynaiina