Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina

genus Pseudodon Gould, 1844: cladomics

9 Recent valid species (9 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 9 (9 total)
analyzed in 90 cladograms in 54 publications
maximum number of terminals in an analysis = 8 (mean = 2.1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 37
maximum number of included terminals = 8, (mean = 3.68 median = 3)
proportion of cladograms that support basic monophyly = 0.92

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pseudodon cladogram distances
PCoA Ordination

NJ Tree of Pseudodon cladogram distances
NJ Tree

Cladograms

Jeratthitikul et al., 2025 [Publ 4476], fig. 2

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
8 8 6 8 8 8 4 0 
  +-------------------- Pilsbryoconcha            
  |                                               
  |              +----- Namkongnaia               
--+  +-----------+                                
  |  |           |  +-- Isannaia                  
  |  |           +--+                             
  |  |              +-- Sundadontina              
  +--+                                            
     |  +-------------- Pseudodon_walpolei **        
     |  |                                         
     |  |           +-- Pseudodon_inoscularis **     
     +--+  +--------+                             
        |  |        +-- Pseudodon_lenyanensis **     
        |  |                                      
        +--+        +-- Pseudodon_mekongi **         
           |  +-----+                             
           |  |     +-- Pseudodon_vondembuschianus **
           +--+                                   
              |  +----- Pseudodon_cambodjensis **    
              +--+                                
                 |  +-- Pseudodon_vagulus **         
                 +--+                             
                    +-- Pseudodon_septuaginta **

Konopleva et al., 2025 [Publ 4446], fig. 4

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
6 6 4 6 39 34 9 2 
  +-------------------------- Lannanaia                 
  |                                                     
--+  +----------------------- Pilsbryoconcha            
  |  |                                                  
  +--+  +-------------------- Songkhlanaia              
     |  |                                               
     |  |              +----- Nyeinchanconcha           
     +--+  +-----------+                                
        |  |           |  +-- Thaiconcha                
        |  |           +--+                             
        |  |              +-- Bineurus                  
        +--+                                            
           |  +-------------- Sundadontina              
           |  |                                         
           |  |           +-- Namkongnaia               
           +--+  +--------+                             
              |  |        +-- Isannaia                  
              |  |                                      
              +--+        +-- Pseudodon_inoscularis **     
                 |  +-----+                             
                 |  |     +-- Pseudodon_lenyanensis **     
                 |  |                                   
                 +--+     +-- Pseudodon_cambodjensis **    
                    |  +--+                             
                    |  |  +-- Pseudodon_vagulus **         
                    +--+                                
                       |  +-- Pseudodon_vondembuschianus **
                       +--+                             
                          +-- Pseudodon_mekongi **

Zieritz et al., 2024 [Publ 4381], fig. 8A

iTtTbiriSoTroTroSiD
7 7 5 7 1 1 1 7 
  +-------------- Namkongnaia               
  |                                         
  |           +-- Pseudodon_vondembuschianus **
--+  +--------+                             
  |  |        +-- Pseudodon_mekongi **         
  |  |                                      
  +--+        +-- Pseudodon_lenyanensis **     
     |  +-----+                             
     |  |     +-- Pseudodon_inoscularis **     
     +--+                                   
        |  +----- Pseudodon_cambodjensis **    
        +--+                                
           |  +-- Pseudodon_walpolei **        
           +--+                             
              +-- Pseudodon_vagulus **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
6 6 4 6 45 33 9 2 
                          +-- Lannanaia                 
  +-----------------------+                             
  |                       +-- Pilsbryoconcha            
  |                                                     
--+  +----------------------- Songkhlanaia              
  |  |                                                  
  |  |                 +----- Nyeinchanconcha           
  +--+  +--------------+                                
     |  |              |  +-- Thaiconcha                
     |  |              +--+                             
     +--+                 +-- Bineurus                  
        |                                               
        |  +----------------- Sundadontina              
        |  |                                            
        +--+  +-------------- Isannaia                  
           |  |                                         
           +--+  +----------- Namkongnaia               
              |  |                                      
              |  |        +-- Pseudodon_inoscularis **     
              +--+  +-----+                             
                 |  |     +-- Pseudodon_lenyanensis **     
                 |  |                                   
                 +--+     +-- Pseudodon_vagulus **         
                    |  +--+                             
                    |  |  +-- Pseudodon_cambodjensis **    
                    +--+                                
                       |  +-- Pseudodon_vondembuschianus **
                       +--+                             
                          +-- Pseudodon_mekongi **

Bolotov et al., 2023 [Publ 4312], fig. S1

iTtTbiriSoTroTroSiD
6 6 4 6 33 28 7 2 
  +----------------------- Pilsbryoconcha            
  |                                                  
  |                 +----- Nyeinchanconcha           
--+  +--------------+                                
  |  |              |  +-- Thaiconcha                
  |  |              +--+                             
  +--+                 +-- Bineurus                  
     |                                               
     |  +----------------- Songkhlanaia              
     |  |                                            
     +--+  +-------------- Sundadontina              
        |  |                                         
        +--+  +----------- Namkongnaia               
           |  |                                      
           |  |        +-- Pseudodon_inoscularis **     
           +--+  +-----+                             
              |  |     +-- Pseudodon_lenyanensis **     
              |  |                                   
              +--+     +-- Pseudodon_cambodjensis **    
                 |  +--+                             
                 |  |  +-- Pseudodon_vagulus **         
                 +--+                                
                    |  +-- Pseudodon_vondembuschianus **
                    +--+                             
                       +-- Pseudodon_mekongi **

Bolotov et al., 2023 [Publ 4312], fig. 2

iTtTbiriSoTroTroSiD
6 6 5 7 33 28 7 5 
  +-------------------- Pilsbryoconcha               
  |                                                  
  |  +----------------- Songkhlanaia                 
--+  |                                               
  |  |        +-------- Sundadontina                 
  |  |  +-----+                                      
  +--+  |     |  +----- Nyeinchanconcha              
     |  |     +--+                                   
     |  |        |  +-- Thaiconcha                   
     |  |        +--+                                
     +--+           +-- Bineurus                     
        |                                            
        |  +----------- Namkongnaia                  
        |  |                                         
        |  |     +----- Pseudodon_vondembuschianus__2 **
        +--+  +--+                                   
           |  |  |  +-- Pseudodon_vondembuschianus__1 **
           |  |  +--+                                
           |  |     +-- Pseudodon_mekongi **            
           +--+                                      
              |     +-- Pseudodon_cambodjensis **       
              |  +--+                                
              |  |  +-- Pseudodon_vagulus **            
              +--+                                   
                 |  +-- Pseudodon_lenyanensis **        
                 +--+                                
                    +-- Pseudodon_inoscularis **

Bolotov et al., 2023 [Publ 4312], fig. S2

iTtTbiriSoTroTroSiD
6 6 4 6 35 30 7 6 
  +----------------------- Songkhlanaia              
  |                                                  
  |                    +-- Nyeinchanconcha           
--+  +-----------------+                             
  |  |                 +-- Sundadontina              
  |  |                                               
  +--+              +----- Namkongnaia               
     |  +-----------+                                
     |  |           |  +-- Thaiconcha                
     |  |           +--+                             
     +--+              +-- Bineurus                  
        |                                            
        |  +-------------- Pilsbryoconcha            
        |  |                                         
        +--+  +----------- Pseudodon_cambodjensis **    
           |  |                                      
           +--+        +-- Pseudodon_lenyanensis **     
              |  +-----+                             
              |  |     +-- Pseudodon_inoscularis **     
              +--+                                   
                 |  +----- Pseudodon_vagulus **         
                 +--+                                
                    |  +-- Pseudodon_vondembuschianus **
                    +--+                             
                       +-- Pseudodon_mekongi **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
6 6 5 7 54 31 9 3 
              +-- Pseudodon_inoscularis **        
     +--------+                                
     |        +-- Pseudodon_lenyanensis **        
     |                                         
  +--+        +-- Pseudodon_vagulus **            
  |  |  +-----+                                
  |  |  |     +-- Pseudodon_cambodjensis **       
  |  +--+                                      
  |     |  +----- Pseudodon_vondembuschianus__2 **
  |     +--+                                   
  |        |  +-- Pseudodon_vondembuschianus__1 **
  |        +--+                                
--+           +-- Pseudodon_mekongi **            
  |                                            
  |     +-------- Sundadontina                 
  |  +--+                                      
  |  |  |  +----- Nyeinchanconcha              
  |  |  +--+                                   
  |  |     |  +-- Thaiconcha                   
  |  |     +--+                                
  +--+        +-- Bineurus                     
     |                                         
     |        +-- Isannaia                     
     |  +-----+                                
     |  |     +-- Pilsbryoconcha               
     +--+                                      
        |  +----- Namkongnaia                  
        +--+                                   
           |  +-- Lannanaia                    
           +--+                                
              +-- Songkhlanaia

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
4 4 2 4 108 13 5 4 
  +-------------------- Bineurus                  
  |                                               
--+  +----------------- Nyeinchanconcha           
  |  |                                            
  +--+              +-- Thaiconcha                
     |  +-----------+                             
     |  |           +-- Pilsbryoconcha            
     +--+                                         
        |  +----------- Sundadontina              
        +--+                                      
           |  +-------- Pseudodon_lenyanensis **     
           +--+                                   
              |  +----- Pseudodon_cambodjensis **    
              +--+                                
                 |  +-- Pseudodon_vondembuschianus **
                 +--+                             
                    +-- Pseudodon_mekongi **

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 4 23 15 6 4 
              +----- Pilsbryoconcha_exilis__2  
  +-----------+                                
  |           |  +-- Pilsbryoconcha_exilis__1  
  |           +--+                             
  |              +-- Namkongnaia               
--+                                            
  |           +----- Nyeinchanconcha           
  |  +--------+                                
  |  |        |  +-- Thaiconcha                
  |  |        +--+                             
  +--+           +-- Bineurus                  
     |                                         
     |  +----------- Sundadontina              
     +--+                                      
        |  +-------- Pseudodon_lenyanensis **     
        +--+                                   
           |  +----- Pseudodon_cambodjensis **    
           +--+                                
              |  +-- Pseudodon_vondembuschianus **
              +--+                             
                 +-- Pseudodon_mekongi **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 32 14 6 4 
  +-------------------- Pilsbryoconcha            
  |                                               
  |                 +-- Nyeinchanconcha           
  |              +--+                             
--+              |  +-- Sundadontina_cumingii     
  |  +-----------+                                
  |  |           |  +-- Thaiconcha                
  |  |           +--+                             
  |  |              +-- Bineurus                  
  +--+                                            
     |           +----- Sundadontina_tumida       
     |  +--------+                                
     |  |        |  +-- Sundadontina_brandti      
     |  |        +--+                             
     +--+           +-- Sundadontina_taskaevi     
        |                                         
        |  +----------- Namkongnaia               
        +--+                                      
           |  +-------- Pseudodon_lenyanensis **     
           +--+                                   
              |  +----- Pseudodon_cambodjensis **    
              +--+                                
                 |  +-- Pseudodon_vondembuschianus **
                 +--+                             
                    +-- Pseudodon_mekongi **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
4 4 2 4 79 22 7 4 
  +-------------------- Pilsbryoconcha            
  |                                               
--+  +----------------- Songkhlanaia              
  |  |                                            
  +--+  +-------------- Namkongnaia               
     |  |                                         
     |  |     +-------- Pseudodon_lenyanensis **     
     +--+  +--+                                   
        |  |  |  +----- Pseudodon_cambodjensis **    
        |  |  +--+                                
        |  |     |  +-- Pseudodon_vondembuschianus **
        +--+     +--+                             
           |        +-- Pseudodon_mekongi **         
           |                                      
           |  +-------- Sundadontina              
           +--+                                   
              |  +----- Nyeinchanconcha           
              +--+                                
                 |  +-- Thaiconcha                
                 +--+                             
                    +-- Bineurus

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
4 7 5 4 135 9 5 7 
     +----- Pseudodon_cambodjensis **    
  +--+                                
  |  |  +-- Pseudodon_mekongi **         
  |  +--+                             
  |     +-- Pseudodon_vondembuschianus **
--+                                   
  |     +-- Pseudodon_inoscularis **     
  |  +--+                             
  |  |  +-- Sundadontina_tumida       
  +--+                                
     |  +-- Nyeinchanconcha           
     +--+                             
        +-- Sundadontina_cumingii

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 4 31 13 6 4 
                 +----- Pilsbryoconcha_exilis       
  +--------------+                                  
  |              |  +-- Pilsbryoconcha_linguaeformis
  |              +--+                               
  |                 +-- Namkongnaia_lemeslei__2     
--+                                                 
  |              +----- Nyeinchanconcha             
  |  +-----------+                                  
  |  |           |  +-- Thaiconcha                  
  |  |           +--+                               
  +--+              +-- Bineurus                    
     |                                              
     |  +-------------- Sundadontina                
     |  |                                           
     +--+           +-- Namkongnaia_lemeslei__1     
        |  +--------+                               
        |  |        +-- Namkongnaia_inkhavilayi     
        +--+                                        
           |  +-------- Pseudodon_lenyanensis **       
           +--+                                     
              |  +----- Pseudodon_cambodjensis **      
              +--+                                  
                 |  +-- Pseudodon_vondembuschianus **  
                 +--+                               
                    +-- Pseudodon_mekongi **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 4 114 13 5 4 
  +-------------------- Bineurus                  
  |                                               
--+  +----------------- Nyeinchanconcha           
  |  |                                            
  +--+              +-- Thaiconcha                
     |  +-----------+                             
     |  |           +-- Pilsbryoconcha            
     +--+                                         
        |  +----------- Sundadontina              
        +--+                                      
           |  +-------- Pseudodon_lenyanensis **     
           +--+                                   
              |  +----- Pseudodon_cambodjensis **    
              +--+                                
                 |  +-- Pseudodon_vondembuschianus **
                 +--+                             
                    +-- Pseudodon_mekongi **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 122 9 5 5 
  +----------- Bineurus                  
  |                                      
  |        +-- Nyeinchanconcha           
  +--------+                             
  |        +-- Sundadontina_cumingii     
--+                                      
  |        +-- Thaiconcha                
  +--------+                             
  |        +-- Pilsbryoconcha            
  |                                      
  |  +-------- Sundadontina_tumida       
  +--+                                   
     |  +----- Pseudodon_cambodjensis **    
     +--+                                
        |  +-- Pseudodon_mekongi **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 45 9 5 5 
  +-------------- Pilsbryoconcha            
  |                                         
  |        +----- Pseudodon_cambodjensis **    
--+  +-----+                                
  |  |     |  +-- Pseudodon_mekongi **         
  |  |     +--+                             
  +--+        +-- Pseudodon_vondembuschianus **
     |                                      
     |  +-------- Sundadontina              
     +--+                                   
        |  +----- Nyeinchanconcha           
        +--+                                
           |  +-- Bineurus                  
           +--+                             
              +-- Thaiconcha

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 44 9 5 5 
  +-------------- Pilsbryoconcha            
  |                                         
  |        +----- Pseudodon_cambodjensis **    
--+  +-----+                                
  |  |     |  +-- Pseudodon_mekongi **         
  |  |     +--+                             
  +--+        +-- Pseudodon_vondembuschianus **
     |                                      
     |  +-------- Sundadontina              
     +--+                                   
        |  +----- Nyeinchanconcha           
        +--+                                
           |  +-- Bineurus                  
           +--+                             
              +-- Thaiconcha

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3 3 2 4 196 9 5 6 
  +----------------- Pilsbryoconcha               
  |                                               
  |              +-- Nyeinchanconcha              
  |           +--+                                
--+           |  +-- Sundadontina_cumingii        
  |  +--------+                                   
  |  |        |  +-- Thaiconcha                   
  |  |        +--+                                
  +--+           +-- Bineurus                     
     |                                            
     |  +----------- Sundadontina_tumida          
     +--+                                         
        |  +-------- Pseudodon_cambodjensis **       
        +--+                                      
           |  +----- Pseudodon_vondembuschianus__1 **
           +--+                                   
              |  +-- Pseudodon_mekongi **            
              +--+                                
                 +-- Pseudodon_vondembuschianus__2 **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 198 9 5 5 
  +-------------- Pilsbryoconcha            
  |                                         
  |        +----- Nyeinchanconcha           
--+  +-----+                                
  |  |     |  +-- Thaiconcha                
  |  |     +--+                             
  +--+        +-- Bineurus                  
     |                                      
     |  +-------- Sundadontina              
     +--+                                   
        |  +----- Pseudodon_cambodjensis **    
        +--+                                
           |  +-- Pseudodon_mekongi **         
           +--+                             
              +-- Pseudodon_vondembuschianus **

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 46 9 5 5 
  +-------------- Pilsbryoconcha            
  |                                         
  |        +----- Pseudodon_cambodjensis **    
--+  +-----+                                
  |  |     |  +-- Pseudodon_mekongi **         
  |  |     +--+                             
  +--+        +-- Pseudodon_vondembuschianus **
     |                                      
     |  +-------- Sundadontina              
     +--+                                   
        |  +----- Nyeinchanconcha           
        +--+                                
           |  +-- Bineurus                  
           +--+                             
              +-- Thaiconcha

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
3 3 1 3 42 11 5 5 
              +-- Nyeinchanconcha       
  +-----------+                         
  |           +-- Bineurus              
  |                                     
--+           +-- Thaiconcha            
  |  +--------+                         
  |  |        +-- Sundadontina          
  +--+                                  
     |  +-------- Pilsbryoconcha        
     +--+                               
        |  +----- Pseudodon_lenyanensis ** 
        +--+                            
           |  +-- Pseudodon_mekongi **     
           +--+                         
              +-- Pseudodon_cambodjensis **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
3 8 0 3 80 5 2 11 
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_cambodjensis **    
  |                             
  +-- Pseudodon_mekongi **         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_vondembuschianus **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
3 3 1 3 42 11 5 5 
           +-- Nyeinchanconcha       
  +--------+                         
  |        +-- Bineurus              
  |                                  
  |        +-- Thaiconcha            
--+--------+                         
  |        +-- Sundadontina          
  |                                  
  |  +-------- Pilsbryoconcha        
  +--+                               
     |  +----- Pseudodon_lenyanensis ** 
     +--+                            
        |  +-- Pseudodon_mekongi **     
        +--+                         
           +-- Pseudodon_cambodjensis **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 43 5 4 5 
        +----- Sundadontina              
  +-----+                                
  |     |  +-- Thaiconcha                
  |     +--+                             
--+        +-- Bineurus                  
  |                                      
  |  +-------- Pilsbryoconcha            
  +--+                                   
     |  +----- Pseudodon_walpolei **        
     +--+                                
        |  +-- Pseudodon_mekongi **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3 3 1 3 198 9 5 5 
  +-------- Nyeinchanconcha           
  |                                   
  +-------- Thaiconcha                
  |                                   
  +-------- Sundadontina_cumingii     
  |                                   
  +-------- Sundadontina_tumida__1    
--+                                   
  +-------- Sundadontina_tumida__2    
  |                                   
  +-------- Pilsbryoconcha            
  |                                   
  +-------- Bineurus                  
  |                                   
  |  +----- Pseudodon_cambodjensis **    
  +--+                                
     |  +-- Pseudodon_mekongi **         
     +--+                             
        +-- Pseudodon_vondembuschianus **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 136 9 5 5 
  +-------------- Pilsbryoconcha            
  |                                         
  |     +-------- Sundadontina_tumida       
  |  +--+                                   
--+  |  |  +----- Pseudodon_cambodjensis **    
  |  |  +--+                                
  |  |     |  +-- Pseudodon_mekongi **         
  |  |     +--+                             
  +--+        +-- Pseudodon_vondembuschianus **
     |                                      
     |        +-- Nyeinchanconcha           
     |     +--+                             
     |     |  +-- Sundadontina_cumingii     
     +-----+                                
           |  +-- Thaiconcha                
           +--+                             
              +-- Bineurus

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
3 8 0 3 49 5 2 11 
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_cambodjensis **    
  |                             
  +-- Pseudodon_mekongi **         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_vondembuschianus **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +-------- Pilsbryoconcha            
  |                                   
--+     +-- Pseudodon_mekongi **         
  |  +--+                             
  |  |  +-- Pseudodon_vondembuschianus **
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +-------- Pilsbryoconcha            
  |                                   
--+     +-- Pseudodon_mekongi **         
  |  +--+                             
  |  |  +-- Pseudodon_vondembuschianus **
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha            
--+                                      
  |  +-------- Bineurus                  
  +--+                                   
     |  +----- Thaiconcha                
     +--+                                
        |  +-- Pseudodon_mekongi **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha            
  |                                      
--+  +-------- Bineurus_anodontinum      
  |  |                                   
  +--+-------- Bineurus_loeiensis        
     |                                   
     |  +----- Thaiconcha                
     +--+                                
        |  +-- Pseudodon_mekongi **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 29 4 4 -- 
  +----------- Sundadontina              
  |                                      
--+        +-- Nyeinchanconcha           
  |  +-----+                             
  |  |     +-- Bineurus                  
  +--+                                   
     |  +----- Pilsbryoconcha            
     +--+                                
        |  +-- Pseudodon_cambodjensis **    
        +--+                             
           +-- Pseudodon_vondembuschianus **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 30 5 4 -- 
  +----------- Pilsbryoconcha            
  |                                      
--+        +-- Pseudodon_mekongi **         
  |  +-----+                             
  |  |     +-- Pseudodon_vondembuschianus **
  +--+                                   
     |  +----- Sundadontina              
     +--+                                
        |  +-- Thaiconcha                
        +--+                             
           +-- Bineurus

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +-------- Pilsbryoconcha            
  |                                   
--+     +-- Pseudodon_mekongi **         
  |  +--+                             
  |  |  +-- Pseudodon_vondembuschianus **
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
2 2 0 2 35 4 3 -- 
  +-------- Pilsbryoconcha            
  |                                   
--+     +-- Pseudodon_mekongi **         
  |  +--+                             
  |  |  +-- Pseudodon_vondembuschianus **
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 38 4 4 -- 
  +----------- Pilsbryoconcha            
  |                                      
--+        +-- Pseudodon_vondembuschianus **
  |  +-----+                             
  |  |     +-- Pseudodon_cambodjensis **    
  +--+                                   
     |  +----- Sundadontina              
     +--+                                
        |  +-- Nyeinchanconcha           
        +--+                             
           +-- Bineurus

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 10 6 -- 
  +----------- Bineurus                  
  |                                      
  |     +----- Namkongnaia               
--+  +--+                                
  |  |  |  +-- Sundadontina              
  |  |  +--+                             
  +--+     +-- Nyeinchanconcha           
     |                                   
     |  +----- Pseudodon_vondembuschianus **
     +--+                                
        |  +-- Pilsbryoconcha            
        +--+                             
           +-- Thaiconcha

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 45 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 43 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus **

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 18 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 23 1 1 -- 
  +-- Pseudodon_cambodjensis **
--+                         
  +-- Pilsbryoconcha

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 2 1 -- 
  +----- Pilsbryoconcha_exilis    
--+                               
  |  +-- Pseudodon_cambodjensis **   
  +--+                            
     +-- Pilsbryoconcha_carinifera

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Pseudodon_vondembuschianus **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 14 6 -- 
     +----- Pilsbryoconcha            
  +--+                                
  |  |  +-- Namkongnaia               
  |  +--+                             
  |     +-- Pseudodon_vondembuschianus **
--+                                   
  |     +-- Nyeinchanconcha           
  |  +--+                             
  |  |  +-- Sundadontina              
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 2 1 -- 
  +----- Pilsbryoconcha_exilis    
--+                               
  |  +-- Pseudodon_cambodjensis **   
  +--+                            
     +-- Pilsbryoconcha_carinifera

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 57 2 2 -- 
-- Pseudodon_lenyanensis **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 3 3 -- 
  +-------- Pilsbryoconcha            
--+                                   
  |  +----- Bineurus                  
  +--+                                
     |  +-- Pseudodon_vondembuschianus **
     +--+                             
        +-- Sundadontina

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 45 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Pseudodon_vondembuschianus **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 53 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 60 2 2 -- 
-- Pseudodon_lenyanensis **

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 57 2 2 -- 
-- Pseudodon_lenyanensis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 26 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 79 4 3 -- 
  +-------- Pilsbryoconcha        
--+                               
  |  +----- Pseudodon_cambodjensis **
  +--+                            
     |  +-- Bineurus              
     +--+                         
        +-- Nyeinchanconcha

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 14 6 -- 
     +----- Pilsbryoconcha            
  +--+                                
  |  |  +-- Namkongnaia               
  |  +--+                             
  |     +-- Pseudodon_vondembuschianus **
--+                                   
  |     +-- Nyeinchanconcha           
  |  +--+                             
  |  |  +-- Sundadontina              
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 3 3 -- 
  +-------- Bineurus                  
--+                                   
  |  +----- Pilsbryoconcha            
  +--+                                
     |  +-- Pseudodon_vondembuschianus **
     +--+                             
        +-- Sundadontina

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
1 1 0 1 48 1 1 -- 
  +-- Pseudodon_lenyanensis **
--+                        
  +-- Sundadontina

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 43 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 18 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 60 2 2 -- 
-- Pseudodon_lenyanensis **

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 57 2 2 -- 
-- Pseudodon_lenyanensis **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 5 4 -- 
  +-------- Pilsbryoconcha   
  |                          
--+  +----- Pseudodon_mekongi **
  |  |                       
  +--+  +-- Nyeinchanconcha  
     |  |                    
     +--+-- Bineurus         
        |                    
        +-- Sundadontina

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 1 1 -- 
  +-- Pseudodon_cambodjensis **
--+                         
  +-- Sundadontina

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha