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Page last updated
Wed 1 October 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Hyrioidea | Hyriidae | Hyriinae

tribe RHIPIDODONTINI Starobogatov, 1970: cladomics
     1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 43 (43 total)
number of subtaxa (rank: genus) = 1
analyzed in 33 cladograms in 21 publications
maximum number of terminals in an analysis = 2 (mean = 1.21, median = 1)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 7
maximum number of included terminals = 2, (mean = 2 median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 0.71

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
21901181140
                 +----- Diplodon_suavidicus__1 **
  +--------------+                            
  |              |  +-- Hyriini               
  |              +--+                         
  |                 +-- Castaliini            
  |                                           
  |           +-------- Westralunio           
  |           |                               
  +-----------+     +-- Hyridella_drapeta     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa    
  |           +--+                            
  |              |  +-- Diplodon_gratus **       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2 **
  |                                           
  |                 +-- Hyridella_australis   
  |  +--------------+                         
  |  |              +-- Cucumerunio           
  +--+                                        
     |  +-------------- Echyridella           
     +--+                                     
        |  +----------- Velesunio_ambiguus    
        +--+                                  
           |  +-------- Microdontia           
           +--+                               
              |  +----- Lortiella             
              +--+                            
                 |  +-- Alathyria             
                 +--+                         
                    +-- Velesunio_angasi

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
22011252--
  +-- Hyridellini
--+              
  +-- Diplodon **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
22014543--
  +-------- Echyridella
--+                    
  |  +----- Hyridellini
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Diplodon **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
22014443--
  +-------- Echyridella
--+                    
  |  +----- Hyridellini
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Diplodon **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
22014443--
  +-------- Echyridella
--+                    
  |  +----- Hyridellini
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Diplodon **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
22014543--
  +-------- Echyridella
--+                    
  |  +----- Hyridellini
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Diplodon **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
219011811411
                    +----- Diplodon_suavidicus__2 **
  +-----------------+                            
  |                 |  +-- Hyriini               
  |                 +--+                         
  |                    +-- Castaliini            
  |                                              
  |              +-------- Westralunio           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1 **
  +--+              +--+                         
     |                 +-- Diplodon_gratus **       
     |                                           
     |  +----------------- Echyridella           
     |  |                                        
     +--+              +-- Cucumerunio           
        |  +-----------+                         
        |  |           +-- Hyridella_australis   
        +--+                                     
           |  +----------- Velesunio_ambiguus    
           +--+                                  
              |  +-------- Alathyria             
              +--+                               
                 |  +----- Velesunio_angasi      
                 +--+                            
                    |  +-- Lortiella             
                    +--+                         
                       +-- Microdontia

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
11012922--
  +----- Diplodon **  
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014033--
  +----- Diplodon **  
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
11011622--
-- Diplodon **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
11013022--
  +-- Castaliini
--+             
  +-- Diplodon **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119864--
  +----------- Echyridella
--+                       
  |  +-------- Hyridellini
  +--+                    
     |  +----- Diplodon **   
     +--+                 
        |  +-- Castaliini 
        +--+              
           +-- Hyriini

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019011--
  +-- Diplodon **  
--+             
  +-- Castaliini

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
11013022--
  +----- Diplodon **  
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119864--
  +----- Diplodon **  
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
11013022--
  +-- Castaliini
--+             
  +-- Diplodon **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113822--
  +----- Diplodon **  
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
11011963--
  +-------- Hyridellini
--+                    
  |  +----- Diplodon **   
  +--+                 
     |  +-- Hyriini    
     +--+              
        +-- Castaliini

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113822--
  +----- Diplodon **  
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

iTtTbiriSoTroTroSiD
1101662--
  +----- Diplodon **  
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
11012922--
  +----- Diplodon **  
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
11013322--
  +-- Castaliini
--+             
  +-- Diplodon **

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
1101843--
  +----- Hyridella_depressa 
--+                         
  |  +-- Hyridella_australis
  +--+                      
     +-- Diplodon **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
11013022--
-- Diplodon_gratus **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019011--
  +-- Castaliini
--+             
  +-- Diplodon **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
11013543--
-- Diplodon_gratus **

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
1101743--
  +-- Hyridellini
--+              
  +-- Diplodon **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014033--
  +-------- Echyridella
--+                    
  |  +----- Diplodon **   
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Hyriini

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
11011043--
           +-- Diplodon **              
  +--------+                         
  |        +-- Castaliini            
--+                                  
  |  +-------- Hyridella_australis__2
  +--+                               
     |  +----- Hyridella_australis__1
     +--+                            
        |  +-- Echyridella           
        +--+                         
           +-- Hyridella_depressa

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
11013022--
-- Diplodon_gratus **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
11011722--
  +----- Hyridellini
--+                 
  |  +-- Diplodon **   
  +--+              
     +-- Castaliini

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
11013022--
  +-- Castaliini
--+             
  +-- Diplodon **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119664--
  +----------- Echyridella
--+                       
  |  +-------- Hyridellini
  +--+                    
     |  +----- Diplodon **   
     +--+                 
        |  +-- Castaliini 
        +--+              
           +-- Hyriini
 
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