MUSSELp
 
MUSSELpdb
  Introduction
Browse
d'basing
 
Taxonomy
  all genera
Unionidae
Margaritiferidae
Hyriidae
Etheriidae
Mycetopodidae
Iridinidae

Species & Genus Richness by Taxonomy
 
Geography
  Summary
North America
Central America
South America
Afrotropics
North Eurasia
East Asia
Australasia

Species Richness & Endemism by Geography
 
 

Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae

tribe PARREYSIINI Henderson, 1935: cladomics
     2 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 18 (18 total)
number of subtaxa (rank: genus) = 2
analyzed in 29 cladograms in 17 publications
maximum number of terminals in an analysis = 7 (mean = 3, median = 3)
maximum number of subtaxa (genus) = 2 (mean = 1.07, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 19
maximum number of included terminals = 7, (mean = 4.05 median = 4)
maximum number of subtaxa (genus) = 2 (mean = 1.11, median = 1)
proportion of cladograms that support basic monophyly = 0.95

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 2
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 7 (mean = 7, median = 7)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.5

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Bolotov et al., 2024 [Publ 4369], fig. S1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
762602565540
  +----------- Parreysia **     
  |                          
--+        +-- Leoparreysiini
  |  +-----+                 
  |  |     +-- Lamellidentini
  +--+                       
     |  +----- Assamnaia **     
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2024 [Publ 4369], fig. 2

iTtTbiriSoTroTroSiD
77525555459
  +----------- Lamellidentini
  |                          
--+        +-- Coelaturini   
  |  +-----+                 
  |  |     +-- Indochinellini
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Assamnaia **     
        +--+                 
           +-- Parreysia **

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
55315349457
  +----------- Parreysia **     
--+                          
  |  +-------- Lamellidentini
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
55015652460
  +----------- Parreysia **     
--+                          
  |  +-------- Lamellidentini
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
55315652459
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
55315349459
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
55315349461
  +-------- Lamellidentini
  |                       
--+     +-- Parreysia **     
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
553110735457
  +-------- Lamellidentini
  |                       
--+     +-- Parreysia **     
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
553111336461
  +-------- Lamellidentini
  |                       
--+     +-- Parreysia **     
  |  +--+                 
  |  |  +-- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
442112134460
           +-------------- Trapezidens             
           |                                       
  +--------+  +----------- Lamellidens_lamellatus  
  |        |  |                                    
  |        +--+  +-------- Lamellidens_jenkinsianus
  |           |  |                                 
  |           |  |  +----- Lamellidens_generosus   
  |           +--+  |                              
  |              |  +----- Lamellidens_mainwaringi 
  |              |  |                              
  |              +--+  +-- Lamellidens_ferrugineus 
--+                 +--+                           
  |                 |  +-- Lamellidens_marginalis  
  |                 |                              
  |                 |  +-- Lamellidens_savadiensis 
  |                 +--+                           
  |                    +-- Lamellidens_corrianus__1
  |                                                
  |  +-------------------- Parreysia **               
  |  |                                             
  +--+  +----------------- Leoparreysiini          
     |  |                                          
     +--+  +-------------- Coelaturini             
        |  |                                       
        |  |        +----- Indochinella            
        +--+  +-----+                              
           |  |     |  +-- Scabies                 
           |  |     +--+                           
           +--+        +-- Radiatula               
              |                                    
              |  +-------- Indonaia_shurtleffiana  
              |  |                                 
              +--+  +----- Indonaia_rugosa         
                 |  |                              
                 |  |  +-- Indonaia_andersoniana   
                 +--+--+                           
                    |  +-- Indonaia_caerulea       
                    |                              
                    |  +-- Indonaia_bonneaudii     
                    +--+                           
                       +-- Lamellidens_corrianus__2

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
331119631459
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
330119631460
  +-------- Parreysia **     
  |                       
--+-------- Lamellidentini
  |                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
330113626460
        +-- Parreysia **     
  +-----+                 
  |     +-- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelaturini   
     +--+                 
        +-- Indochinellini

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
330113626460
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
33112524459
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
331119431459
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Lyubas et al., 2022 [Publ 4255], fig. S1

iTtTbiriSoTroTroSiD
22012900--
1

Lyubas et al., 2022 [Publ 4255], fig. 7

iTtTbiriSoTroTroSiD
22012900--
1

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
220181224--
  +----------- Lamellidentini
--+                          
  |  +-------- Parreysia **     
  +--+                       
     |  +----- Leoparreysiini
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. S2

iTtTbiriSoTroTroSiD
110114142--
  +----- Indochinellini
--+                    
  |  +-- Parreysia **     
  +--+                 
     +-- Leoparreysiini

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
11014422--
  +-- Coelaturini   
  |                 
--+-- Indochinellini
  |                 
  +-- Parreysia **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
11011611--
-- Parreysia **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015563--
  +-------- Lamellidentini
--+                       
  |  +----- Parreysia **     
  +--+                    
     |  +-- Leoparreysiini
     +--+                 
        +-- Indochinellini

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
11014422--
  +----- Parreysia **     
--+                    
  |  +-- Coelaturini   
  +--+                 
     +-- Indochinellini

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11013922--
  +----- Parreysia **     
--+                    
  |  +-- Coelaturini   
  +--+                 
     +-- Indochinellini

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
110122224--
  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Parreysia **     
     +--+                    
        |  +-- Coelaturini   
        +--+                 
           +-- Indochinellini

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
110148233--
  +-------- Lamellidentini
--+                       
  |  +----- Parreysia **     
  +--+                    
     |  +-- Leoparreysiini
     +--+                 
        +-- Indochinellini

Imsong & Murali, 2023 [Publ 4370], fig. 2

iTtTbiriSoTroTroSiD
1101222--
  +----- Lamellidentini
--+                    
  |  +-- Indochinellini
  +--+                 
     +-- Parreysia **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11013922--
  +-- Coelaturini   
  |                 
--+-- Indochinellini
  |                 
  +-- Parreysia **
 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."