species Anodonta cygnea (Linnaeus, 1758): synonymy

Synonymy

Anodonta ovata Kuster of Authors

Kuster of Authors.

Mytilus cygneus Linnaeus, 1758

Lister, 1678. Hist. Anim. Anglic.: 29, pl. 1, fig. 3. [pre-linnaean]
Lister, 1685. Conch.: pl. 193, fig. 8. [pre-linnaean]
Klein, 1753. Tent. Method. Ostrac., etc.: pl. 9, fig. 26. [pre-linnaean]
Linnaeus, 1758. Syst. Nat. 1: 706, sp. 218.
Linnaeus, 1767. Syst. Nat. 1(2): 1158, sp. 257.
HOLOTYPE LS_257, Europe, a l’embouchure des fleuves.

Anodonta cygnea — Simpson (1900), Kobelt (1904), Haas (1940), Modell (1942), Heard (1975), ... Nagel (2025), Campbell & Lydeard (2026), Wu et al. (2026), Prie et al. (2026), Wu et al. (2026)
Anodonta (Cygnaeana) cygnaea [sic] — Germain (1904)
Anodonta (Groupe des Cygneana) [sic] cygnea — Caziot (1907)
Anodonta cygneus — Ortmann (1912), Heard & Guckert (1970), Kantor & Sysoev (2005)
“Anodonta cygnea” ** — Hannibal (1912)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta cygnaea [sic] — Germain (1931), Daget (1998), Dhora (2009)
Anodonta (s.s.) cygnea — Zhadin (1938), Haas (1969), Haas (1969), Kat (1983), Stadnichenko (1984), Falkner (1994), Vinarski & Kantor (2016)
Anodonta (s.s.) cygneus — Zhadin (1965), Habe (1977)
Anodonta (s.s.) cygnea cygnea — Falkner et al. (2001), Falkner et al. (2002)
Anodonta cygnea cygnea — Bogatov (2022)

Mytilus maxima Schröter, 1779

Schröter, 1779. Gesch. Flußconch.: 159, pl. 1, fig. 1.

Anodonta cygnaea [sic] forme maxima — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Mya arenaria Schröter, 1779

(misidentified reference to Mya arenaria Linnaeus, 1758)
Schröter, 1779. Gesch. Flußconch.: 165, pl. 2, fig. 1.

Anodonta (Cygnaeana) arenaria — Germain (1904)
Anodonta (Groupe des Cygneana) [sic] arenaria — Caziot (1907)
Anodonta cygnaea [sic] forme arenaria — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)
Anodonta cygnea — Araujo et al. (2009)

Mytilus zellensis Gmelin, 1791

(renamed Anodonta cellensis Pfeiffer, 1821)
Schröter, 1779. Gesch. Flußconch.: pl. 2, fig. 1.
Gmelin, 1791. Syst. Nat. 1(6): 3362, sp. 51.

Anodonta cygnea — Simpson (1900), Graf (2007)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cellensis [sic] — Zhadin (1938), Zhadin (1965)
Anodonta cellensis cellensis [sic] — Modell (1945)
Anodonta (s.s.) cygnea — Haas (1969)
Anodonta zellensis — Zatravkin (1983), Starobogatov et al. (2004), Mezhzherin et al. (2014)
Anodonta zellensis zellensis — Kantor & Sysoev (2005), Kantor et al. (2010)
Anodonta rellensis [sic] — Gozhik (2006)
Anodonta (s.s.) zellensis — Vinarski & Kantor (2016)
Anodonta cygnea zellensis — Bogatov (2022)

Mytilus stagnalis Gmelin, 1791

Schröter, 1779. Gesch. Flußconch.: pl. 1, fig. 1.
Gmelin, 1791. Syst. Nat. 1(6): 3352, sp. 50.

Anodonta cygnea — Simpson (1900), Graf (2007)
Anodonta (Gravidiana) stagnalis — Germain (1904)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta cellensis [sic] stagnalis — Modell (1945)
Anodonta stagnalis — Zatravkin (1983), Starobogatov et al. (2004), Kantor & Sysoev (2005), Gozhik (2006), Kantor et al. (2010), Mezhzherin et al. (2014)
Anodonta (s.s.) stagnalis — Stadnichenko (1984), Vinarski & Kantor (2016)
Anodonta cygnea stagnalis — Bogatov (2022)

Anodonta cygnaea ‘Linnaeus’ of Authors

(error for Mytilus cygneus Linnaeus, 1758)
‘Linnaeus’ of Authors.
Poiret, 1801. Coq. Fluv., et Terr. de l’Aisne: 109 [’Anodontites cygnæa’].
Gassies, 1849. Moll. Agenais: 189 [as ‘cygnæa’].
Dupuy, 1852. Hist. Natur. Moll. France (6): 601.
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 268.
Tiberi, 1879. Bull. Soc. Malac. Ital. 5: 143.
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 140.
Locard, 1882. Prodr. Malac. Franç. 1: 269.
Bourguignat, 1882. Natural. Sicil. 2 (1): 20 [1].
Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 89.
Locard, 1890. Contr. Faune Malacol. Franc. 14: 70 [26].
Baichère, 1890. Bull. Soc. Malac. France 7: 126.
Fagot, 1892. Bull. Soc. Ramond 27: 39.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 234.
Locard, 1899. Conchyliol. Portug.: 267.
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 289.
Germain, 1931. Faune de France 22: 752.
‘Linnaeus’ Daget, 1998. Cat. Bivalves Africains: 19.
Dhora, 2009. Arch. Biol. Sci. Belgrade 61(3): 547.

Anodonta cygnea — Simpson (1900)
Anodonta (Cygnaeana) cygnaea [sic] — Germain (1904)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta cygnaea [sic] — Germain (1931)

Anodonta variabilis Draparnaud, 1801

Draparnaud, 1801. Tab. Moll. Fr.: 108.

Anodonta (s.s.) cygnea — Vinarski & Kantor (2016)

Mytilus avonensis Montagu, 1803

(renamed Anodonta avonica Bourguignat, 1881)
Lister, 1685. Conch.: pl. 154, fig. 9?. [pre-linnaean]
Montagu, 1803. Test. Brit. 1-2: 172.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta avonensis — Germain (1931)
Anodonta anatina avonensis — Modell (1945)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Anodonta sulcata Lamarck, 1819

specimen image from publication Bruguière, 1797. Encycl. Méth. 2: pl. 202, figs. 1a-b [as Anodontites sp.].
Lamarck, 1819. Hist. Natur. Anim. sans Vert. 6: 85, sp. 3.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Anodonta intermedia Lamarck, 1819

Schröter, 1779. Gesch. Flußconch.: pl. 1, figs. 2-3.
Martini & Chemnitz, 1785. Conchyl. Cab. 8: 186, pl. 86, fig. 763 [as ‘Mytilus anatinus’].
specimen image from publication Bruguière, 1797. Encycl. Méth. 2: pl. 201, fig. 2 [as Anodontites sp.].
Lamarck, 1819. Hist. Natur. Anim. sans Vert. 6: 86, sp. 10.
POSSIBLE SYNTYPE MNHN_MPL_0004, dans le Loir, près Châteaudun [in the Loir, near Châteaudun].

Anodonta cygnea — Simpson (1900)
Anodonta (Intermediana) intermedia — Germain (1904)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta intermedia — Germain (1931)
Anodonta anatina intermedia — Modell (1945)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Mytilus incrassatus Sheppard, 1821

Sheppard, 1821. Trans. Linn. Soc. London 13: 85, pl. 5, fig. 4.

Anodonta cygnea — Simpson (1900)
Anodonta (Incrassatiana) incrassatus — Germain (1904)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta intermedia — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Mytilus macula Sheppard, 1821

(renamed Mytilus maculatus Brown, 1844)
Sheppard, 1821. Trans. Linn. Soc. London 13: 86, pl. 5, fig. 6.

Anodonta cygnea — Simpson (1900)
Anodonta (Tricassiniana) maculata — Germain (1904)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Anodonta cellensis Pfeiffer, 1821

(new name for Mytilus zellensis Gmelin, 1791)
Pfeiffer, 1821. Natur. Deutsch. Land Sussw. Moll. 1: 110, pl. 6, fig. 1.

Anodonta cygnea — Simpson (1900), Hannibal (1912)
Anodonta (Cygnaeana) arenaria — Germain (1904)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta cygnaea [sic] forme arenaria — Germain (1931)
Anodonta cellensis — Petr (1992)
Anodonta zellensis zellensis — Kantor & Sysoev (2005), Kantor et al. (2010)
Anodonta (s.s.) zellensis — Vinarski & Kantor (2016)

Anodon paludosus Turton, 1822

Turton, 1822. Conch. Ins. Brit.: 240, pl. 15, fig. 6.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Mytilus cellensis ‘Gmelin’ of Authors

(error for Mytilus zellensis Gmelin, 1791)
‘Gmelin’ of Authors.
Menke, 1828. Syn. Method. Moll.: 63.
Menke, 1830. Syn. Method. Moll.: 106.
Küster, 1839. Conchyl. Cab. 9(1): 16.
Mørch, 1853. Cat. Conch. Yoldi 2: 48.
Clessin, 1877. Deutsche Excurs. Moll. Fauna: 438.
Drouët, 1881. Unionidae Russie: 22.
Bourguignat, 1882. Natural. Sicil. 2 (1): 20 [1].
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 211.
Zhadin, 1938. Faune USSR Mollusques 4(1): 119.
Modell, 1945. Jena. Z. Med. Naturw. 78: 81.

Anodonta zellensis zellensis — Kantor & Sysoev (2005)
Anodonta (s.s.) zellensis — Vinarski & Kantor (2016)

Musculus latus Fleming, 1828

(intro'd as syn of Mytilus cygneus Linnaeus, 1758)
‘Lister’ Fleming, 1828. Hist. Brit. Anim.: 415.

Anodonta oblonga Millet, 1831

Millet, 1831. Mém. Soc. Agric. Sci. Angers 1: 242, pl. 12, fig. 1.

Anodonta cygnea — Simpson (1900), Araujo et al. (2009)
Anodonta (Jourdheuiliana) oblonga — Germain (1904)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)
Colletopterum oblongum — Kantor et al. (2010)

Anodonta confervigera Schlüter, 1838

Schlüter, 1838. Kurzgef. Syst. Verz. Conch. Samml.: ??.

Anodonta cellensis var. solearis Küster, 1839

(intro'd as intra-sp. of Anodonta cellensis Pfeiffer, 1821)
Küster, 1839. Conchyl. Cab. 9(1): 25.

Anodonta cellensis [sic] solearis — Modell (1945)
Anodonta (s.s.) cygnea — Haas (1969)
Anodonta (s.s.) cygnea solearis — Falkner et al. (2001)
Anodonta cygnea — Lopes-Lima et al. (2017)

Anodonta cellensis orata Held in Küster, 1839

(intro'd as intra-sp. of Anodonta cellensis Pfeiffer, 1821)
Held in Küster, 1839. Conchyl. Cab. 9(1): 25.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta callosa Held in Küster, 1839

Held in Küster, 1839. Conchyl. Cab. 9(1): 36, pl. 9, fig. 1.

Anodonta cygnea — Simpson (1900)
Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Anodonta cariosa Küster, 1842

Küster, 1842. Conch. Cab. Anodonta 9: 43, pl. 4, fig. 3; pl. 5, fig. 1; pl. 10, figs. 1-2.

Anodonta cygnea — Simpson (1900)
Anodonta (Cygnaeana) cariosa — Germain (1904)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cellensis [sic] var. cariosa — Zhadin (1938)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Anodonta arealis Küster, 1842

Küster, 1842. Conch. Cab. Anodonta 9: 47, pl. 9, figs. 2-4.

Anodonta cygnea — Simpson (1900)
Anodonta (Arealiana) arealis — Germain (1904)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta tumida Küster, 1842

Küster, 1842. Conch. Cab. Anodonta 9: 48, pl. 12, figs. 1-2.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Anodonta subluxata Küster, 1842

Küster, 1842. Conch. Cab. Anodonta 9: 52, pl. 13, figs. 1-3.

Anodonta cygnea — Simpson (1900)
Anodonta (Groupe des Westerlundiana) subluxata — Caziot (1907)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta avonensis var. subluxata — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)
Anodonta anatina — Nagel (2025)

Anodonta sondermannii Küster, 1842

Küster, 1842. Conch. Cab. Anodonta 9: 54, pl. 13, fig. 4.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Anodonta opalina Küster, 1842

Küster, 1842. Conch. Cab. Anodonta 9: 60, pl. 16, fig. 1-2.

Anodonta cygnea — Simpson (1900)
Anodonta grandis var. gigantea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Mytilus maculatus Brown, 1844

(new name for Mytilus macula Sheppard, 1821)
Brown, 1844. Ill. Rec. Conch. Gr. Brit. & Ireland: 80.
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 285 [as ‘Anodonta maculata’].

Anodonta (Tricassiniana) maculata — Germain (1904)
Anodonta maculata — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta elongata Potiez & Michaud, 1844

(homonym of Anodonta elongata Holandre, 1836)
Potiez & Michaud, 1844. Galérie Moll. at. Mus. Douai 2: 141, pl. 55, fig. 1.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)

Anodon cygneus var. contorta Brown, 1844

(intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758)
Brown, 1844. Ill. Rec. Conch. Gr. Brit. & Ireland: 81, pl. 30, figs. 7-8.
Brown, 1845. Ill. Land & Fr. Water Conch. Gr. Brit. & Ireland: 106, pl. 18, figs. 1-2.

Anodonta (s.s.) cygnea — Vinarski & Kantor (2016)

Anodonta ranarum Morelet, 1845

Morelet, 1845. Moll. Terr. Fluv. Portugal: 104, pl. 12, fig. 2.
HOLOTYPE BMNH_93-2-4-1737, afflu. de la Guardiana, Portugal.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Anodonta glabra Stabile, 1845

(Ziegler) Stabile, 1845. Faun. Lug.: 58, pl. 2, fig. 68.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Vinarski & Kantor (2016)

Anodonta lucasii Deshayes, 1847

Deshayes, 1847. Exploration scientifique de l’Algérie: pl. 108, figs. 1-3.
HOLOTYPE BMNH_93-2-4-1950, Algiers.

Anodonta lucasi — Simpson (1900), IUCN (2010), Van Damme et al. (2010)
Anodonta (Group of A. cygnea) lucasi — Simpson (1914)
Anodonta cygnea — Van Damme (1984), Khalloufi & Boumaïza (2005), Graf & Cummings (2011)
Anodonta cygnaea [sic] — Daget (1998)

Anodonta arelatensis Dupuy, 1850

Jacquemin, 1835. Guide Ror. Arles: 124. [nomen nudum]
Dupuy, 1850. Hist. Natur. Moll. France (4): pl. 19, fig. 14.
‘Jacquemin’ Dupuy, 1852. Hist. Natur. Moll. France (6): 611.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta idrina Spinelli, 1851

Spinelli, 1851. Cat. Moll. Terr. Fluv. Prov. Brescia: 19, figs. 5-6.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969)

Anodon lucasii Morelet, 1851

(reference to Anodonta lucasii Deshayes, 1847)
Morelet, 1851. J. de Conch. 2: 359.

Anodonta lucasi — Simpson (1900)

Anodonta grossa Gallenstein, 1852

‘(Ziegler)’ Gallenstein, 1852. Jb. Naturh. Landesmus. Kärnten 1: 122 (sep. 66).

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta assimilis Gallenstein, 1852

‘(Ziegler)’ Gallenstein, 1852. Jb. Naturh. Landesmus. Kärnten 1: 122 (sep. 66).

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta fuscata Gallenstein, 1852

(homonym of Anodonta inflata var. fuscata Rafinesque, 1831)
‘(Ziegler)’ Gallenstein, 1852. Jb. Naturh. Landesmus. Kärnten 1: 123 (sep. 67).

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta vetula Gallenstein, 1852

‘(Ziegler)’ Gallenstein, 1852. Jb. Naturh. Landesmus. Kärnten 1: 124 (sep. 68).

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta deplanata Gallenstein, 1852

‘(Ziegler)’ Gallenstein, 1852. Jb. Naturh. Landesmus. Kärnten 1: 124 (sep. 68).

Anodonta cellensis [sic] deplanata — Modell (1945)
Anodonta (s.s.) cygnea — Haas (1969)
Anodonta (s.s.) cygnea deplanata — Falkner et al. (2001)
Anodonta cygnea — Lopes-Lima et al. (2017)

Unio (Anodonta) piscinalis var. latissima Gallenstein, 1852

(intro'd as intra-sp. of Anodonta piscinalis Nilsson, 1822)
‘Kokeil’ Anton, 1839. Verz. Conchylien: 16, no. 584 var. [nomen nudum]
‘(Kokeil)’ Gallenstein, 1852. Jb. Naturh. Landesmus. Kärnten 1: 125 (sep. 69).

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta platyrhincha Gallenstein, 1852

‘(Kokeil)’ Gallenstein, 1852. Jb. Naturh. Landesmus. Kärnten 1: 125 (sep. 69).

Anodonta anatina platyrhyncha [sic] — Modell (1945)
Anodonta (s.s.) cygnea — Haas (1969)

Anodontes europaea Leach, 1852

Lister, 1678. Hist. Anim. Anglic.: 29, pl. 1, fig. 3. [pre-linnaean]
Leach, 1852. Syn. Moll. Gr. Brit.: 326 [as ‘europæa’].

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Vinarski & Kantor (2016)

Anodonta cellensis inflata Rossmässler, 1853

(homonym of Anodonta inflata Rafinesque, 1831; intro'd as intra-sp. of Anodonta cellensis Pfeiffer, 1821)
Rossmässler, 1853. Zeitschr. f. Malakozool. 10: 15.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cygnea cordata Rossmässler, 1859

(intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758)
Rossmässler, 1859. Icon. Land- und Süssw.-Moll. 3 (5-6): 136, nos. 970-971, pls. 88-89, figs. 968a-b.
LECTOTYPE SMF_5140 (by Zilch, 1967: 110), Lache b. Platschütz b. Allenburg [Germany].

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)
Anodonta cygnea — Nagel (2025)

Anodonta leprosa benacensis Gredler, 1859

(intro'd as intra-sp. of Anodonta leprosa Drouët, 1883)
Spinelli, 1851. Cat. Moll. Terr. Fluv. Prov. Brescia: 19. [nomen nudum]
‘Villa’ Gredler, 1859. Verh. Zool.-Bot. Ges. Wien 9: 260.
Villa & Villa, 1871. Bull. Malac. Italiano: 94 [as ‘Anodonta benacensis’]. [nomen nudum]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta atrovirens Stabile, 1859

(homonym of Anodonta atrovirens Philippi, 1849)
‘Shuttleworth’ Stabile, 1859. Atti soc. geol. Milano 1: 169, 182, sp. 43.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta atrovirens mut. normalis Stabile, 1859

(intro'd as intra-sp. of Anodonta atrovirens Stabile, 1859)
Stabile, 1859. Atti soc. geol. Milano 1: 169, 183, sp. 44.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta atrovirens mut. attenuata Stabile, 1859

(homonym of Anodonta attenuata Held, 1836; intro'd as intra-sp. of Anodonta atrovirens Stabile, 1859)
Stabile, 1859. Atti soc. geol. Milano 1: 169, 183, sp. 45.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta piscinalis var. rostrata anom. contracta Stabile, 1859

(intro'd as intra-sp. of Anodonta rostrata Rossmässler, 1836)
Stabile, 1859. Atti soc. geol. Milano 1: 169, 183, sp. 47. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cygnaea var. minor de Joannis, 1859

(intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 269.

Anodonta cygnaea var. radiata de Joannis, 1859

(intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758; homonym of Anodonta (Lastena) ohiensis var. radiata Rafinesque, 1820)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 268, pl. 1.

Anodonta (Gravidiana) stagnalis — Germain (1904)

Anodonta cygnaea var. radiata de Joannis, 1859

(intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758; homonym of Anodonta (Lastena) ohiensis var. radiata Rafinesque, 1820)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 268, pl. 1.

Anodonta (Gravidiana) stagnalis radiata — Germain (1904)

Anodonta psammita Bourguignat, 1862

Bourguignat, 1862. Malacol. Lac Quatre-Cantons: 58, pl. 4, figs. 1-3.
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 297.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta helvetica Bourguignat, 1862

Bourguignat, 1862. Malacol. Lac Quatre-Cantons: 59, pl. 3, figs. 1-2.
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 234.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cygnea lingua Mørch, 1863

(intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758)
‘Yoldi’ Mørch, 1853. Cat. Conch. Yoldi 2: 48 [as ‘A. lingua’]. [nomen nudum]
‘(Yoldi)’ Mørch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): 83.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cygnea lirata Mørch, 1863

(intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758)
Mørch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): 83.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cygnea lyngbyana Mørch, 1863

(intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758)
Mørch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): 84.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cygnea forchhammeri Mørch, 1863

(intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758)
Mørch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): 84.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta embia Bourguignat, 1864

Bourguignat, 1864. Mal. Alg. 2: 297, pl. 25, figs. 1-4; pl. 26, fig. 1.

Anodonta lucasi — Simpson (1900), IUCN (2010)
Anodonta (Group of A. cygnea) lucasi — Simpson (1914)
Anodonta cygnea — Van Damme (1984), Khalloufi & Boumaïza (2005)
Anodonta cygnaea [sic] — Daget (1998)

Anodonta letourneuxi Bourguignat, 1864

Bourguignat, 1864. Mal. Alg. 2: 299, pl. 26, figs. 2-6.

Anodonta numidica — Simpson (1900)
Anodonta (Group of A. cygnea) numidica — Simpson (1914)
Anodonta cygnaea [sic] — Daget (1998)

Anodonta rostrata var. major Brusina, 1866

(intro'd as intra-sp. of Anodonta rostrata Rossmässler, 1836)
Brusina, 1866. Moll. Dalm.: 131.

Anodonta cygnea var. rostrata Brot, 1867

(homonym of Anodonta rostrata Rossmässler, 1836; intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): 30, pl. 2, figs. 1-2.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. dilatata Brot, 1867

(intro'd as intra-sp. of Anodonta cellensis Pfeiffer, 1821)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): 37, pl. 6, fig. 4.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis rostrata Brot, 1867

(homonym of Anodonta rostrata Rossmässler, 1836; intro'd as intra-sp. of Anodonta cellensis Pfeiffer, 1821)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): 37, pl. 4, fig. 1.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta anatina rostrata Brot, 1867

(homonym of Anodonta rostrata Rossmässler, 1836; intro'd as intra-sp. of Mytilus anatinus Linnaeus, 1758)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): 40, pl. 7, fig. 3.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta anatina major Brot, 1867

(intro'd as intra-sp. of Mytilus anatinus Linnaeus, 1758; homonym of Anodonta rostrata var. major Brusina, 1866)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): 42, pl. 7, fig. 1; pl. 6, fig. 3.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta anatina elongata Brot, 1867

(homonym of Anodonta elongata Holandre, 1836; intro'd as intra-sp. of Mytilus anatinus Linnaeus, 1758)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): 43, pl. 6, fig. 1.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta pictetiana rostrata Brot, 1867

(homonym of Anodonta rostrata Rossmässler, 1836; intro'd as intra-sp. of Anodonta pictetiana Brot, 1867)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): 46, pl. 8, fig. 2.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta pictetiana elongata Brot, 1867

(homonym of Anodonta elongata Holandre, 1836; intro'd as intra-sp. of Anodonta pictetiana Brot, 1867)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): 46, pl. 8, fig. 3.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta piscinalis major Brot, 1867

(homonym of Anodonta rostrata var. major Brusina, 1866; intro'd as intra-sp. of Anodonta piscinalis Nilsson, 1822)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): 47, pl. 5, fig. 1.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cygnea var. rossmasleri Malzine, 1867

(intro'd as intra-sp. of Mytilus cygneus Linnaeus, 1758)
‘L.’ Rossmässler, 1835. Icon. Land- und Süssw.-Moll. 1 (1): pl. 3, fig. 67 [as ‘Anodonta cygnea’].
Malzine, 1867. Essai faun. malac. Belg.: 34.

Anodonta (s.s.) cygnea — Haas (1969)

Anodon simplicidus G.B. Sowerby II, 1867

specimen image from publication ‘Parreyss’ G.B. Sowerby II, 1867. Conch. Icon. 17: pl. 12, sp. 40.

Anodonta cygnea — Simpson (1900)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)

Anodonta cellensis var. minor Brot, 1867

(intro'd as intra-sp. of Anodonta cellensis Pfeiffer, 1821; homonym of Anodonta cygnaea var. minor de Joannis, 1859)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): 35.

Anodonta cygnaea [sic] — Germain (1931)

Anodonta capitata Clessin, 1874

Clessin, 1874. Conchyl. Cab.: 125, pl. 39, figs. 1-2.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta mutabilis diminuata Clessin, 1876

(intro'd as intra-sp. of Anodonta mutabilis Clessin, 1877; renamed Anodonta diminuta Bourguignat, 1881)
Clessin, 1876. Conchyl. Cab.: 236, pl. 87, fig. 1.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta mutabilis fragilissima Clessin, 1876

(intro'd as intra-sp. of Anodonta mutabilis Clessin, 1877; renamed Anodonta fragillima Bourguignat, 1881)
Clessin, 1876. Conchyl. Cab.: 237, pl. 87, fig. 2.

Anodonta (Ventricosiana) fragilissima — Germain (1904)
Anodonta cygnaea [sic] forme fragilissima — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta mutabilis Clessin, 1877

Clessin, 1877. Deutsche Excurs. Moll. Fauna: 434.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969), Vinarski & Kantor (2016)

Anodonta mutabilis var. lacustrina Clessin, 1877

(intro'd as intra-sp. of Anodonta mutabilis Clessin, 1877)
Clessin, 1877. Deutsche Excurs. Moll. Fauna: 444, fig. 291.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta gravida Drouët, 1879

Drouët, 1879. J. de Conch. 27: 142.

Anodonta palustris gravida — Modell (1945)
Anodonta (s.s.) cygnea — Haas (1969)
Anodonta (s.s.) cygnea gravida — Falkner (1994), Bank (2006)
Anodonta cygnea — Lopes-Lima et al. (2021)

Anodonta kleciaki Drouët, 1881

Drouët, 1881. J. de Conch. 29: 28.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta savensis Drouët, 1881

(renamed Anodonta savica Bourguignat, 1881)
Drouët, 1881. J. de Conch. 29: 28.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta moesica Drouët, 1881

Drouët, 1881. J. de Conch. 29: 29.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta pammegala Bourguignat, 1881

Schröter, 1779. Gesch. Flußconch.: pl. 1, fig. 1.
‘L.’ Rossmässler, 1837. Icon. Land- und Süssw.-Moll. 1 (5-6): 23, pl. 25, fig. 342 [as ‘Anodonta cygnea’].
‘Linné’ Küster, 1853. Conch. Cab. Anodonta: 58, pl. 15 [as ‘Anodonta cygnea’].
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 107.

Anodonta cygnaea [sic] forme maxima — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta eucypha Bourguignat, 1881

‘L.’ Rossmässler, 1835. Icon. Land- und Süssw.-Moll. 1 (1): pl. 3, fig. 67 [as ‘A. cygnea’].
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 108.

Anodonta (Pammegaliana) eucypha — Germain (1904)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta cellensis [sic] eucypha — Modell (1945)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta eumorphia ‘Letourneux, in litt.’ Bourguignat, 1881

‘Letourneux, in litt.’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 109.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta thiessae Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 112 [as ‘thiessæ’].

Anodonta (s.s.) cygnea — Haas (1969)
Anodonta (s.s.) cygnea gravida — Bank (2006)

Anodonta callidaea ‘Letourneux, in litt.’ Bourguignat, 1881

‘Letourneux, in litt.’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 115 [as ‘callidæa’].

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta gallica Bourguignat, 1881

Sheppard, 1821. Trans. Linn. Soc. London 13: pl. 4, fig. 1 [as ‘Mytilus anatinus’].
Chenu, 1845. Biblio. Conch. 1: 270, pl. 26, fig. 1.
Brown, 1845. Ill. Land & Fr. Water Conch. Gr. Brit. & Ireland: pl. 12, fig. 1 [as ‘A. cellensis’]. [in part]
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 123.

Anodonta gallica — Rochebrune (1903)
Anodonta (Ventricosiana) gallica — Germain (1904)
Anodonta cygnaea [sic] forme ventricosa — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)
Anodonta cygnea — Araujo et al. (2009)

Anodonta locardi Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 126.
Bourguignat in Locard, 1884. Contr. Faune Malacol. Franc. 8: 12.
SYNTYPE MNHN_22312, fossés des forts de la Vitriolerie.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta charpyi Bourguignat, 1881

‘Dupuy’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 127.
SYNTYPE MNHN_22175, Le Drugeon, Doubs.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta fragillima Bourguignat, 1881

(new name for Anodonta mutabilis fragilissima Clessin, 1876)
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 129.
Bourguignat, 1882. Natural. Sicil. 2 (1): 20 [1].
Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 88.
Bourguignat in Locard, 1884. Contr. Faune Malacol. Franc. 8: 13.

Anodonta (Ventricosiana) fragilissima — Germain (1904)
Anodonta cygnaea [sic] forme fragilissima — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta acyrta Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 130.

Anodonta (Ventricosiana) fragilissima acyrta — Germain (1904)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta stataria ‘Ray, in litt.’ Bourguignat, 1881

‘Ray, in litt.’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 132.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta livronica ‘Fagot, in litt.’ Bourguignat, 1881

‘Fagot, in litt.’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 133.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta saintsimoniana Fagot in Bourguignat, 1881

Fagot in Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 142 [as ‘saint-simoniana’].
Fagot in Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 89.
Fagot in Locard, 1884. Contr. Faune Malacol. Franc. 8: 20.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta fagoti Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 144.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta condatina ‘Letourneux, in litt.” Bourguignat, 1881

‘Letourneux, in litt.” Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 147.

Anodonta (Cygnaeana) condatina — Germain (1904)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta nansoutyana Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 148.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta rhodani Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 152.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta ellipsopsis Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 156.
Bourguignat in Locard, 1884. Contr. Faune Malacol. Franc. 8: 21.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta ressmanni Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 159.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta lacuum Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 171.

Anodonta lacuum — Rochebrune (1903)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta adamii Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 191.
Bourguignat, 1882. Natural. Sicil. 2 (1): 22 [3].
Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 95.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta broti Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 210.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta humberti Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 213.

Anodonta (s.s.) cygnea — Haas (1969)
Anodonta anatina — Nagel (2025)

Anodonta sturmi Bourguignat, 1881

(new name for Mytilus anatinus Sturm, 1803)
‘Gmel. Linn.’ Sturm, 1803. Deutsch. Fauna 6(1): unnumberd [11], text figs. a-c [as ‘Mytilus anatinus’].
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 223.
SYNTYPE MNHN_22555, Heuilly-sur-Saône (Côte-d’Or).

Anodonta cygnaea [sic] forme sturmi — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta lutetiana Bourguignat, 1881

‘Mabille, mss.’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 223.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta serbica ‘Letourneux, in litt.’ Bourguignat, 1881

‘Letourneux, in litt.’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 241.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta pilariana Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 249.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta danica ‘Mörch, in litt.’ Bourguignat, 1881

‘Mörch, in litt.’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 252.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta ruvida Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 259.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta kusteri Bourguignat, 1881

‘Pfeiffer’ Küster, 1853. Conch. Cab. 9: 56, pl. 11*, fig. 3 [as ‘A. ventricosa’]. [in part]
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 284.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta clessini Bourguignat, 1881

Clessin, 1876. Conchyl. Cab.: 83, pl. 14, figs. 3-4 [as ‘A. anatina’]. [in part]
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 287.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta carotae Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 292 [as ‘carotæ’].
Bourguignat, 1882. Natural. Sicil. 2 (1): 23 [4].
Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 110 [as ‘carotæ’].

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta truncata Bourguignat, 1881

‘Parreyss, mss.’ Brusina, 1866. Moll. Dalm.: 131. [nomen nudum]
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 300.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta avonica Bourguignat, 1881

(new name for Mytilus avonensis Montagu, 1803)
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 304.

Anodonta (Avoniana) avonica — Germain (1904)
Anodonta avonensis — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta desori ‘Coutagne, in litt.’ Bourguignat, 1881

‘Coutagne, in litt.’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 305.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta pfeifferi Bourguignat, 1881

C. Pfeiffer, 1825. Naturgesch. Dtsch. Land-Süßw. Moll. 2: pl. 3, figs. 1-3 [as ‘A. ventricosa’]. [in part]
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 320.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta monterosati Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 330.
Bourguignat, 1882. Natural. Sicil. 2 (1): 23 [4].
Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 97.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta savica Bourguignat, 1881

(new name for Anodonta savensis Drouët, 1881)
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 340.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta ilysoeca Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 341 [as ‘ilysœca’].

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta resima Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 345.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta aristidis Bourguignat, 1881

Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 347.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta brusinae ‘Letourneux, in litt.’ Bourguignat, 1881

‘Letourneux, in litt.’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 353 [as ‘brusinæ’].

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta balatonica Servain, 1881

Servain, 1881. Malac. Lac Balaton: 106.

Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta tihanyca Servain, 1881

Servain, 1881. Malac. Lac Balaton: 108.

Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta tissoti Servain, 1881

Servain, 1881. Malac. Lac Balaton: 111.

Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta hydatina Servain, 1881

Servain, 1881. Malac. Lac Balaton: 113.
Servain, 1882. Hist. Moll. Acéph. Francfort: 45.

Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta aquatica Servain, 1881

Servain, 1881. Malac. Lac Balaton: 115.

Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta briandiana Servain, 1881

Servain, 1881. Malac. Lac Balaton: 117.

Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta renoufi Servain, 1881

Servain, 1881. Malac. Lac Balaton: 119.

Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta hazayana Servain, 1881

Servain, 1881. Malac. Lac Balaton: 121.

Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta dubreili Servain, 1881

Servain, 1881. Malac. Lac Balaton: 123.

Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta parmata Drouët, 1881

Drouët, 1881. Unionidae Russie: 26.

Anodonta (s.s.) cygnea — Haas (1969)
Anodonta parmata — Kantor et al. (2010)

Anodonta sieversi Drouët, 1881

Drouët, 1881. Unionidae Russie: 28.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta wimmeri Drouët, 1881

Drouët, 1881. J. de Conch. 29: 251.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta dokici Drouët, 1881

Drouët, 1881. J. de Conch. 29: 251.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta nymphigena Drouët, 1881

Drouët, 1881. J. de Conch. 29: 252.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta dealbata Drouët, 1881

Drouët, 1881. J. de Conch. 29: 254.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. acuta Drouët, 1881

(intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791)
Drouët, 1881. Unionidae Russie: 22.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta diminuta Bourguignat, 1881

(new name for Anodonta mutabilis diminuata Clessin, 1876)
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 229.

Anodonta klecaki ‘Drouët’ of Authors

(error for Anodonta kleciaki Drouët, 1881)
‘Drouët’ of Authors.
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 275.
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 261.

Anodonta ellipsopsis var. minor Bourguignat, 1881

(intro'd as intra-sp. of Anodonta ellipsopsis Bourguignat, 1881; homonym of Anodonta cygnaea var. minor de Joannis, 1859)
Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 159.

Anodonta macrostena Servain, 1882

Servain, 1882. Hist. Moll. Acéph. Francfort: 32.

Anodonta (Macrosteniana) macrostena — Germain (1904)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta maganica Servain, 1882

Servain, 1882. Hist. Moll. Acéph. Francfort: 36.

Anodonta (s.s.) cellensis [sic] var. maganica — Zhadin (1938)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta telmoeca Servain, 1882

(correct name for Anodonta telmaeca Servain, 1882)
Servain, 1882. Hist. Moll. Acéph. Francfort: 31 [as ‘telmæca’], 38 [as ‘telmœca’].

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta bythioeca Servain, 1882

Servain, 1882. Hist. Moll. Acéph. Francfort: 40 [as ‘bythiœca’].

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta manica Servain, 1882

Servain, 1882. Hist. Moll. Acéph. Francfort: 42.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta morini Servain, 1882

Servain, 1882. Hist. Moll. Acéph. Francfort: 45.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta ocnera Servain, 1882

Servain, 1882. Hist. Moll. Acéph. Francfort: 47.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta complacita Servain, 1882

Servain, 1882. Hist. Moll. Acéph. Francfort: 49.
Servain, 1888. Bull. Soc. Malac. France 5: 332.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta codopsis Servain, 1882

Servain, 1882. Hist. Moll. Acéph. Francfort: 53.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cypholena Servain, 1882

Servain, 1882. Hist. Moll. Acéph. Francfort: 58.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta delpretei Bourguignat, 1882

Bourguignat, 1882. Natural. Sicil. 2 (1): 21 [2] [as ‘del pretei’].
Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 91 [as ‘del pretei’].

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta raimondoi Bourguignat, 1882

Bourguignat, 1882. Natural. Sicil. 2 (1): 23 [4].
Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 110.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta forschammeri “Mörch” of Authors

(error for Anodonta cygnea forchhammeri Mørch, 1863)
“Mörch” of Authors.
Locard, 1882. Prodr. Malac. Franç. 1: 268.
Locard, 1890. Contr. Faune Malacol. Franc. 14: 65 [21].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 230.
Germain, 1931. Faune de France 22: 753.

Anodonta cygnaea [sic] — Germain (1931)

Anodonta blauneri Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 91.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta alseria Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 93.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta utriculosa Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 94.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta anxurensis Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 95.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta stabilei Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 96.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta padana Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 97.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta pinii Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 98.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta longirostris Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 101.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta scapulosa Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 104.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta romana Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 106.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta leprosa Drouët, 1883

‘Parreyss’ Zelebor, 1851. Ber. Mit. Freunde Naturw. Wien 7: 232, sp. 9.
‘Parreys’ de Betta, 1870. Moll. prov. Veron: 137. [nomen nudum]
‘Parreys’ Bourguignat, 1881. Mater. Moll. Acécphales Syst. Europ. 1: 193. [nomen nudum]
‘Parryss’ Bourguignat, 1882. Natural. Sicil. 2 (1): 23 [4]. [nomen nudum]
Drouët, 1883. Unionidae de l’Italie: 107.
Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 109.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta utinensis Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 109.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta sebinensis Drouët, 1883

(homonym of Anodonta sebinensis Adami in Bourguignat, 1881)
‘(Adami)’ Drouët, 1883. Unionidae de l’Italie: 117.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta villae Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 120.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta paupercula Drouët, 1883

Drouët, 1883. Unionidae de l’Italie: 121.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cristata Drouët, 1883

(homonym of Anodonta complanata cristata Moquin-Tandon, 1855)
Drouët, 1883. Unionidae de l’Italie: 124.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta doriana Issel in Bourguignat, 1883

Issel in Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 85.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta oblonga mantuacina Bourguignat, 1883

(intro'd as intra-sp. of Anodonta oblonga Millet, 1831)
Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 90.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta isseli Bourguignat, 1883

Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 91.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta antinoriana Bourguignat, 1883

Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 94.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta eporediana Issel in Bourguignat, 1883

Issel in Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 97.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta gestroi Bourguignat, 1883

Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 99.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta arturi Bourguignat, 1883

Bourguignat, 1883. Aperçu Unionid. Pénins. Ital.: 111.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta henriquesi da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 123.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta paulinoi da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 125.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta subregularis da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 127.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta silvai Bourguignat in da Silva e Castro, 1883

Bourguignat in da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 128.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta giraldesi da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 130.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta alleniana da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 131.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta embiella Hagenmüller in da Silva e Castro, 1883

Hagenmüller in da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 133.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta bofilliana Bourguignat in da Silva e Castro, 1883

Bourguignat in da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 134.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta machadoiana da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 135.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta capelloiana da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 136.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta bocageana da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 138.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta tamegana da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 141.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta mengoiana da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 143.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta barbosana da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 144.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta specialis da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 145.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta rosai da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 148.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta ribeiroiana da Silva e Castro, 1883

da Silva e Castro, 1883. J. Sci. Math. Phys. Nat. 35: 149.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta reneana Péchaud, 1884

Péchaud, 1884. Bull. Soc. Malac. France 1: 189.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta nevirnensis Locard, 1884

‘Péchaud’ Locard, 1884. Contr. Faune Malacol. Franc. 8: 7.
‘Pechaud’ Locard, 1890. Contr. Faune Malacol. Franc. 14: 63 [19].
‘Pechaud’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 228.
LECTOTYPE MNHN_22403 (by Graf, 2011: 492), Saint-Laurent d’Ain.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta hecartiana Locard, 1884

Locard, 1884. Contr. Faune Malacol. Franc. 8: 9.
Locard, 1890. Contr. Faune Malacol. Franc. 14: 63 [19].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 228.
LECTOTYPE MNHN_22258 (by Graf, 2011: 478), La Saône à Lyon.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta perroudi Locard, 1884

Locard, 1884. Contr. Faune Malacol. Franc. 8: 17.
Locard, 1890. Contr. Faune Malacol. Franc. 14: 73 [29].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 238, fig. 244.
LECTOTYPE MNHN_22451 (by Graf, 2011: 497), Rhône au confluent de la Mulatière.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta euthymeana Locard, 1884

Locard, 1884. Contr. Faune Malacol. Franc. 8: 27.
Locard, 1890. Contr. Faune Malacol. Franc. 14: 74 [30].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 237.
LECTOTYPE MNHN_22201 (by Graf, 2011: 472), Le Menthon (Ain).

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta palustris Pini, 1884

(homonym of Anodonta palustris Férussac, 1822)
Pini, 1884. Atti Soc. Ital. Sci. Nat. 27: 86, text fig.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta henriquezi ‘Castro’ of Authors

(error for Anodonta henriquesi da Silva e Castro, 1883)
‘Castro’ of Authors.
Locard, 1884. Contr. Faune Malacol. Franc. 8: 11.
Locard, 1890. Contr. Faune Malacol. Franc. 14: 68 [24].
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 206.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 232.
Locard, 1899. Conchyliol. Portug.: 263.
Germain, 1931. Faune de France 22: 753.

Anodonta locardi var. major Locard, 1884

(intro'd as intra-sp. of Anodonta locardi Bourguignat, 1881; homonym of Anodonta rostrata var. major Brusina, 1866)
Locard, 1884. Contr. Faune Malacol. Franc. 8: 12. [infra-subsp.]

Anodonta locardi var. curta Locard, 1884

(intro'd as intra-sp. of Anodonta locardi Bourguignat, 1881; homonym of Anodonta macrostena var. curta Servain, 1882)
Locard, 1884. Contr. Faune Malacol. Franc. 8: 13. [infra-subsp.]

Anodonta fragillima var. elongata Locard, 1884

(intro'd as intra-sp. of Anodonta fragillima Bourguignat, 1881; homonym of Anodonta elongata Holandre, 1836)
Locard, 1884. Contr. Faune Malacol. Franc. 8: 14. [infra-subsp.]

Anodonta macrostena var. major Locard, 1884

(homonym of Anodonta rostrata var. major Brusina, 1866; intro'd as intra-sp. of Anodonta macrostena Servain, 1882)
Locard, 1884. Contr. Faune Malacol. Franc. 8: 15. [infra-subsp.]

Anodonta oblonga var. major Locard, 1884

(intro'd as intra-sp. of Anodonta oblonga Millet, 1831; homonym of Anodonta rostrata var. major Brusina, 1866)
Locard, 1884. Contr. Faune Malacol. Franc. 8: 19. [infra-subsp.]

Anodonta oblonga var. elongata Locard, 1884

(homonym of Anodonta elongata Holandre, 1836; intro'd as intra-sp. of Anodonta oblonga Millet, 1831)
Locard, 1884. Contr. Faune Malacol. Franc. 8: 19. [infra-subsp.]

Anodonta oblonga var. curta Locard, 1884

(intro'd as intra-sp. of Anodonta oblonga Millet, 1831; homonym of Anodonta macrostena var. curta Servain, 1882)
Locard, 1884. Contr. Faune Malacol. Franc. 8: 19. [infra-subsp.]

Anodonta schroederi Bourguignat in Schröder, 1885

Bourguignat in Schröder, 1885. Bull. Soc. Malac. France 2: 212.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta potimia Schröder, 1885

Schröder, 1885. Bull. Soc. Malac. France 2: 214.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta perlora Servain in Schröder, 1885

Servain in Schröder, 1885. Bull. Soc. Malac. France 2: 217.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta tricassinaeformis Schröder, 1885

Schröder, 1885. Bull. Soc. Malac. France 2: 218 [as ‘tricassinæformis’].

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta rhynchonella Schröder, 1885

Schröder, 1885. Bull. Soc. Malac. France 2: 220.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta journeopsis Schröder, 1885

Schröder, 1885. Bull. Soc. Malac. France 2: 222.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta josei Bourguignat in da Silva e Castro, 1885

Bourguignat in da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 278.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta enhydra da Silva e Castro, 1885

da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 279.

Anodonta (s.s.) cygnea — Haas (1969)
Anodonta cygnea — Araujo et al. (2009)

Anodonta tuberculata da Silva e Castro, 1885

da Silva e Castro, 1885. Bull. Soc. Malac. France 2: 281.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta epipedacra Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 335.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta eupelina Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 336.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta eucaca Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 338.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta foeda Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 340 [as ‘fœda’].

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta immunda Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 341.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta misara Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 343.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta rayopsis Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 344.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta duregica Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 346.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta tigurica Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 348.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta ruvidella Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 349.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta hypaeschra Servain, 1885

Servain, 1885. Bull. Soc. Malac. France 2: 351 [as ‘hypæschra’].

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta ambrosiana Adami, 1886

Adami, 1886. Bull. Soc. Malac. Ital. 11: 250, fig. 30.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cobelliana Adami, 1886

Adami, 1886. Bull. Soc. Malac. Ital. 11: 252, fig. 31.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cobelliana var. toblinica Adami, 1886

(intro'd as intra-sp. of Anodonta cobelliana Adami, 1886)
Adami, 1886. Bull. Soc. Malac. Ital. 11: 254.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta strobeliana Adami, 1886

Adami, 1886. Bull. Soc. Malac. Ital. 11: 255, fig. 32.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta strobeliana var. medoacensis Adami, 1886

(intro'd as intra-sp. of Anodonta strobeliana Adami, 1886)
Adami, 1886. Bull. Soc. Malac. Ital. 11: 257.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta zenii Adami, 1886

Adami, 1886. Bull. Soc. Malac. Ital. 11: 258, fig. 28.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta coupha Servain, 1887

Servain, 1887. Bull. Soc. Malac. France 4: 261.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta nocturna Servain, 1888

Servain, 1888. Bull. Soc. Malac. France 5: 327.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta quadrangulata Servain, 1888

Servain, 1888. Bull. Soc. Malac. France 5: 328, pl. 9, fig. 1.

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta visurgisina Bourguignat in Servain, 1888

Bourguignat in Servain, 1888. Bull. Soc. Malac. France 5: 332, pl. 8, fig. 3.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta moctera Servain, 1888

Servain, 1888. Bull. Soc. Malac. France 5: 334, pl. 9, fig. 4.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta eusomata Servain, 1888

Servain, 1888. Bull. Soc. Malac. France 5: 335.

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta maxima Drouët, 1889

(reference to Mytilus maxima Schröter, 1779)
‘Schrœter’ Drouët, 1889. Mém. Acad. Sci. Dijon 1: 84.

Anodonta cygnaea [sic] forme maxima — Germain (1931)

Anodonta cellensis var. solearis forma vasta Westerlund, 1890

(intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791; intro'd as intra-sp. of Anodonta cellensis var. solearis Küster, 1839)
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 212. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. ovata forma firma Westerlund, 1890

(intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791; intro'd as intra-sp. of Anodonta ovata Kuster of Authors)
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 213. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. ovata forma gracilis Westerlund, 1890

(homonym of Anodonta gracilis Lea, 1851; intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791; intro'd as intra-sp. of Anodonta ovata Kuster of Authors)
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 213. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. ovata forma ampla Westerlund, 1890

(intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791; intro'd as intra-sp. of Anodonta ovata Kuster of Authors)
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 213. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. ovata forma herbea Westerlund, 1890

(intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791; intro'd as intra-sp. of Anodonta ovata Kuster of Authors)
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 213. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. ovata forma tenella Westerlund, 1890

(intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791; intro'd as intra-sp. of Anodonta ovata Kuster of Authors)
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 213. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. ovata forma tenera Westerlund, 1890

(intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791; intro'd as intra-sp. of Anodonta ovata Kuster of Authors)
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 213. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. ovata forma erosa Westerlund, 1890

(intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791; intro'd as intra-sp. of Anodonta ovata Kuster of Authors)
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 213. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. ovata forma pallida Westerlund, 1890

(intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791; intro'd as intra-sp. of Anodonta ovata Kuster of Authors; homonym of Anodon pallida J.G. Anthony, 1865)
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 214. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta cellensis var. ovata forma coacta Westerlund, 1890

(intro'd as intra-sp. of Mytilus zellensis Gmelin, 1791; intro'd as intra-sp. of Anodonta ovata Kuster of Authors)
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 214. [infra-subsp.]

Anodonta (s.s.) cygnea — Haas (1969)

Anodonta catocyrta Locard, 1890

‘Coutagne’ Locard, 1890. Contr. Faune Malacol. Franc. 14: 65 [21], 150 [106].
‘Coutagne’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 229.
LECTOTYPE MNHN_22173 (by Graf, 2011: 467), a Grosne, à La Ferté (Saône-et-Loire).

Anodonta (Gravidiana) catocyrta — Germain (1904)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta gabilloti Locard, 1890

Locard, 1890. Contr. Faune Malacol. Franc. 14: 67 [23], 151 [107].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 231.
LECTOTYPE MNHN_22230 (by Graf, 2011: 476), La Loire, à Balbigny (Loire).

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta trinurcina Locard, 1890

Locard, 1890. Contr. Faune Malacol. Franc. 14: 70 [26], 153 [109].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 234.
LECTOTYPE MNHN_22589 (by Graf, 2011: 509), Tournus (Saône-et-Loire).

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta annesiaca Locard, 1890

Locard, 1890. Contr. Faune Malacol. Franc. 14: 72 [28], 154 [110].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 236.
LECTOTYPE MNHN_22102 (by Graf, 2011: 458), lac d’Annecy (Savoie).

Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta noeli Locard, 1890

Dupuy, 1852. Hist. Natur. Moll. France (6): pl. 18, fig. 13 [as ‘Unio oblonga’].
‘Bourguignat et Locard’ Locard, 1890. Contr. Faune Malacol. Franc. 14: 73 [29], 155 [111].
‘Bourguignat et Locard’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 236, fig. 243.
‘Bourguignat et Locard’ Locard, 1899. Conchyliol. Portug.: 269.
LECTOTYPE MNHN_22406 (by Graf, 2011: 493), La Saône, à Auxonne (Côte-d’Or).

Anodonta noeli — Rochebrune (1903)
Anodonta (Cygnaeana) noeli — Germain (1904)
Anodonta cygnaea [sic] — Germain (1931)
Anodonta (s.s.) cygnea — Haas (1969)

Anodonta (Liouvillea) pallaryi — Bédé (1932), Haas (1969), Haas (1969)
Anodonta pallaryi — Thiele (1934), IUCN (2010), Van Damme et al. (2010)
Anodonta cygnea — Haas (1940), Van Damme (1984), Khalloufi & Boumaïza (2005), Graf & Cummings (2011)
Anodonta cygnaea [sic] — Daget (1998)
Anodonta anatina — Gomes-dos-Santos et al. (2019)

Anodonta (Liouvillea) theryi Bédé, 1932

Bédé, 1932. Bull. Soc. Sci. Nat. Maroc 12: 226, text fig. 3.

Anodonta (Liouvillea) theryi — Bédé (1932)
Anodonta cygnea — Haas (1940), Van Damme (1984), Khalloufi & Boumaïza (2005)
Anodonta (Liouvillea) pallaryi — Haas (1969)
Anodonta cygnaea [sic] — Daget (1998)
Anodonta pallaryi — IUCN (2010)

Anodonta cellensis [sic] micheli — Modell (1945)
Anodonta (s.s.) cygnea — Haas (1969)
Anodonta (s.s.) zellensis micheli — Stadnichenko (1984)
Anodonta zellensis micheli — Kantor & Sysoev (2005), Kantor et al. (2010)
Anodonta (s.s.) zellensis — Vinarski & Kantor (2016)
Anodonta cygnea — Nagel (2025)

Taxonomic History (since 1900)

Anodonta cygnea — Simpson (1900), Kobelt (1904), Hannibal (1912), Haas (1940), Modell (1942), ... Nagel (2025), Campbell & Lydeard (2026), Wu et al. (2026), Prie et al. (2026), Wu et al. (2026)
Anodonta lucasi — Simpson (1900), IUCN (2010), Van Damme et al. (2010)
Anodonta numidica — Simpson (1900)
Anodonta grandis var. gigantea — Simpson (1900)
Anodonta cygnea var. anatina — Simpson (1900)
Anodonta gallica — Rochebrune (1903)
Anodonta noeli — Rochebrune (1903)
Anodonta lacuum — Rochebrune (1903)
Anodonta (Gravidiana) stagnalis — Germain (1904)
Anodonta (Gravidiana) stagnalis radiata — Germain (1904)
Anodonta (Gravidiana) catocyrta — Germain (1904)
Anodonta (Pammegaliana) eucypha — Germain (1904)
Anodonta (Ventricosiana) gallica — Germain (1904)
Anodonta (Ventricosiana) fragilissima — Germain (1904)
Anodonta (Ventricosiana) fragilissima acyrta — Germain (1904)
Anodonta (Ventricosiana) fragilissima viridis — Germain (1904)
Anodonta (Ventricosiana) joannisi — Germain (1904)
Anodonta (Cygnaeana) cygnaea [sic] — Germain (1904)
Anodonta (Cygnaeana) arenaria — Germain (1904)
Anodonta (Cygnaeana) cariosa — Germain (1904)
Anodonta (Cygnaeana) noeli — Germain (1904)
Anodonta (Cygnaeana) cariosula — Germain (1904)
Anodonta (Cygnaeana) cariosula elongata — Germain (1904)
Anodonta (Cygnaeana) condatina — Germain (1904)
Anodonta (Cygnaeana) marsolinae — Germain (1904)
Anodonta (Vietuliana) vietula — Germain (1904)
Anodonta (Macrosteniana) macrostena — Germain (1904)
Anodonta (Incrassatiana) incrassatus — Germain (1904)
Anodonta (Intermediana) intermedia — Germain (1904)
Anodonta (Jourdheuiliana) oblonga — Germain (1904)
Anodonta (Arealiana) arealis — Germain (1904)
Anodonta (Avoniana) avonica — Germain (1904)
Anodonta (Tricassiniana) maculata — Germain (1904)
Anodonta (Anatiniana) chivoti — Germain (1904)
Anodonta (Groupe des Cygneana) [sic] cygnea — Caziot (1907)
Anodonta (Groupe des Cygneana) [sic] arenaria — Caziot (1907)
Anodonta (Groupe des Westerlundiana) subluxata — Caziot (1907)
Anodonta piscinalis plattenica — Kobelt & Haas (1910)
Anodonta cygneus — Ortmann (1912), Heard & Guckert (1970), Kantor & Sysoev (2005)
“Anodonta cygnea” ** — Hannibal (1912)
Anodonta (Group of A. cygnea) cygnea — Simpson (1914)
Anodonta (Group of A. cygnea) lucasi — Simpson (1914)
Anodonta (Group of A. cygnea) numidica — Simpson (1914)
Anodonta cygnaea [sic] — Germain (1931), Daget (1998), Dhora (2009)
Anodonta cygnaea [sic] forme arenaria — Germain (1931)
Anodonta cygnaea [sic] forme fragilissima — Germain (1931)
Anodonta cygnaea [sic] forme ventricosa — Germain (1931)
Anodonta cygnaea [sic] forme maxima — Germain (1931)
Anodonta cygnaea [sic] forme sturmi — Germain (1931)
Anodonta anatina — Germain (1931), Gomes-dos-Santos et al. (2019), Nagel (2025)
Anodonta avonensis — Germain (1931)
Anodonta avonensis var. subluxata — Germain (1931)
Anodonta intermedia — Germain (1931)
Anodonta maculata — Germain (1931)
Anodonta (Liouvillea) pallaryi — Bédé (1932), Haas (1969), Haas (1969)
Anodonta (Liouvillea) theryi — Bédé (1932)
Anodonta pallaryi — Thiele (1934), IUCN (2010), Van Damme et al. (2010)
Anodonta (s.s.) cygnea — Zhadin (1938), Haas (1969), Haas (1969), Kat (1983), Stadnichenko (1984), Falkner (1994), Vinarski & Kantor (2016)
Anodonta (s.s.) cellensis [sic] — Zhadin (1938), Zhadin (1965)
Anodonta (s.s.) cellensis [sic] var. maganica — Zhadin (1938)
Anodonta (s.s.) cellensis [sic] var. cariosa — Zhadin (1938)
Anodonta palustris gravida — Modell (1945)
Anodonta anatina platyrhyncha [sic] — Modell (1945)
Anodonta anatina avonensis — Modell (1945)
Anodonta anatina intermedia — Modell (1945)
Anodonta cellensis cellensis [sic] — Modell (1945)
Anodonta cellensis [sic] stagnalis — Modell (1945)
Anodonta cellensis [sic] solearis — Modell (1945)
Anodonta cellensis [sic] deplanata — Modell (1945)
Anodonta cellensis [sic] micheli — Modell (1945)
Anodonta cellensis [sic] eucypha — Modell (1945)
Anodonta (s.s.) cygneus — Zhadin (1965), Habe (1977)
Anodonta stagnalis — Zatravkin (1983), Starobogatov et al. (2004), Kantor & Sysoev (2005), Gozhik (2006), Kantor et al. (2010), Mezhzherin et al. (2014)
Anodonta zellensis — Zatravkin (1983), Starobogatov et al. (2004), Mezhzherin et al. (2014)
Anodonta (s.s.) zellensis micheli — Stadnichenko (1984)
Anodonta (s.s.) stagnalis — Stadnichenko (1984), Vinarski & Kantor (2016)
Anodonta cellensis — Petr (1992)
Anodonta (s.s.) cygnea gravida — Falkner (1994), Bank (2006)
Anodonta (s.s.) cygnea cygnea — Falkner et al. (2001), Falkner et al. (2002)
Anodonta (s.s.) cygnea solearis — Falkner et al. (2001)
Anodonta (s.s.) cygnea deplanata — Falkner et al. (2001)
Anodonta zellensis zellensis — Kantor & Sysoev (2005), Kantor et al. (2010)
Anodonta zellensis micheli — Kantor & Sysoev (2005), Kantor et al. (2010)
Anodonta (s.s.) anatina watersoni — Bank (2006)
Anodonta rellensis [sic] — Gozhik (2006)
Colletopterum oblongum — Kantor et al. (2010)
Anodonta parmata — Kantor et al. (2010)
Anodonta (s.s.) zellensis — Vinarski & Kantor (2016)
Anodonta cygnea cygnea — Bogatov (2022)
Anodonta cygnea zellensis — Bogatov (2022)
Anodonta cygnea stagnalis — Bogatov (2022)

MUSSELpdb

Taxonomy

Geography

species Anodonta cygnea (Linnaeus, 1758): synonymy

Synonymy