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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini | Cyprogenia

species Cyprogenia stegaria (Rafinesque, 1820): synonymy
     3 specimens + 3 Genbank vouchers

Obovaria stegaria Rafinesque, 1820
Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 312, pl. 82, figs. 4-5 [also as ‘Unio stegaria’].
Rafinesque, 1864. [in] Binney & Tryon, reprint: 55.

Obovaria stegariaSimpson (1900), Simpson (1914)
Cyprogenia stegariaVanatta (1915), Ortmann (1918), Frierson (1927), Haas (1930), Thiele (1934), ... Klymus et al. (2020), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023), FMCS (2025)
Cyprogenia irroratusOrtmann & Walker (1922)
Cyprogenia irrorataBurch (1975)

Obovaria stegaria var. fasciolata Rafinesque, 1820
Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 312.
Rafinesque, 1864. [in] Binney & Tryon, reprint: 55.
(intro'd as intra-sp. of Obovaria stegaria Rafinesque, 1820)

Vanatta (1915)

Obovaria stegaria var. tuberculata Rafinesque, 1820
Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 312.
Rafinesque, 1864. [in] Binney & Tryon, reprint: 55.
(intro'd as intra-sp. of Obovaria stegaria Rafinesque, 1820; homonym of Obovaria striata var. tuberculata Rafinesque, 1820)
Type(s): LECTOTYPE ANSP_20241, Ohio River.

Cyprogenia stegariaVanatta (1915), Williams et al. (2008), Watters et al. (2009)

Unio irroratus Lea, 1828
Lea, 1828. Trans. Amer. Phil. Soc. 3 [O. 1]: 269, pl. 5, fig. 5-7.
Type(s): PARATYPE USNM_84299, Cincinnati, Ohio.

Cyprogenia irrorataSimpson (1900), Ortmann (1909), Ortmann (1909), Ortmann (1912), Simpson (1914), Ortmann (1924), Ortmann (1925), Burch (1975)
Unio (s.s.) irroratusCall (1900)
Cryptogenia [sic] irrorataOrtmann (1909)
Unio irroratusde Gregorio (1914)
Cyprogenia stegariaVanatta (1915), Ortmann (1918), Frierson (1927), Haas (1930), Thiele (1934), ... Parmalee & Bogan (1998), Graf & Cummings (2007), Williams et al. (2008), Watters et al. (2009), Graf & Cummings (2021)
Cyprogenia irroratusOrtmann & Walker (1922)
Cyprogenia irrorata irrorataITIS (2012)

Unio verrucosus albus Hildreth, 1828
Hildreth, 1828. Amer. J. Sci. 14: 281.
(intro'd as intra-sp. of Unio verrucosus Barnes, 1823)

Cyprogenia irrorataSimpson (1900), Simpson (1914)
Cyprogenia stegariaFrierson (1927), Haas (1969), R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)

Unio stegarius Say, 1834
‘Rafinesque’ of Authors.
Say, 1834. Amer. Conch. 6: ??.
Ferussac, 1835. Mag. Zool. 5: 28.
Conrad, 1838. Monogr. Fam. Unionidae 1(10): 83, pl. 46, fig. 1.
Reeve, 1864. Conch. Icon. 16: pl. 12, sp. 45.
(reference to Obovaria stegaria Rafinesque, 1820)

Cyprogenia irrorataSimpson (1900), Simpson (1914)
Cyprogenia stegariaR.I. Johnson (1980), Watters et al. (2009)

Cyprogenia irrorata var. pusilla Simpson, 1900
Simpson, 1900. Proc. U.S. Nat. Mus. 22: 610.
Simpson, 1914. Descr. Cat. Naiades: 328.
(intro'd as intra-sp. of Unio irroratus Lea, 1828)
Type(s): LECTOTYPE USNM_152004 (by R.I. Johnson, 1975: 18), Mammoth Cave (Green R.), Kentucky.

Cyprogenia irrorata var. pusillaSimpson (1900), Simpson (1914)
Cyprogenia stegaria-pusillaFrierson (1927)
Cyprogenia stegariaHaas (1969), R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Cyprogenia irrorata pusillaITIS (2012)

Taxonomic History (since 1900)

Obovaria stegariaSimpson (1900), Simpson (1914)
Cyprogenia irrorataSimpson (1900), Ortmann (1909), Ortmann (1909), Ortmann (1912), Simpson (1914), Ortmann (1924), Ortmann (1925), Burch (1975)
Unio (s.s.) irroratusCall (1900)
Cyprogenia irrorata var. pusillaSimpson (1900), Simpson (1914)
Cryptogenia [sic] irrorataOrtmann (1909)
Unio irroratusde Gregorio (1914)
Cyprogenia stegariaVanatta (1915), Ortmann (1918), Frierson (1927), Haas (1930), Thiele (1934), ... Klymus et al. (2020), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023), FMCS (2025)
Vanatta (1915)
Cyprogenia irroratusOrtmann & Walker (1922)
Cyprogenia stegaria-pusillaFrierson (1927)
Cyprogenia irrorata irrorataITIS (2012)
Cyprogenia irrorata pusillaITIS (2012)

 
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