Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Oxynaiina

genus Pseudocuneopsis Huang, Dai, Chen & Wu et al., 2022: cladomics

7 Recent valid species (7 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 7 (7 total)
analyzed in 43 cladograms in 26 publications
maximum number of terminals in an analysis = 7 (mean = 2.51, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 29
maximum number of included terminals = 7, (mean = 3.24 median = 2)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pseudocuneopsis cladogram distances
PCoA Ordination

NJ Tree of Pseudocuneopsis cladogram distances
NJ Tree

Cladograms

Wang et al., 2025 [Publ 4488], fig. 3B

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
7 7 5 7 58 16 4 0 
           +-------- Nodularia                    
  +--------+                                      
  |        |  +----- Schistodesmus                
  |        +--+                                   
  |           |  +-- Tchangsinaia                 
--+           +--+                                
  |              +-- Cuneopsis                    
  |                                               
  |  +-------------- Pseudocuneopsis_capitata **     
  |  |                                            
  +--+  +----------- Pseudocuneopsis_perflora **     
     |  |                                         
     +--+  +-------- Pseudocuneopsis_liuovatus **    
        |  |                                      
        +--+     +-- Pseudocuneopsis_sichuanensis ** 
           |  +--+                                
           |  |  +-- Pseudocuneopsis_yemaoi **       
           +--+                                   
              |  +-- Pseudocuneopsis_wuana **        
              +--+                                
                 +-- Pseudocuneopsis_yangshuoensis **

Chen et al., 2023 [Publ 4302], fig. 1

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
2 2 0 2 39 12 6 -- 
  +-- Pseudocuneopsis_sichuanensis **
--+                               
  +-- Pseudocuneopsis_capitata **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
6 6 4 6 80 13 5 1 
        +-------- Nodularia                    
        |                                      
  +-----+     +-- Tongchuania                  
  |     |  +--+                                
  |     |  |  +-- Schistodesmus                
  |     +--+                                   
  |        |  +-- Tchangsinaia                 
--+        +--+                                
  |           +-- Cuneopsis                    
  |                                            
  |  +----------- Pseudocuneopsis_capitata **     
  |  |                                         
  +--+  +-------- Pseudocuneopsis_perflora **     
     |  |                                      
     +--+     +-- Pseudocuneopsis_sichuanensis ** 
        |  +--+                                
        |  |  +-- Pseudocuneopsis_yemaoi **       
        +--+                                   
           |  +-- Pseudocuneopsis_wuana **        
           +--+                                
              +-- Pseudocuneopsis_yangshuoensis **

Chen et al., 2025 [Publ 4448], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 6 3 2 1 3 
  +-------------- Nodularia                    
  |                                            
--+  +----------- Pseudocuneopsis_perflora **     
  |  |                                         
  +--+  +-------- Pseudocuneopsis_capitata **     
     |  |                                      
     +--+     +-- Pseudocuneopsis_yangshuoensis **
        |  +--+                                
        |  |  +-- Pseudocuneopsis_wuana **        
        +--+                                   
           |  +-- Pseudocuneopsis_yemaoi **       
           +--+                                
              +-- Pseudocuneopsis_sichuanensis **

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 10 7 2 2 
  +-------- Pseudocuneopsis_capitata **     
  |                                      
--+     +-- Pseudocuneopsis_sichuanensis ** 
  |  +--+                                
  |  |  +-- Pseudocuneopsis_yemaoi **       
  +--+                                   
     |  +-- Pseudocuneopsis_wuana **        
     +--+                                
        +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2025 [Publ 4432], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 23 20 4 2 
     +-------- Nodularia                    
  +--+                                      
  |  |  +----- Schistodesmus                
  |  +--+                                   
  |     |  +-- Tchangsinaia                 
--+     +--+                                
  |        +-- Cuneopsis                    
  |                                         
  |  +-------- Pseudocuneopsis_capitata **     
  |  |                                      
  +--+     +-- Pseudocuneopsis_sichuanensis ** 
     |  +--+                                
     |  |  +-- Pseudocuneopsis_yemaoi **       
     +--+                                   
        |  +-- Pseudocuneopsis_wuana **        
        +--+                                
           +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2025 [Publ 4401], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 23 20 4 2 
     +-------- Nodularia                    
  +--+                                      
  |  |  +----- Schistodesmus                
  |  +--+                                   
  |     |  +-- Tchangsinaia                 
--+     +--+                                
  |        +-- Cuneopsis                    
  |                                         
  |  +-------- Pseudocuneopsis_capitata **     
  |  |                                      
  +--+     +-- Pseudocuneopsis_sichuanensis ** 
     |  +--+                                
     |  |  +-- Pseudocuneopsis_yemaoi **       
     +--+                                   
        |  +-- Pseudocuneopsis_wuana **        
        +--+                                
           +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 71 19 4 2 
     +-------- Nodularia                    
  +--+                                      
  |  |  +----- Schistodesmus                
  |  +--+                                   
  |     |  +-- Tchangsinaia                 
--+     +--+                                
  |        +-- Cuneopsis                    
  |                                         
  |  +-------- Pseudocuneopsis_capitata **     
  |  |                                      
  +--+     +-- Pseudocuneopsis_sichuanensis ** 
     |  +--+                                
     |  |  +-- Pseudocuneopsis_yemaoi **       
     +--+                                   
        |  +-- Pseudocuneopsis_wuana **        
        +--+                                
           +-- Pseudocuneopsis_yangshuoensis **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 84 12 4 3 
           +----- Schistodesmus                
  +--------+                                   
  |        |  +-- Tchangsinaia                 
  |        +--+                                
--+           +-- Cuneopsis                    
  |                                            
  |  +----------- Nodularia                    
  +--+                                         
     |  +-------- Pseudocuneopsis_capitata **     
     +--+                                      
        |  +----- Pseudocuneopsis_yangshuoensis **
        +--+                                   
           |  +-- Pseudocuneopsis_sichuanensis ** 
           +--+                                
              +-- Pseudocuneopsis_yemaoi **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 4 20 7 5 3 
  +-------------- Cuneopsis                    
  |                                            
  |           +-- Nodularia                    
  +-----------+                                
--+           +-- Schistodesmus                
  |                                            
  |  +----------- Tchangsinaia                 
  |  |                                         
  +--+----------- Inversiunio                  
     |                                         
     |  +-------- Pseudocuneopsis_capitata **     
     +--+                                      
        |  +----- Pseudocuneopsis_sichuanensis ** 
        +--+                                   
           |  +-- Pseudocuneopsis_wuana **        
           +--+                                
              +-- Pseudocuneopsis_yangshuoensis **

Dai et al., 2024 [Publ 4358], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 2 2 2 3 
           +-- Schistodesmus                
  +--------+                                
  |        +-- Nodularia                    
--+                                         
  |  +-------- Pseudocuneopsis_capitata **     
  +--+                                      
     |  +----- Pseudocuneopsis_yangshuoensis **
     +--+                                   
        |  +-- Pseudocuneopsis_sichuanensis ** 
        +--+                                
           +-- Pseudocuneopsis_yemaoi **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 4 20 7 5 3 
  +-------------------- Cuneopsis                    
  |                                                  
--+                 +-- Nodularia                    
  |  +--------------+                                
  |  |              +-- Schistodesmus                
  +--+                                               
     |  +-------------- Tchangsinaia                 
     +--+                                            
        |  +----------- Inversiunio                  
        +--+                                         
           |  +-------- Pseudocuneopsis_capitata **     
           +--+                                      
              |  +----- Pseudocuneopsis_sichuanensis ** 
              +--+                                   
                 |  +-- Pseudocuneopsis_wuana **        
                 +--+                                
                    +-- Pseudocuneopsis_yangshuoensis **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 4 49 10 4 3 
     +-------- Pseudocuneopsis_capitata **     
  +--+                                      
  |  |  +----- Pseudocuneopsis_yangshuoensis **
  |  +--+                                   
  |     |  +-- Pseudocuneopsis_yemaoi **       
--+     +--+                                
  |        +-- Pseudocuneopsis_sichuanensis ** 
  |                                         
  |  +-------- Nodularia                    
  +--+                                      
     |  +----- Schistodesmus                
     +--+                                   
        |  +-- Tchangsinaia                 
        +--+                                
           +-- Cuneopsis

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 20 7 5 4 
  +----------------- Cuneopsis                    
  |                                               
--+              +-- Nodularia                    
  |  +-----------+                                
  |  |           +-- Schistodesmus                
  +--+                                            
     |  +----------- Tchangsinaia                 
     +--+                                         
        |  +-------- Inversiunio                  
        +--+                                      
           |  +----- Pseudocuneopsis_capitata **     
           +--+                                   
              |  +-- Pseudocuneopsis_yangshuoensis **
              +--+                                
                 +-- Pseudocuneopsis_sichuanensis **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 12 6 -- 
  +-- Pseudocuneopsis_sichuanensis **
--+                               
  +-- Pseudocuneopsis_capitata **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 51 20 4 -- 
           +-- Pseudocuneopsis_sichuanensis **
  +--------+                               
  |        +-- Pseudocuneopsis_capitata **    
--+                                        
  |  +-------- Nodularia                   
  +--+                                     
     |  +----- Schistodesmus               
     +--+                                  
        |  +-- Tchangsinaia                
        +--+                               
           +-- Cuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
2 2 0 2 94 10 4 -- 
              +-- Pseudocuneopsis_capitata **    
  +-----------+                               
  |           +-- Pseudocuneopsis_sichuanensis **
--+                                           
  |  +----------- Nodularia                   
  |  |                                        
  +--+        +-- Schistodesmus_lampreyanus   
     |  +-----+                               
     |  |     +-- Cuneopsis                   
     +--+                                     
        |  +----- Tchangsinaia                
        +--+                                  
           |  +-- Schistodesmus_xinyuensis    
           +--+                               
              +-- Schistodesmus_spinosus

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 52 17 4 -- 
           +-- Pseudocuneopsis_capitata **    
  +--------+                               
  |        +-- Pseudocuneopsis_sichuanensis **
--+                                        
  |  +-------- Nodularia                   
  +--+                                     
     |  +----- Schistodesmus               
     +--+                                  
        |  +-- Tchangsinaia                
        +--+                               
           +-- Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 72 10 4 -- 
           +-- Pseudocuneopsis_capitata **    
  +--------+                               
  |        +-- Pseudocuneopsis_sichuanensis **
--+                                        
  |  +-------- Nodularia                   
  +--+                                     
     |  +----- Schistodesmus               
     +--+                                  
        |  +-- Tchangsinaia                
        +--+                               
           +-- Cuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
2 2 0 2 94 10 4 -- 
     +----- Nodularia                   
  +--+                                  
  |  |  +-- Pseudocuneopsis_capitata **    
  |  +--+                               
--+     +-- Pseudocuneopsis_sichuanensis **
  |                                     
  |  +----- Tchangsinaia                
  +--+                                  
     |  +-- Schistodesmus               
     +--+                               
        +-- Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
2 2 0 2 16 7 4 -- 
           +-- Pseudocuneopsis_sichuanensis **
  +--------+                               
  |        +-- Pseudocuneopsis_capitata **    
--+                                        
  |  +-------- Nodularia                   
  +--+                                     
     |  +----- Tchangsinaia                
     +--+                                  
        |  +-- Cuneopsis                   
        +--+                               
           +-- Schistodesmus

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
2 2 0 2 90 10 4 -- 
  +-------- Nodularia                   
  |                                     
  |     +-- Pseudocuneopsis_capitata **    
--+-----+                               
  |     +-- Pseudocuneopsis_sichuanensis **
  |                                     
  |  +----- Schistodesmus               
  +--+                                  
     |  +-- Tchangsinaia                
     +--+                               
        +-- Cuneopsis

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 43 11 6 -- 
  +-- Pseudocuneopsis_sichuanensis **
--+                               
  +-- Pseudocuneopsis_capitata **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
2 2 0 2 90 10 4 -- 
  +-------- Nodularia                   
  |                                     
  |     +-- Pseudocuneopsis_capitata **    
--+-----+                               
  |     +-- Pseudocuneopsis_sichuanensis **
  |                                     
  |  +----- Schistodesmus               
  +--+                                  
     |  +-- Tchangsinaia                
     +--+                               
        +-- Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
2 2 0 2 10 9 5 -- 
  +-------------- Nodularia                   
  |                                           
--+           +-- Pseudocuneopsis_capitata **    
  |  +--------+                               
  |  |        +-- Pseudocuneopsis_sichuanensis **
  +--+                                        
     |  +-------- Schistodesmus               
     +--+                                     
        |  +----- Cuneopsis                   
        +--+                                  
           |  +-- Tchangsinaia                
           +--+                               
              +-- Inversiunio

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 43 11 6 -- 
  +-- Pseudocuneopsis_sichuanensis **
--+                               
  +-- Pseudocuneopsis_capitata **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
2 2 0 2 37 13 5 -- 
              +-- Pseudocuneopsis_capitata **    
  +-----------+                               
  |           +-- Pseudocuneopsis_sichuanensis **
--+                                           
  |  +----------- Nodularia                   
  +--+                                        
     |  +-------- Schistodesmus               
     +--+                                     
        |  +----- Cuneopsis                   
        +--+                                  
           |  +-- Tchangsinaia                
           +--+                               
              +-- Inversiunio

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 39 11 6 -- 
              +-- Tchangsinaia                
  +-----------+                               
  |           +-- Inversiunio                 
  |                                           
--+           +-- Pseudocuneopsis_sichuanensis **
  |  +--------+                               
  |  |        +-- Pseudocuneopsis_capitata **    
  +--+                                        
     |  +-------- Cuneopsis                   
     +--+                                     
        |  +----- Nodularia                   
        +--+                                  
           |  +-- Diaurora                    
           +--+                               
              +-- Schistodesmus

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 11 6 -- 
           +-- Tchangsinaia                
  +--------+                               
  |        +-- Inversiunio                 
  |                                        
--+     +----- Cuneopsis                   
  |  +--+                                  
  |  |  |  +-- Pseudocuneopsis_sichuanensis **
  |  |  +--+                               
  +--+     +-- Pseudocuneopsis_capitata **    
     |                                     
     |  +----- Nodularia                   
     +--+                                  
        |  +-- Diaurora                    
        +--+                               
           +-- Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 6 4 -- 
  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Schistodesmus           
     +--+                              
        |  +-- Cuneopsis               
        +--+                           
           +-- Tchangsinaia

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 4 4 -- 
  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Tchangsinaia            
     +--+                              
        |  +-- Cuneopsis               
        +--+                           
           +-- Schistodesmus

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 10 4 -- 
  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Schistodesmus           
     +--+                              
        |  +-- Tchangsinaia            
        +--+                           
           +-- Cuneopsis

Wu et al., 2024 [Publ 4379], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 12 12 3 -- 
  +----- Nodularia               
  |                              
--+  +-- Cuneopsis               
  |  |                           
  +--+-- Tchangsinaia            
     |                           
     +-- Pseudocuneopsis_capitata **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 6 4 -- 
  +----------- Pseudocuneopsis_sichuanensis **
--+                                        
  |  +-------- Nodularia                   
  +--+                                     
     |  +----- Tchangsinaia                
     +--+                                  
        |  +-- Cuneopsis                   
        +--+                               
           +-- Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 6 4 -- 
  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Schistodesmus           
     +--+                              
        |  +-- Tchangsinaia            
        +--+                           
           +-- Cuneopsis

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 4 4 -- 
  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Tchangsinaia            
     +--+                              
        |  +-- Cuneopsis               
        +--+                           
           +-- Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 6 4 -- 
  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Tchangsinaia            
     +--+                              
        |  +-- Cuneopsis               
        +--+                           
           +-- Schistodesmus

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 53 5 4 -- 
        +-- Cuneopsis               
  +-----+                           
  |     +-- Schistodesmus           
--+                                 
  |  +----- Pseudocuneopsis_capitata **
  +--+                              
     |  +-- Nodularia               
     +--+                           
        +-- Tchangsinaia

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 13 6 4 -- 
  +-------------- Tchangsinaia            
--+                                       
  |  +----------- Cuneopsis_heudei        
  +--+                                    
     |  +-------- Nodularia               
     +--+                                 
        |  +----- Pseudocuneopsis_capitata **
        +--+                              
           |  +-- Cuneopsis_rufescens     
           +--+                           
              +-- Schistodesmus

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 7 4 -- 
  +-------- Pseudocuneopsis_capitata **
  |                                 
--+     +-- Cuneopsis               
  |  +--+                           
  |  |  +-- Schistodesmus           
  +--+                              
     |  +-- Tchangsinaia            
     +--+                           
        +-- Nodularia

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 29 5 4 -- 
  +----------- Pseudocuneopsis_capitata **
--+                                    
  |  +-------- Nodularia               
  +--+                                 
     |  +----- Schistodesmus           
     +--+                              
        |  +-- Cuneopsis               
        +--+                           
           +-- Tchangsinaia

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 4 4 -- 
        +-- Pseudocuneopsis_capitata **
  +-----+                           
  |     +-- Nodularia               
--+                                 
  |  +----- Schistodesmus           
  +--+                              
     |  +-- Cuneopsis               
     +--+                           
        +-- Tchangsinaia

Hou et al., 2025 [Publ 4432], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 10 8 2 -- 
-- Pseudocuneopsis_capitata **