Bivalvia | Unionida | Hyrioidea | Hyriidae | Hyriinae

tribe RHIPIDODONTINI Starobogatov, 1970: cladomics

1 genus

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 43 (43 total)
number of subtaxa (rank: genus) = 1
analyzed in 35 cladograms in 22 publications
maximum number of terminals in an analysis = 3 (mean = 1.26, median = 1)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 8
maximum number of included terminals = 3, (mean = 2.13 median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 0.63

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Rhipidodontini cladogram distances
PCoA Ordination

NJ Tree of Rhipidodontini cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
3 26 32 1 629 15 4 0 
                             +----- Diplodon_suavidicus__2 **
  +--------------------------+                            
  |                          |  +-- Hyriini               
  |                          +--+                         
  |                             +-- Castaliini            
  |                                                       
  |                             +-- Hyridella_depressa    
--+                       +-----+                         
  |                       |     +-- Hyridella_drapeta     
  |  +--------------------+                               
  |  |                    |  +----- Diplodon_suavidicus__1 **
  |  |                    +--+                            
  |  |                       |  +-- Diplodon_chilensis **    
  |  |                       +--+                         
  +--+                          +-- Diplodon_gratus **       
     |                                                    
     |                          +-- Cucumerunio           
     |  +-----------------------+                         
     |  |                       +-- Hyridella_australis   
     |  |                                                 
     +--+  +----------------------- Echyridella           
        |  |                                              
        |  |              +-------- Velesunio__1          
        +--+  +-----------+                               
           |  |           |  +----- Alathyria_jacksoni__1 
           |  |           +--+                            
           |  |              |  +-- Velesunio_ambiguus    
           +--+              +--+                         
              |                 +-- Alathyria_profuga     
              |                                           
              |  +----------------- Velesunio__3          
              |  |                                        
              +--+  +-------------- Microdontia           
                 |  |                                     
                 +--+           +-- Lortiella             
                    |  +--------+                         
                    |  |        +-- Velesunio_angasi      
                    +--+                                  
                       |  +-------- Alathyria_pertexta__2 
                       +--+                               
                          |  +----- Velesunio__2          
                          +--+                            
                             |  +-- Alathyria_jacksoni__2 
                             +--+                         
                                +-- Alathyria_pertexta__1

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 1 12 5 2 -- 
  +-- Hyridellini
--+              
  +-- Diplodon **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
2 19 0 1 18 11 4 21.5 
                    +----- Diplodon_suavidicus__2 **
  +-----------------+                            
  |                 |  +-- Hyriini               
  |                 +--+                         
  |                    +-- Castaliini            
  |                                              
  |              +-------- Westralunio           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1 **
  +--+              +--+                         
     |                 +-- Diplodon_gratus **       
     |                                           
     |  +----------------- Echyridella           
     |  |                                        
     +--+              +-- Cucumerunio           
        |  +-----------+                         
        |  |           +-- Hyridella_australis   
        +--+                                     
           |  +----------- Velesunio_ambiguus    
           +--+                                  
              |  +-------- Alathyria             
              +--+                               
                 |  +----- Velesunio_angasi      
                 +--+                            
                    |  +-- Lortiella             
                    +--+                         
                       +-- Microdontia

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 1 44 4 3 -- 
  +-------- Echyridella
--+                    
  |  +----- Hyridellini
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Diplodon **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 1 45 4 3 -- 
  +-------- Echyridella
--+                    
  |  +----- Hyridellini
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Diplodon **

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iTtTbiriSoTroTroSiD
2 19 0 1 18 11 4 20.5 
                 +----- Diplodon_suavidicus__1 **
  +--------------+                            
  |              |  +-- Hyriini               
  |              +--+                         
  |                 +-- Castaliini            
  |                                           
  |           +-------- Westralunio           
  |           |                               
  +-----------+     +-- Hyridella_drapeta     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa    
  |           +--+                            
  |              |  +-- Diplodon_gratus **       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2 **
  |                                           
  |                 +-- Hyridella_australis   
  |  +--------------+                         
  |  |              +-- Cucumerunio           
  +--+                                        
     |  +-------------- Echyridella           
     +--+                                     
        |  +----------- Velesunio_ambiguus    
        +--+                                  
           |  +-------- Microdontia           
           +--+                               
              |  +----- Lortiella             
              +--+                            
                 |  +-- Alathyria             
                 +--+                         
                    +-- Velesunio_angasi

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 1 45 4 3 -- 
  +-------- Echyridella
--+                    
  |  +----- Hyridellini
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Diplodon **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 1 44 4 3 -- 
  +-------- Echyridella
--+                    
  |  +----- Hyridellini
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Diplodon **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
-- Diplodon_gratus **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 3 3 -- 
  +-------- Echyridella
--+                    
  |  +----- Diplodon **   
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Hyriini

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
1 1 0 1 8 4 3 -- 
  +----- Hyridella_depressa 
--+                         
  |  +-- Hyridella_australis
  +--+                      
     +-- Diplodon **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 2 2 -- 
  +----- Hyridellini
--+                 
  |  +-- Diplodon **   
  +--+              
     +-- Castaliini

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 29 2 2 -- 
  +----- Diplodon **  
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 140 3 3 -- 
  +-------- Echyridella
--+                    
  |  +----- Diplodon **   
  +--+                 
     |  +-- Castaliini 
     +--+              
        +-- Hyriini

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 7 4 3 -- 
  +-- Hyridellini
--+              
  +-- Diplodon **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +-- Castaliini
--+             
  +-- Diplodon **

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 6 3 -- 
  +-------- Hyridellini
--+                    
  |  +----- Diplodon **   
  +--+                 
     |  +-- Hyriini    
     +--+              
        +-- Castaliini

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 35 4 3 -- 
-- Diplodon_gratus **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 137 2 2 -- 
  +----- Diplodon **  
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 137 2 2 -- 
  +----- Diplodon **  
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 199 6 4 -- 
  +----------- Echyridella
--+                       
  |  +-------- Hyridellini
  +--+                    
     |  +----- Diplodon **   
     +--+                 
        |  +-- Castaliini 
        +--+              
           +-- Hyriini

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
-- Diplodon_gratus **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 2 -- 
  +-- Castaliini
--+             
  +-- Diplodon **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +-- Castaliini
--+             
  +-- Diplodon **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 1 1 -- 
  +-- Diplodon **  
--+             
  +-- Castaliini

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +----- Diplodon **  
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +-- Castaliini
--+             
  +-- Diplodon **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
1 1 0 1 197 6 4 -- 
  +----------- Echyridella
--+                       
  |  +-------- Hyridellini
  +--+                    
     |  +----- Diplodon **   
     +--+                 
        |  +-- Castaliini 
        +--+              
           +-- Hyriini

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 6 6 2 -- 
  +----- Diplodon **  
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 3 3 -- 
  +----- Diplodon **  
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
1 1 0 1 199 6 4 -- 
  +----- Diplodon **  
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
1 1 0 1 10 4 3 -- 
           +-- Diplodon **              
  +--------+                         
  |        +-- Castaliini            
--+                                  
  |  +-------- Hyridella_australis__2
  +--+                               
     |  +----- Hyridella_australis__1
     +--+                            
        |  +-- Echyridella           
        +--+                         
           +-- Hyridella_depressa

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 29 2 2 -- 
  +----- Diplodon **  
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 1 1 -- 
  +-- Castaliini
--+             
  +-- Diplodon **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 2 2 -- 
-- Diplodon **