Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina

genus Pseudodon Gould, 1844: cladomics

9 Recent valid species (9 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 9 (9 total)
analyzed in 98 cladograms in 58 publications
maximum number of terminals in an analysis = 7 (mean = 1.79, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 32
maximum number of included terminals = 7, (mean = 3.41 median = 3)
proportion of cladograms that support basic monophyly = 0.88

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pseudodon cladogram distances
PCoA Ordination

NJ Tree of Pseudodon cladogram distances
NJ Tree

Cladograms

Zieritz et al., 2026 [Publ 4812], fig. 5a

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
7 7 7 9 1 1 1 0 
  +-------------------- Namkongnaia               
  |                                               
--+  +----------------- Pseudodon_mekongi **         
  |  |                                            
  |  |           +----- Pseudodon_lenyanensis__2 **  
  +--+  +--------+                                
     |  |        |  +-- Pseudodon_inoscularis **     
     |  |        +--+                             
     +--+           +-- Pseudodon_lenyanensis__1 **  
        |                                         
        |  +----------- Pseudodon_vondembuschianus **
        +--+                                      
           |  +-------- Pseudodon_vagulus **         
           +--+                                   
              |  +----- Pseudodon_cokelatus__2 **    
              +--+                                
                 |  +-- Pseudodon_walpolei **        
                 +--+                             
                    +-- Pseudodon_cokelatus__1 **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
6 7 6 7 624 34 9 10 
                          +-- Lannanaia                 
  +-----------------------+                             
  |                       +-- Pilsbryoconcha            
  |                                                     
--+  +----------------------- Songkhlanaia              
  |  |                                                  
  |  |                    +-- Nyeinchanconcha           
  +--+  +-----------------+                             
     |  |                 +-- Sundadontina              
     |  |                                               
     |  |                 +-- Isannaia                  
     +--+              +--+                             
        |              |  +-- Namkongnaia               
        |  +-----------+                                
        |  |           |  +-- Thaiconcha                
        |  |           +--+                             
        +--+              +-- Bineurus                  
           |                                            
           |  +-------------- Pseudodon_walpolei **        
           |  |                                         
           +--+  +----------- Pseudodon_vagulus **         
              |  |                                      
              |  |     +----- Pseudodon_vondembuschianus **
              +--+  +--+                                
                 |  |  |  +-- Pseudodon_mekongi **         
                 |  |  +--+                             
                 +--+     +-- Schepmania                
                    |                                   
                    |  +----- Pseudodon_lenyanensis__2 **  
                    +--+                                
                       |  +-- Pseudodon_inoscularis **     
                       +--+                             
                          +-- Pseudodon_lenyanensis__1 **

Jeratthitikul et al., 2025 [Publ 4476], fig. 2

iTtTbiriSoTroTroSiD
7 7 5 7 8 8 4 10 
  +-------------------- Pilsbryoconcha            
  |                                               
  |              +----- Namkongnaia               
--+  +-----------+                                
  |  |           |  +-- Isannaia                  
  |  |           +--+                             
  +--+              +-- Sundadontina              
     |                                            
     |  +-------------- Pseudodon_walpolei **        
     |  |                                         
     +--+           +-- Pseudodon_inoscularis **     
        |  +--------+                             
        |  |        +-- Pseudodon_lenyanensis **     
        +--+                                      
           |  +-------- Pseudodon_mekongi **         
           +--+                                   
              |  +----- Pseudodon_vondembuschianus **
              +--+                                
                 |  +-- Pseudodon_vagulus **         
                 +--+                             
                    +-- Pseudodon_septuaginta **

Zieritz et al., 2024 [Publ 4381], fig. 8A

iTtTbiriSoTroTroSiD
6 6 4 6 1 1 1 3 
  +-------------- Namkongnaia               
  |                                         
--+  +----------- Pseudodon_mekongi **         
  |  |                                      
  +--+        +-- Pseudodon_lenyanensis **     
     |  +-----+                             
     |  |     +-- Pseudodon_inoscularis **     
     +--+                                   
        |  +----- Pseudodon_vondembuschianus **
        +--+                                
           |  +-- Pseudodon_walpolei **        
           +--+                             
              +-- Pseudodon_vagulus **

Bolotov et al., 2023 [Publ 4312], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 5 33 28 7 4 
  +-------------------- Pilsbryoconcha            
  |                                               
--+  +----------------- Songkhlanaia              
  |  |                                            
  |  |        +-------- Sundadontina              
  +--+  +-----+                                   
     |  |     |  +----- Nyeinchanconcha           
     |  |     +--+                                
     |  |        |  +-- Thaiconcha                
     +--+        +--+                             
        |           +-- Bineurus                  
        |                                         
        |  +----------- Namkongnaia               
        |  |                                      
        +--+  +-------- Pseudodon_mekongi **         
           |  |                                   
           +--+     +-- Pseudodon_vondembuschianus **
              |  +--+                             
              |  |  +-- Pseudodon_vagulus **         
              +--+                                
                 |  +-- Pseudodon_lenyanensis **     
                 +--+                             
                    +-- Pseudodon_inoscularis **

Bolotov et al., 2023 [Publ 4312], fig. S2

iTtTbiriSoTroTroSiD
5 5 3 5 35 30 7 8 
  +-------------------- Songkhlanaia              
  |                                               
  |                 +-- Nyeinchanconcha           
--+  +--------------+                             
  |  |              +-- Sundadontina              
  |  |                                            
  +--+           +----- Namkongnaia               
     |  +--------+                                
     |  |        |  +-- Thaiconcha                
     |  |        +--+                             
     +--+           +-- Bineurus                  
        |                                         
        |  +----------- Pilsbryoconcha            
        |  |                                      
        +--+  +-------- Pseudodon_vondembuschianus **
           |  |                                   
           +--+     +-- Pseudodon_mekongi **         
              |  +--+                             
              |  |  +-- Pseudodon_vagulus **         
              +--+                                
                 |  +-- Pseudodon_lenyanensis **     
                 +--+                             
                    +-- Pseudodon_inoscularis **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 54 31 9 6 
              +-- Pseudodon_inoscularis **     
        +-----+                             
        |     +-- Pseudodon_lenyanensis **     
  +-----+                                   
  |     |  +----- Pseudodon_mekongi **         
  |     +--+                                
  |        |  +-- Pseudodon_vagulus **         
  |        +--+                             
  |           +-- Pseudodon_vondembuschianus **
--+                                         
  |     +-------- Sundadontina              
  |  +--+                                   
  |  |  |  +----- Nyeinchanconcha           
  |  |  +--+                                
  |  |     |  +-- Thaiconcha                
  |  |     +--+                             
  +--+        +-- Bineurus                  
     |                                      
     |        +-- Isannaia                  
     |  +-----+                             
     |  |     +-- Pilsbryoconcha            
     +--+                                   
        |  +----- Namkongnaia               
        +--+                                
           |  +-- Lannanaia                 
           +--+                             
              +-- Songkhlanaia

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 45 33 9 6 
                          +-- Lannanaia                 
  +-----------------------+                             
  |                       +-- Pilsbryoconcha            
  |                                                     
--+  +----------------------- Songkhlanaia              
  |  |                                                  
  |  |                 +----- Nyeinchanconcha           
  +--+  +--------------+                                
     |  |              |  +-- Thaiconcha                
     |  |              +--+                             
     +--+                 +-- Bineurus                  
        |                                               
        |  +----------------- Sundadontina              
        |  |                                            
        +--+  +-------------- Isannaia                  
           |  |                                         
           +--+  +----------- Namkongnaia               
              |  |                                      
              +--+        +-- Pseudodon_inoscularis **     
                 |  +-----+                             
                 |  |     +-- Pseudodon_lenyanensis **     
                 +--+                                   
                    |  +----- Pseudodon_mekongi **         
                    +--+                                
                       |  +-- Pseudodon_vagulus **         
                       +--+                             
                          +-- Pseudodon_vondembuschianus **

Konopleva et al., 2025 [Publ 4446], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 39 34 9 6 
  +-------------------------- Lannanaia                 
  |                                                     
--+  +----------------------- Pilsbryoconcha            
  |  |                                                  
  +--+  +-------------------- Songkhlanaia              
     |  |                                               
     |  |              +----- Nyeinchanconcha           
     +--+  +-----------+                                
        |  |           |  +-- Thaiconcha                
        |  |           +--+                             
        |  |              +-- Bineurus                  
        +--+                                            
           |  +-------------- Sundadontina              
           |  |                                         
           |  |           +-- Namkongnaia               
           +--+  +--------+                             
              |  |        +-- Isannaia                  
              |  |                                      
              +--+        +-- Pseudodon_inoscularis **     
                 |  +-----+                             
                 |  |     +-- Pseudodon_lenyanensis **     
                 +--+                                   
                    |  +----- Pseudodon_mekongi **         
                    +--+                                
                       |  +-- Pseudodon_vondembuschianus **
                       +--+                             
                          +-- Pseudodon_vagulus **

Bolotov et al., 2023 [Publ 4312], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 33 28 7 6 
  +----------------------- Pilsbryoconcha            
  |                                                  
  |                 +----- Nyeinchanconcha           
--+  +--------------+                                
  |  |              |  +-- Thaiconcha                
  |  |              +--+                             
  +--+                 +-- Bineurus                  
     |                                               
     |  +----------------- Songkhlanaia              
     |  |                                            
     +--+  +-------------- Sundadontina              
        |  |                                         
        +--+  +----------- Namkongnaia               
           |  |                                      
           +--+        +-- Pseudodon_inoscularis **     
              |  +-----+                             
              |  |     +-- Pseudodon_lenyanensis **     
              +--+                                   
                 |  +----- Pseudodon_mekongi **         
                 +--+                                
                    |  +-- Pseudodon_vondembuschianus **
                    +--+                             
                       +-- Pseudodon_vagulus **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
3 6 4 3 136 9 5 10 
        +-- Pseudodon_mekongi **         
  +-----+                             
  |     +-- Pseudodon_vondembuschianus **
  |                                   
--+     +-- Pseudodon_inoscularis **     
  |  +--+                             
  |  |  +-- Sundadontina_tumida       
  +--+                                
     |  +-- Nyeinchanconcha           
     +--+                             
        +-- Sundadontina_cumingii

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 108 13 5 8 
  +----------------- Bineurus                  
  |                                            
--+  +-------------- Nyeinchanconcha           
  |  |                                         
  +--+           +-- Thaiconcha                
     |  +--------+                             
     |  |        +-- Pilsbryoconcha            
     +--+                                      
        |  +-------- Sundadontina              
        +--+                                   
           |  +----- Pseudodon_lenyanensis **     
           +--+                                
              |  +-- Pseudodon_mekongi **         
              +--+                             
                 +-- Pseudodon_vondembuschianus **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 31 13 6 8 
              +----- Pilsbryoconcha_exilis       
  +-----------+                                  
  |           |  +-- Pilsbryoconcha_linguaeformis
  |           +--+                               
  |              +-- Namkongnaia_lemeslei__2     
--+                                              
  |           +----- Nyeinchanconcha             
  |  +--------+                                  
  |  |        |  +-- Thaiconcha                  
  |  |        +--+                               
  +--+           +-- Bineurus                    
     |                                           
     |  +----------- Sundadontina                
     |  |                                        
     +--+        +-- Namkongnaia_lemeslei__1     
        |  +-----+                               
        |  |     +-- Namkongnaia_inkhavilayi     
        +--+                                     
           |  +----- Pseudodon_lenyanensis **       
           +--+                                  
              |  +-- Pseudodon_mekongi **           
              +--+                               
                 +-- Pseudodon_vondembuschianus **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 32 14 6 8 
  +----------------- Pilsbryoconcha            
  |                                            
  |              +-- Nyeinchanconcha           
  |           +--+                             
--+           |  +-- Sundadontina_cumingii     
  |  +--------+                                
  |  |        |  +-- Thaiconcha                
  |  |        +--+                             
  |  |           +-- Bineurus                  
  +--+                                         
     |        +----- Sundadontina_tumida       
     |  +-----+                                
     |  |     |  +-- Sundadontina_brandti      
     |  |     +--+                             
     +--+        +-- Sundadontina_taskaevi     
        |                                      
        |  +-------- Namkongnaia               
        +--+                                   
           |  +----- Pseudodon_lenyanensis **     
           +--+                                
              |  +-- Pseudodon_mekongi **         
              +--+                             
                 +-- Pseudodon_vondembuschianus **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
3 3 1 3 42 11 5 8 
           +-- Nyeinchanconcha           
  +--------+                             
  |        +-- Bineurus                  
  |                                      
  |        +-- Thaiconcha                
--+--------+                             
  |        +-- Sundadontina              
  |                                      
  |  +-------- Pilsbryoconcha            
  +--+                                   
     |  +----- Pseudodon_lenyanensis **     
     +--+                                
        |  +-- Pseudodon_mekongi **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 23 15 6 8 
           +----- Pilsbryoconcha_exilis__2  
  +--------+                                
  |        |  +-- Pilsbryoconcha_exilis__1  
  |        +--+                             
  |           +-- Namkongnaia               
--+                                         
  |        +----- Nyeinchanconcha           
  |  +-----+                                
  |  |     |  +-- Thaiconcha                
  |  |     +--+                             
  +--+        +-- Bineurus                  
     |                                      
     |  +-------- Sundadontina              
     +--+                                   
        |  +----- Pseudodon_lenyanensis **     
        +--+                                
           |  +-- Pseudodon_mekongi **         
           +--+                             
              +-- Pseudodon_vondembuschianus **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 114 13 5 8 
  +----------------- Bineurus                  
  |                                            
--+  +-------------- Nyeinchanconcha           
  |  |                                         
  +--+           +-- Thaiconcha                
     |  +--------+                             
     |  |        +-- Pilsbryoconcha            
     +--+                                      
        |  +-------- Sundadontina              
        +--+                                   
           |  +----- Pseudodon_lenyanensis **     
           +--+                                
              |  +-- Pseudodon_mekongi **         
              +--+                             
                 +-- Pseudodon_vondembuschianus **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
3 3 1 3 42 11 5 8 
              +-- Nyeinchanconcha           
  +-----------+                             
  |           +-- Bineurus                  
  |                                         
--+           +-- Thaiconcha                
  |  +--------+                             
  |  |        +-- Sundadontina              
  +--+                                      
     |  +-------- Pilsbryoconcha            
     +--+                                   
        |  +----- Pseudodon_lenyanensis **     
        +--+                                
           |  +-- Pseudodon_mekongi **         
           +--+                             
              +-- Pseudodon_vondembuschianus **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
3 3 1 3 79 22 7 8 
  +-------------------- Pilsbryoconcha            
  |                                               
--+  +----------------- Songkhlanaia              
  |  |                                            
  +--+  +-------------- Namkongnaia               
     |  |                                         
     |  |        +----- Pseudodon_lenyanensis **     
     +--+  +-----+                                
        |  |     |  +-- Pseudodon_mekongi **         
        |  |     +--+                             
        +--+        +-- Pseudodon_vondembuschianus **
           |                                      
           |  +-------- Sundadontina              
           +--+                                   
              |  +----- Nyeinchanconcha           
              +--+                                
                 |  +-- Thaiconcha                
                 +--+                             
                    +-- Bineurus

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 38 4 4 -- 
  +----------- Pilsbryoconcha            
  |                                      
--+        +-- Pseudodon_mekongi **         
  |  +-----+                             
  |  |     +-- Pseudodon_vondembuschianus **
  +--+                                   
     |  +----- Sundadontina              
     +--+                                
        |  +-- Nyeinchanconcha           
        +--+                             
           +-- Bineurus

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 123 9 5 -- 
  +-------- Bineurus                  
  |                                   
  |     +-- Nyeinchanconcha           
  +-----+                             
  |     +-- Sundadontina_cumingii     
--+                                   
  |     +-- Thaiconcha                
  +-----+                             
  |     +-- Pilsbryoconcha            
  |                                   
  |  +----- Sundadontina_tumida       
  +--+                                
     |  +-- Pseudodon_mekongi **         
     +--+                             
        +-- Pseudodon_vondembuschianus **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
2 2 0 2 199 9 5 -- 
  +----- Nyeinchanconcha           
  |                                
  +----- Thaiconcha                
  |                                
  +----- Sundadontina_cumingii     
  |                                
  +----- Sundadontina_tumida__1    
--+                                
  +----- Sundadontina_tumida__2    
  |                                
  +----- Pilsbryoconcha            
  |                                
  +----- Bineurus                  
  |                                
  |  +-- Pseudodon_mekongi **         
  +--+                             
     +-- Pseudodon_vondembuschianus **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
2 2 0 2 197 9 5 -- 
  +----------- Pilsbryoconcha            
  |                                      
  |     +----- Sundadontina_tumida       
--+  +--+                                
  |  |  |  +-- Pseudodon_mekongi **         
  |  |  +--+                             
  |  |     +-- Pseudodon_vondembuschianus **
  +--+                                   
     |     +-- Nyeinchanconcha           
     |  +--+                             
     |  |  +-- Sundadontina_cumingii     
     +--+                                
        |  +-- Thaiconcha                
        +--+                             
           +-- Bineurus

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 3 46 9 5 -- 
  +-------------- Pilsbryoconcha               
  |                                            
  |        +----- Pseudodon_vondembuschianus__2 **
--+  +-----+                                   
  |  |     |  +-- Pseudodon_mekongi **            
  |  |     +--+                                
  +--+        +-- Pseudodon_vondembuschianus__1 **
     |                                         
     |  +-------- Sundadontina                 
     +--+                                      
        |  +----- Nyeinchanconcha              
        +--+                                   
           |  +-- Bineurus                     
           +--+                                
              +-- Thaiconcha

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 44 9 5 -- 
  +-------------- Pilsbryoconcha            
  |                                         
--+           +-- Pseudodon_mekongi **         
  |  +--------+                             
  |  |        +-- Pseudodon_vondembuschianus **
  +--+                                      
     |  +-------- Sundadontina              
     +--+                                   
        |  +----- Nyeinchanconcha           
        +--+                                
           |  +-- Bineurus                  
           +--+                             
              +-- Thaiconcha

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
2 7 0 3 49 5 2 12 
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_vondembuschianus **
  |                             
  +-- Pseudodon_mekongi__1 **      
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_mekongi__2 **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 3 45 9 5 -- 
  +-------------- Pilsbryoconcha               
  |                                            
  |        +----- Pseudodon_vondembuschianus__2 **
--+  +-----+                                   
  |  |     |  +-- Pseudodon_mekongi **            
  |  |     +--+                                
  +--+        +-- Pseudodon_vondembuschianus__1 **
     |                                         
     |  +-------- Sundadontina                 
     +--+                                      
        |  +----- Nyeinchanconcha              
        +--+                                   
           |  +-- Bineurus                     
           +--+                                
              +-- Thaiconcha

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
2 7 0 3 80 5 2 12 
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_vondembuschianus **
  |                             
  +-- Pseudodon_mekongi__1 **      
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_mekongi__2 **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 137 9 5 -- 
  +----------- Pilsbryoconcha            
  |                                      
  |     +----- Sundadontina_tumida       
--+  +--+                                
  |  |  |  +-- Pseudodon_mekongi **         
  |  |  +--+                             
  |  |     +-- Pseudodon_vondembuschianus **
  +--+                                   
     |     +-- Nyeinchanconcha           
     |  +--+                             
     |  |  +-- Sundadontina_cumingii     
     +--+                                
        |  +-- Thaiconcha                
        +--+                             
           +-- Bineurus

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 199 9 5 -- 
  +----------- Pilsbryoconcha            
  |                                      
  |     +----- Nyeinchanconcha           
--+  +--+                                
  |  |  |  +-- Thaiconcha                
  |  |  +--+                             
  +--+     +-- Bineurus                  
     |                                   
     |  +----- Sundadontina              
     +--+                                
        |  +-- Pseudodon_mekongi **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 43 5 4 -- 
     +----- Pilsbryoconcha    
  +--+                        
  |  |  +-- Pseudodon_walpolei **
  |  +--+                     
--+     +-- Pseudodon_mekongi ** 
  |                           
  |  +----- Sundadontina      
  +--+                        
     |  +-- Thaiconcha        
     +--+                     
        +-- Bineurus

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 29 4 4 -- 
  +----------- Sundadontina              
  |                                      
--+        +-- Nyeinchanconcha           
  |  +-----+                             
  |  |     +-- Bineurus                  
  +--+                                   
     |  +----- Pilsbryoconcha            
     +--+                                
        |  +-- Pseudodon_vondembuschianus **
        +--+                             
           +-- Pseudodon_mekongi **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 60 2 2 -- 
-- Pseudodon_lenyanensis **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 67 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
1 1 0 1 48 1 1 -- 
  +-- Pseudodon_lenyanensis **
--+                        
  +-- Sundadontina

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 57 2 2 -- 
-- Pseudodon_lenyanensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 5 4 -- 
  +-------- Pilsbryoconcha   
  |                          
--+  +----- Pseudodon_mekongi **
  |  |                       
  +--+  +-- Nyeinchanconcha  
     |  |                    
     +--+-- Bineurus         
        |                    
        +-- Sundadontina

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 86 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 18 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Zieritz et al., 2026 [Publ 4812], fig. 4a

iTtTbiriSoTroTroSiD
1 1 0 1 8 8 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 30 5 4 -- 
  +----------- Pilsbryoconcha   
--+                             
  |  +-------- Pseudodon_mekongi **
  +--+                          
     |  +----- Sundadontina     
     +--+                       
        |  +-- Thaiconcha       
        +--+                    
           +-- Bineurus

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 57 2 2 -- 
-- Pseudodon_lenyanensis **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 84 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 3 3 -- 
  +-------- Pilsbryoconcha            
--+                                   
  |  +----- Bineurus                  
  +--+                                
     |  +-- Pseudodon_vondembuschianus **
     +--+                             
        +-- Sundadontina

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 80 4 3 -- 
  +-------- Pilsbryoconcha            
--+                                   
  |  +----- Pseudodon_vondembuschianus **
  +--+                                
     |  +-- Bineurus                  
     +--+                             
        +-- Nyeinchanconcha

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 53 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 35 4 3 -- 
  +-------- Pilsbryoconcha   
--+                          
  |  +----- Pseudodon_mekongi **
  +--+                       
     |  +-- Thaiconcha       
     +--+                    
        +-- Bineurus

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 45 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 3 3 -- 
  +-------- Bineurus                  
--+                                   
  |  +----- Pilsbryoconcha            
  +--+                                
     |  +-- Pseudodon_vondembuschianus **
     +--+                             
        +-- Sundadontina

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 14 6 -- 
     +----- Pilsbryoconcha   
  +--+                       
  |  |  +-- Namkongnaia      
  |  +--+                    
  |     +-- Pseudodon_mekongi **
--+                          
  |     +-- Nyeinchanconcha  
  |  +--+                    
  |  |  +-- Sundadontina     
  +--+                       
     |  +-- Thaiconcha       
     +--+                    
        +-- Bineurus

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 26 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
1 1 0 1 35 4 3 -- 
  +-------- Pilsbryoconcha   
--+                          
  |  +----- Pseudodon_mekongi **
  +--+                       
     |  +-- Thaiconcha       
     +--+                    
        +-- Bineurus

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 35 4 3 -- 
  +-------- Pilsbryoconcha   
--+                          
  |  +----- Bineurus         
  +--+                       
     |  +-- Pseudodon_mekongi **
     +--+                    
        +-- Thaiconcha

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 72 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Pseudodon_mekongi **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 43 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 94 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 14 6 -- 
     +----- Pilsbryoconcha   
  +--+                       
  |  |  +-- Namkongnaia      
  |  +--+                    
  |     +-- Pseudodon_mekongi **
--+                          
  |     +-- Nyeinchanconcha  
  |  +--+                    
  |  |  +-- Sundadontina     
  +--+                       
     |  +-- Thaiconcha       
     +--+                    
        +-- Bineurus

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 23 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 60 2 2 -- 
-- Pseudodon_lenyanensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 139 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 72 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 10 6 -- 
  +----------- Bineurus         
  |                             
  |     +----- Namkongnaia      
--+  +--+                       
  |  |  |  +-- Sundadontina     
  |  |  +--+                    
  +--+     +-- Nyeinchanconcha  
     |                          
     |  +----- Pseudodon_mekongi **
     +--+                       
        |  +-- Pilsbryoconcha   
        +--+                    
           +-- Thaiconcha

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 94 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
1 1 0 1 35 4 3 -- 
  +-------- Pilsbryoconcha      
  |                             
--+  +----- Bineurus_anodontinum
  |  |                          
  +--+----- Bineurus_loeiensis  
     |                          
     |  +-- Pseudodon_mekongi **   
     +--+                       
        +-- Thaiconcha

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 72 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 43 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 2 1 -- 
  +----- Pilsbryoconcha_exilis     
--+                                
  |  +-- Pseudodon_vondembuschianus **
  +--+                             
     +-- Pilsbryoconcha_carinifera

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 94 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 90 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Pseudodon_mekongi **

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 57 2 2 -- 
-- Pseudodon_lenyanensis **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 2 1 -- 
  +----- Pilsbryoconcha_exilis     
--+                                
  |  +-- Pseudodon_vondembuschianus **
  +--+                             
     +-- Pilsbryoconcha_carinifera

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 35 4 3 -- 
  +-------- Pilsbryoconcha   
--+                          
  |  +----- Pseudodon_mekongi **
  +--+                       
     |  +-- Thaiconcha       
     +--+                    
        +-- Bineurus

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 45 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 35 4 3 -- 
  +-------- Pilsbryoconcha   
--+                          
  |  +----- Pseudodon_mekongi **
  +--+                       
     |  +-- Thaiconcha       
     +--+                    
        +-- Bineurus

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 18 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Sundadontina

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 90 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 -- 
  +-- Pilsbryoconcha   
--+                    
  +-- Pseudodon_mekongi **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Pseudodon_mekongi **
--+                    
  +-- Pilsbryoconcha