Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Rectidentini

subtribe RECTIDENTINA Modell, 1942: cladomics

5 genera

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 21 (21 total)
number of subtaxa (rank: genus) = 5
analyzed in 61 cladograms in 40 publications
maximum number of terminals in an analysis = 16 (mean = 5.25, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.46, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 58
maximum number of included terminals = 16, (mean = 5.47 median = 4)
maximum number of subtaxa (genus) = 3 (mean = 2.53, median = 3)
proportion of cladograms that support basic monophyly = 0.95

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 54
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 16 (mean = 5.41, median = 3.5)
maximum number of subtaxa (genus) = 3 (mean = 2.57, median = 3)
proportion of cladograms that support rigorous monophyly = 0.94

Cladograms

Wu et al., 2024 [Publ 4350], fig. 5b

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
4 13 11 3 33 9 2 0 
  +----------- Hyriopsis_bialata **  
  |                               
--+        +-- Rectidens **          
  |  +-----+                      
  |  |     +-- Ctenodesmina       
  +--+                            
     |  +----- Ensidens **           
     +--+                         
        |  +-- Hyriopsis_desowitzi **
        +--+                      
           +-- Contradentina

Zieritz et al., 2020 [Publ 4102], fig. 4

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
11 11 7 3 20 15 2 19 
  +----------------- Contradentina        
  |                                       
--+  +-------------- Ctenodesmina         
  |  |                                    
  +--+  +----------- Ensidens **             
     |  |                                 
     +--+  +-------- Hyriopsis_desowitzi **  
        |  |                              
        |  +-------- Rectidens **            
        +--+                              
           |  +----- Hyriopsis_panhai **     
           |  |                           
           +--+  +-- Hyriopsis_khoratensis **
              |  |                        
              +--+-- Hyriopsis_bialata **    
                 |                        
                 +-- Hyriopsis_myersiana **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
16 16 14 3 67 19 1 25 
  +-------------------------- Contradentina          
  |                                                  
--+  +----------------------- Ensidens **               
  |  |                                               
  +--+                    +-- Hyriopsis_desowitzi **    
     |  +-----------------+                          
     |  |                 +-- Hyriopsis_delaportei **   
     +--+                                            
        |  +----------------- Rectidens **              
        |  |                                         
        +--+  +-------------- Hyriopsis_panhai **       
           |  |                                      
           +--+  +----------- Hyriopsis_myersiana **    
              |  |                                   
              +--+        +-- Hyriopsis_khoratensis **  
                 |  +-----+                          
                 |  |     +-- Hyriopsis_bialata **      
                 +--+                                
                    |  +----- Hyriopsis_sakhonensis **  
                    +--+                             
                       |  +-- Hyriopsis_phuphaniensis **
                       +--+                          
                          +-- Hyriopsis_kratiensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
13 13 11 3 16 15 1 22 
  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Rectidens **    
     +--+                
        +-- Hyriopsis **

Konopleva et al., 2019 [Publ 4041], fig. 3

iTtTbiriSoTroTroSiD
12 12 0 3 22 19 1 19 
  +----- Contradentina        
  |                           
  |  +-- Ensidens_ingallsianus **
  |  |                        
  |  +-- Ensidens_telus **       
  |  |                        
  |  +-- Ensidens_dugasti **     
--+  |                        
  |  +-- Hyriopsis_khoratensis **
  |  |                        
  |  +-- Hyriopsis_bialata **    
  |  |                        
  |  +-- Hyriopsis_myersiana **  
  |  |                        
  +--+-- Hyriopsis_desowitzi **  
     |                        
     +-- Rectidens **            
     |                        
     +-- Hyriopsis_panhai **     
     |                        
     +-- Ensidens_jaculus__1 **  
     |                        
     +-- Ensidens_jaculus__2 **  
     |                        
     +-- Ensidens_spiculus **    
     |                        
     +-- Ensidens_sagittarius **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
12 12 10 3 106 23 1 21 
  +----------------- Contradentina        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+  +----------- Hyriopsis_desowitzi **  
     |  |                                 
     +--+  +-------- Rectidens **            
        |  |                              
        +--+     +-- Hyriopsis_khoratensis **
           |  +--+                        
           |  |  +-- Hyriopsis_bialata **    
           +--+                           
              |  +-- Hyriopsis_myersiana **  
              +--+                        
                 +-- Hyriopsis_panhai **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
12 12 10 3 100 23 1 21 
  +-------------------- Contradentina        
--+                                          
  |  +----------------- Ensidens **             
  +--+                                       
     |  +-------------- Hyriopsis_desowitzi **  
     +--+                                    
        |  +----------- Rectidens **            
        +--+                                 
           |  +-------- Hyriopsis_panhai **     
           +--+                              
              |  +----- Hyriopsis_myersiana **  
              +--+                           
                 |  +-- Hyriopsis_khoratensis **
                 +--+                        
                    +-- Hyriopsis_bialata **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
11 11 7 3 114 12 1 19 
  +-------------- Contradentina        
  |                                    
--+  +----------- Ensidens **             
  |  |                                 
  +--+  +-------- Hyriopsis_desowitzi **  
     |  |                              
     |  +-------- Rectidens **            
     +--+                              
        |  +----- Hyriopsis_panhai **     
        |  |                           
        +--+  +-- Hyriopsis_myersiana **  
           |  |                        
           +--+-- Hyriopsis_bialata **    
              |                        
              +-- Hyriopsis_khoratensis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
11 11 5 3 190 12 1 18 
  +-------- Contradentina        
  |                              
  |  +----- Hyriopsis_desowitzi **  
--+  |                           
  |  +----- Rectidens **            
  |  |                           
  +--+----- Ensidens **             
     |                           
     |  +-- Hyriopsis_bialata **    
     |  |                        
     |  +-- Hyriopsis_myersiana **  
     +--+                        
        +-- Hyriopsis_panhai **     
        |                        
        +-- Hyriopsis_khoratensis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
11 11 6 3 190 12 1 19 
  +-------- Contradentina        
  |                              
  |  +----- Hyriopsis_desowitzi **  
--+  |                           
  |  +----- Hyriopsis_myersiana **  
  |  |                           
  |  +----- Hyriopsis_panhai **     
  +--+                           
     |  +-- Hyriopsis_bialata **    
     +--+                        
     |  +-- Hyriopsis_khoratensis **
     |                           
     |  +-- Rectidens **            
     +--+                        
        +-- Ensidens **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
11 11 7 3 128 11 1 17 
  +----------- Contradentina        
  |                                 
  |        +-- Hyriopsis_desowitzi **  
--+        |                        
  |  +-----+-- Rectidens **            
  |  |     |                        
  |  |     +-- Ensidens **             
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+----- Hyriopsis_khoratensis **
        |                           
        |  +-- Hyriopsis_myersiana **  
        +--+                        
           +-- Hyriopsis_bialata **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
11 11 8 3 41 16 1 19 
  +----------------- Contradentina        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+  +----------- Hyriopsis_desowitzi **  
     |  |                                 
     +--+----------- Rectidens **            
        |                                 
        |  +-------- Hyriopsis_panhai **     
        +--+                              
           |  +----- Hyriopsis_myersiana **  
           +--+                           
              |  +-- Hyriopsis_khoratensis **
              +--+                        
                 +-- Hyriopsis_bialata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
11 11 9 3 128 12 1 20 
  +----------------- Contradentina        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+           +-- Hyriopsis_bialata **    
     |  +--------+                        
     |  |        +-- Hyriopsis_khoratensis **
     +--+                                 
        |  +-------- Hyriopsis_panhai **     
        +--+                              
           |  +----- Hyriopsis_myersiana **  
           +--+                           
              |  +-- Rectidens **            
              +--+                        
                 +-- Hyriopsis_desowitzi **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
11 11 9 3 188 12 1 20 
  +----------------- Contradentina        
  |                                       
--+  +-------------- Ensidens **             
  |  |                                    
  +--+  +----------- Hyriopsis_desowitzi **  
     |  |                                 
     +--+  +-------- Rectidens **            
        |  |                              
        +--+     +-- Hyriopsis_panhai **     
           |  +--+                        
           |  |  +-- Hyriopsis_khoratensis **
           +--+                           
              |  +-- Hyriopsis_bialata **    
              +--+                        
                 +-- Hyriopsis_myersiana **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iTtTbiriSoTroTroSiD
11 11 9 3 10 10 1 21 
  +-------------------- Contradentina          
  |                                            
--+  +----------------- Ensidens **               
  |  |                                         
  |  |              +-- Hyriopsis_desowitzi **    
  +--+  +-----------+                          
     |  |           +-- Hyriopsis_delaportei **   
     |  |                                      
     +--+  +----------- Rectidens **              
        |  |                                   
        |  |     +----- Hyriopsis_myersiana **    
        +--+  +--+                             
           |  |  |  +-- Hyriopsis_bialata **      
           |  |  +--+                          
           +--+     +-- Hyriopsis_panhai **       
              |                                
              |  +----- Hyriopsis_khoratensis **  
              +--+                             
                 |  +-- Hyriopsis_phuphaniensis **
                 +--+                          
                    +-- Hyriopsis_sakhonensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 27

iTtTbiriSoTroTroSiD
10 10 6 2 1 1 1 19 
  +----- Ctenodesmina
--+                  
  |  +-- Rectidens **   
  +--+               
     +-- Hyriopsis **

Muanta et al., 2019 [Publ 3921], fig. 2

iTtTbiriSoTroTroSiD
9 9 8 2 0 0 0 19 
  +-- Hyriopsis **
--+            
  +-- Ensidens **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
8 8 0 2 75 10 1 17 
  +----- Contradentina        
  |                           
  |  +-- Ensidens_telus **       
  |  |                        
--+  +-- Ensidens_ingallsianus **
  |  |                        
  |  +-- Ensidens_jaculus **     
  |  |                        
  |  +-- Hyriopsis_desowitzi **  
  +--+                        
     +-- Hyriopsis_myersiana **  
     |                        
     +-- Hyriopsis_khoratensis **
     |                        
     +-- Ensidens_spiculus **    
     |                        
     +-- Ensidens_sagittarius **

Pfeiffer et al., 2021 [Publ 4126], fig. 30

iTtTbiriSoTroTroSiD
7 7 5 2 0 0 0 16 
  +-- Rectidens **
--+            
  +-- Ensidens **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
7 7 2 3 73 4 1 16 
  +-------- Contradentina        
  |                              
--+  +----- Hyriopsis_desowitzi **  
  |  |                           
  |  +----- Rectidens **            
  +--+                           
     +----- Ensidens **             
     |                           
     |  +-- Hyriopsis_khoratensis **
     +--+                        
        +-- Hyriopsis_myersiana **

Zieritz et al., 2016 [Publ 3804], fig. S2-1

iTtTbiriSoTroTroSiD
6 6 3 2 0 0 0 15 
  +-- Rectidens **
--+            
  +-- Hyriopsis **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 3 41 13 2 14 
  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 3 33 9 2 13 
  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
4 4 2 3 33 9 2 13 
  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
4 4 2 3 33 9 2 13 
  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 3 28 10 2 13 
  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Rectidens **    
        +--+                
           +-- Hyriopsis **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
4 4 2 2 87 4 1 13 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Ensidens **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
4 13 3 3 33 9 2 8 
  +-- Ensidens **           
  |                      
  +-- Hyriopsis_desowitzi **
  |                      
  +-- Rectidens **          
--+                      
  +-- Hyriopsis_bialata **  
  |                      
  +-- Ctenodesmina       
  |                      
  +-- Contradentina

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 3 33 9 2 13 
  +-------------- Ctenodesmina       
--+                                  
  |  +----------- Contradentina      
  +--+                               
     |  +-------- Ensidens **           
     +--+                            
        |  +----- Hyriopsis_bialata **  
        +--+                         
           |  +-- Hyriopsis_desowitzi **
           +--+                      
              +-- Rectidens **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
4 4 2 2 87 4 1 13 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Ensidens **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 57 6 2 13 
  +----------- Ctenodesmina 
--+                         
  |  +-------- Contradentina
  +--+                      
     |  +----- Ensidens **     
     +--+                   
        |  +-- Hyriopsis **    
        +--+                
           +-- Rectidens **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 32 3 1 13 
  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Hyriopsis **    
     +--+                
        +-- Rectidens **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 54 5 1 13 
  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Hyriopsis **    
     +--+                
        +-- Rectidens **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iTtTbiriSoTroTroSiD
3 3 0 3 4 4 1 13.5 
  +----- Contradentina
  |                   
--+  +-- Ensidens **     
  |  |                
  |  +-- Hyriopsis__1 ** 
  +--+                
     +-- Hyriopsis__2 ** 
     |                
     +-- Rectidens **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
3 3 1 3 73 3 1 13 
  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Rectidens **    
     +--+                
        +-- Hyriopsis **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 33 3 1 13 
  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Hyriopsis **    
     +--+                
        +-- Rectidens **

Zieritz et al., 2024 [Publ 4381], fig. 6A

iTtTbiriSoTroTroSiD
3 3 1 2 0 0 0 13 
  +-- Ensidens ** 
--+            
  +-- Rectidens **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
3 3 0 3 73 3 1 12 
  +----- Contradentina
  |                   
--+  +-- Ensidens **     
  |  |                
  +--+-- Hyriopsis **    
     |                
     +-- Rectidens **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
3 3 1 3 54 5 1 13 
  +-------- Contradentina
--+                      
  |  +----- Ensidens **     
  +--+                   
     |  +-- Hyriopsis **    
     +--+                
        +-- Rectidens **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 25 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 86 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 45 5 2 -- 
  +-------- Ctenodesmina 
--+                      
  |  +----- Contradentina
  +--+                   
     |  +-- Ensidens **     
     +--+                
        +-- Hyriopsis **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 43 3 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Ensidens **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
2 2 0 2 90 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
2 2 0 2 21 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 84 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 46 5 2 -- 
  +-------- Ctenodesmina 
--+                      
  |  +----- Contradentina
  +--+                   
     |  +-- Ensidens **     
     +--+                
        +-- Hyriopsis **

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 18 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
2 2 0 2 46 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
2 2 0 2 94 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
2 2 0 2 94 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 43 3 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Ensidens **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 51 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 47 5 2 -- 
  +-------- Ctenodesmina 
--+                      
  |  +----- Contradentina
  +--+                   
     |  +-- Ensidens **     
     +--+                
        +-- Hyriopsis **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
2 2 0 2 90 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
2 5 0 2 120 3 1 13 
  +-- Lens     
  |            
  +-- Ensidens ** 
  |            
--+-- Physunio 
  |            
  +-- Rectidens **
  |            
  +-- Yaukthwa

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 18 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 51 2 1 -- 
  +----- Contradentina
--+                   
  |  +-- Hyriopsis **    
  +--+                
     +-- Rectidens **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 3 1 -- 
  +-- Ensidens **     
--+                
  +-- Contradentina

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 8 6 1 -- 
  +-- Ensidens **     
--+                
  +-- Contradentina

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 3 1 -- 
  +-- Ensidens **     
--+                
  +-- Contradentina