MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
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Species Richness & Endemism of
freshwater mussels
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Data last updated
Tue 02 June 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini | Cambarunio
species Cambarunio simus (Lea, 1838): synonymy
3 specimens + 2 GenBank vouchers
Synonymy
Unio cumberlandicus Lea, 1838
(renamed Margaron (Unio) cumberlandianus Lea, 1852)
Lea, 1836. Synopsis Fam. Naiades: 27 [as ‘cumberlandianus’]. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 25, pl. 7, fig. 19.
Type(s): HOLOTYPE USNM_85254, Tenn.
Lampsilis (sec. Eurynia) nebulosus — Simpson (1900)
Lampsilis (Eurynia) nebulosa — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Ligumia nebulosa — Haas (1969)
Villosa iris — Parmalee & Bogan (1998)
Cambarunio simus — Watters (2018)
Unio notatus Lea, 1838
Lea, 1836. Synopsis Fam. Naiades: 26. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 28, pl. 8, fig. 22.
Type(s): HOLOTYPE (MONOTYPY) USNM_85075, Cumberland R., Tenn.
Lampsilis (sec. Eurynia) nebulosus — Simpson (1900)
Lampsilis (Eurynia) nebulosa — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Micromya nebulosa — Ortmann (1924)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Ligumia nebulosa — Haas (1969)
Villosa nebulosa — Burch (1975)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Cambarunio simus — Watters (2018)
Unio simus Lea, 1838
Lea, 1836. Synopsis Fam. Naiades: 29. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 26, pl. 8, fig. 20.
Type(s): HOLOTYPE USNM_85342, Cumberland R., Tenn.
Lampsilis (sec. Eurynia) simus — Simpson (1900)
Lampsilis (Eurynia) sima — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Micromya nebulosa — Ortmann (1924)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Ligumia nebulosa — Haas (1969)
Villosa nebulosa — Burch (1975)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Villosa sima — Williams et al. (2017)
Cambarunio simus — Watters (2018), Burlakova et al. (2019), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023), FMCS (2025), Campbell & Lydeard (2026)
Margaron (Unio) cumberlandianus Lea, 1852
(new name for Unio cumberlandicus Lea, 1838)
Lea, 1836. Synopsis Fam. Naiades: 27 [as ‘Margarita (Unio) cumberlandianus’]. [nomen nudum]
Lea, 1838. Synopsis Fam. Naiades: 20 [as ‘Margarita (Unio) cumberlandianus’]. [nomen nudum]
Lea, 1852. Synopsis Fam. Naiades: 30.
Lampsilis (sec. Eurynia) nebulosus — Simpson (1900)
Lampsilis (Eurynia) nebulosa — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Micromya nebulosa — Ortmann (1924)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Ligumia nebulosa — Haas (1969)
Villosa nebulosa — Burch (1975)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
Villosa iris ‘Lea’ of Authors
(misidentified reference to Unio iris Lea, 1829)
‘Lea’ of Authors.
Campbell et al., 2005. Invert. Biol. 124(2): 155. [in part]
Zanatta & Murphy, 2006. MPE 41(1): 198. [in part]
Cambarunio simus — Burlakova et al. (2019)
Unio notatus Conrad, 1853
(reference to Unio notatus Lea, 1838)
‘Lea’ Conrad, 1853. Proc. Acad. Nat. Sci. 6: 253.
Lampsilis (sec. Eurynia) simus — Simpson (1900)
Lampsilis (Eurynia) sima — Simpson (1914)
Unio proximus Küster, 1861
(misidentified reference to Unio proximus Lea, 1852)
‘Lea’ Küster, 1861. Conchyl. Cab. 9(2): 248, pl. 83, fig. 4.
Lampsilis (sec. Eurynia) simus — Simpson (1900)
Lampsilis (Eurynia) sima — Simpson (1914)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Villosa iris — Parmalee & Bogan (1998)
Unio spatulatus Sowerby, 1868
(misidentified reference to Unio spatulatus Lea, 1845)
‘Lea’ Sowerby, 1868. Conch. Icon. 16: pl. 65, sp. 328.
Lampsilis (sec. Eurynia) simus — Simpson (1900)
Lampsilis (Eurynia) sima — Simpson (1914)
Villosa iris — Parmalee & Bogan (1998)
Taxonomic History (since 1900)
Lampsilis (sec. Eurynia) nebulosus — Simpson (1900)
Lampsilis (sec. Eurynia) simus — Simpson (1900)
Lampsilis (Eurynia) nebulosa — Simpson (1914)
Lampsilis (Eurynia) sima — Simpson (1914)
Eurynia nebulosus — Ortmann (1918)
Micromya nebulosa — Ortmann (1924)
Lampsilis (Ligumia) nebulosa — Frierson (1927)
Ligumia nebulosa — Haas (1969)
Villosa nebulosa — Burch (1975)
Villosa iris — Parmalee & Bogan (1998), Williams et al. (2008)
“Villosa iris” ** — Campbell et al. (2005), Zanatta & Murphy (2006)
Villosa sima — Williams et al. (2017)
Cambarunio simus — Watters (2018), Burlakova et al. (2019), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023), FMCS (2025), Campbell & Lydeard (2026)
