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Data last updated
Thu 18 Dec 2025

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Pseudocuneopsis Huang, Dai, Chen & Wu et al., 2022: cladomics

7 Recent valid species (7 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 7 (7 total)
analyzed in 43 cladograms in 26 publications
maximum number of terminals in an analysis = 7 (mean = 2.51, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 29
maximum number of included terminals = 7, (mean = 3.24 median = 2)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Pseudocuneopsis cladogram distances
PCoA Ordination

NJ Tree of Pseudocuneopsis cladogram distances
NJ Tree

Cladograms

Wang et al., 2025 [Publ 4488], fig. 3B

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
7 7 5 7 58 26 7 0 
  +-------------------------- Aculamprotula                
  |                                                        
--+  +----------------------- Unio                         
  |  |                                                     
  +--+  +-------------------- Middendorffinaia             
     |  |                                                  
     |  |           +-------- Nodularia                    
     +--+  +--------+                                      
        |  |        |  +----- Schistodesmus                
        |  |        +--+                                   
        |  |           |  +-- Tchangsinaia                 
        +--+           +--+                                
           |              +-- Cuneopsis                    
           |                                               
           |  +-------------- Pseudocuneopsis_capitata **     
           |  |                                            
           +--+  +----------- Pseudocuneopsis_perflora **     
              |  |                                         
              +--+  +-------- Pseudocuneopsis_liuovatus **    
                 |  |                                      
                 +--+     +-- Pseudocuneopsis_sichuanensis ** 
                    |  +--+                                
                    |  |  +-- Pseudocuneopsis_yemaoi **       
                    +--+                                   
                       |  +-- Pseudocuneopsis_wuana **        
                       +--+                                
                          +-- Pseudocuneopsis_yangshuoensis **

Dai et al., 2025 [Publ 4440], fig. 1

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
2 2 0 2 43 20 10 -- 
  +----------------------- Globunio                    
  |                                                    
--+  +-------------------- Aculamprotula               
  |  |                                                 
  +--+  +----------------- Middendorffinaia            
     |  |                                              
     +--+  +-------------- Nodularia                   
        |  |                                           
        |  |  +----------- Cuneopsis                   
        +--+  |                                        
           |  |        +-- Diaurora                    
           |  +--------+                               
           +--+        +-- Schistodesmus               
              |                                        
              |        +-- Pseudocuneopsis_sichuanensis **
              |  +-----+                               
              |  |     +-- Pseudocuneopsis_capitata **    
              +--+                                     
                 |  +----- Unio                        
                 +--+                                  
                    |  +-- Tchangsinaia                
                    +--+                               
                       +-- Inversiunio

Chen et al., 2025 [Publ 4448], fig. 2

iTtTbiriSoTroTroSiD
6 6 4 6 3 2 1 3 
  +-------------- Nodularia                    
  |                                            
--+  +----------- Pseudocuneopsis_perflora **     
  |  |                                         
  +--+  +-------- Pseudocuneopsis_capitata **     
     |  |                                      
     +--+     +-- Pseudocuneopsis_yangshuoensis **
        |  +--+                                
        |  |  +-- Pseudocuneopsis_wuana **        
        +--+                                   
           |  +-- Pseudocuneopsis_yemaoi **       
           +--+                                
              +-- Pseudocuneopsis_sichuanensis **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
6 6 4 6 80 22 7 1 
  +-------------------- Aculamprotula                
  |                                                  
  |  +----------------- Unio                         
--+  |                                               
  |  |        +-------- Nodularia                    
  |  |        |                                      
  +--+  +-----+     +-- Tongchuania                  
     |  |     |  +--+                                
     |  |     |  |  +-- Schistodesmus                
     |  |     +--+                                   
     |  |        |  +-- Tchangsinaia                 
     +--+        +--+                                
        |           +-- Cuneopsis                    
        |                                            
        |  +----------- Pseudocuneopsis_capitata **     
        |  |                                         
        +--+  +-------- Pseudocuneopsis_perflora **     
           |  |                                      
           +--+     +-- Pseudocuneopsis_sichuanensis ** 
              |  +--+                                
              |  |  +-- Pseudocuneopsis_yemaoi **       
              +--+                                   
                 |  +-- Pseudocuneopsis_wuana **        
                 +--+                                
                    +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 71 44 7 2 
  +-------------------- Aculamprotula                
  |                                                  
--+  +----------------- Unio                         
  |  |                                               
  +--+  +-------------- Middendorffinaia             
     |  |                                            
     |  |     +-------- Nodularia                    
     +--+  +--+                                      
        |  |  |  +----- Schistodesmus                
        |  |  +--+                                   
        |  |     |  +-- Tchangsinaia                 
        +--+     +--+                                
           |        +-- Cuneopsis                    
           |                                         
           |  +-------- Pseudocuneopsis_capitata **     
           |  |                                      
           +--+     +-- Pseudocuneopsis_sichuanensis ** 
              |  +--+                                
              |  |  +-- Pseudocuneopsis_yemaoi **       
              +--+                                   
                 |  +-- Pseudocuneopsis_wuana **        
                 +--+                                
                    +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2025 [Publ 4432], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 23 21 5 2 
  +-------------- Middendorffinaia             
  |                                            
  |     +-------- Nodularia                    
--+  +--+                                      
  |  |  |  +----- Schistodesmus                
  |  |  +--+                                   
  |  |     |  +-- Tchangsinaia                 
  +--+     +--+                                
     |        +-- Cuneopsis                    
     |                                         
     |  +-------- Pseudocuneopsis_capitata **     
     |  |                                      
     +--+     +-- Pseudocuneopsis_sichuanensis ** 
        |  +--+                                
        |  |  +-- Pseudocuneopsis_yemaoi **       
        +--+                                   
           |  +-- Pseudocuneopsis_wuana **        
           +--+                                
              +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2025 [Publ 4401], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 23 21 5 2 
  +-------------- Middendorffinaia             
  |                                            
  |     +-------- Nodularia                    
--+  +--+                                      
  |  |  |  +----- Schistodesmus                
  |  |  +--+                                   
  |  |     |  +-- Tchangsinaia                 
  +--+     +--+                                
     |        +-- Cuneopsis                    
     |                                         
     |  +-------- Pseudocuneopsis_capitata **     
     |  |                                      
     +--+     +-- Pseudocuneopsis_sichuanensis ** 
        |  +--+                                
        |  |  +-- Pseudocuneopsis_yemaoi **       
        +--+                                   
           |  +-- Pseudocuneopsis_wuana **        
           +--+                                
              +-- Pseudocuneopsis_yangshuoensis **

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 10 8 3 2 
        +----------- Tchangsinaia                 
        |                                         
  +-----+        +-- Cuneopsis_kiangsiensis       
  |     |  +-----+                                
  |     |  |     +-- Cuneopsis_celtiformis        
  |     +--+                                      
  |        |  +----- Cuneopsis_szechenyii         
  |        +--+                                   
--+           |  +-- Cuneopsis_heudei             
  |           +--+                                
  |              +-- Cuneopsis_demangei           
  |                                               
  |  +-------------- Cuneopsis_rufescens          
  |  |                                            
  +--+  +----------- Middendorffinaia             
     |  |                                         
     +--+  +-------- Pseudocuneopsis_capitata **     
        |  |                                      
        +--+     +-- Pseudocuneopsis_sichuanensis ** 
           |  +--+                                
           |  |  +-- Pseudocuneopsis_yemaoi **       
           +--+                                   
              |  +-- Pseudocuneopsis_wuana **        
              +--+                                
                 +-- Pseudocuneopsis_yangshuoensis **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
4 4 2 4 20 10 7 3 
  +----------------- Unio                         
  |                                               
  +----------------- Aculamprotula                
  |                                               
--+  +-------------- Cuneopsis                    
  |  |                                            
  |  |           +-- Nodularia                    
  |  +-----------+                                
  +--+           +-- Schistodesmus                
     |                                            
     |  +----------- Tchangsinaia                 
     |  |                                         
     +--+----------- Inversiunio                  
        |                                         
        |  +-------- Pseudocuneopsis_capitata **     
        +--+                                      
           |  +----- Pseudocuneopsis_sichuanensis ** 
           +--+                                   
              |  +-- Pseudocuneopsis_wuana **        
              +--+                                
                 +-- Pseudocuneopsis_yangshuoensis **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 4 20 10 7 3 
  +-------------------------- Unio                         
  |                                                        
--+  +----------------------- Aculamprotula                
  |  |                                                     
  +--+  +-------------------- Cuneopsis                    
     |  |                                                  
     +--+                 +-- Nodularia                    
        |  +--------------+                                
        |  |              +-- Schistodesmus                
        +--+                                               
           |  +-------------- Tchangsinaia                 
           +--+                                            
              |  +----------- Inversiunio                  
              +--+                                         
                 |  +-------- Pseudocuneopsis_capitata **     
                 +--+                                      
                    |  +----- Pseudocuneopsis_sichuanensis ** 
                    +--+                                   
                       |  +-- Pseudocuneopsis_wuana **        
                       +--+                                
                          +-- Pseudocuneopsis_yangshuoensis **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 4 49 14 6 3 
  +----------------- Aculamprotula                
  |                                               
--+  +-------------- Unio                         
  |  |                                            
  |  |     +-------- Pseudocuneopsis_capitata **     
  +--+  +--+                                      
     |  |  |  +----- Pseudocuneopsis_yangshuoensis **
     |  |  +--+                                   
     |  |     |  +-- Pseudocuneopsis_yemaoi **       
     +--+     +--+                                
        |        +-- Pseudocuneopsis_sichuanensis ** 
        |                                         
        |  +-------- Nodularia                    
        +--+                                      
           |  +----- Schistodesmus                
           +--+                                   
              |  +-- Tchangsinaia                 
              +--+                                
                 +-- Cuneopsis

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 84 25 8 3 
  +----------------------- Globunio                     
  |                                                     
--+  +-------------------- Aculamprotula                
  |  |                                                  
  +--+  +----------------- Unio                         
     |  |                                               
     |  |           +----- Schistodesmus                
     +--+  +--------+                                   
        |  |        |  +-- Tchangsinaia                 
        |  |        +--+                                
        +--+           +-- Cuneopsis                    
           |                                            
           |  +----------- Nodularia                    
           +--+                                         
              |  +-------- Pseudocuneopsis_capitata **     
              +--+                                      
                 |  +----- Pseudocuneopsis_yangshuoensis **
                 +--+                                   
                    |  +-- Pseudocuneopsis_sichuanensis ** 
                    +--+                                
                       +-- Pseudocuneopsis_yemaoi **

Dai et al., 2024 [Publ 4358], fig. 3

iTtTbiriSoTroTroSiD
4 4 2 4 2 2 2 3 
           +-- Schistodesmus                
  +--------+                                
  |        +-- Nodularia                    
--+                                         
  |  +-------- Pseudocuneopsis_capitata **     
  +--+                                      
     |  +----- Pseudocuneopsis_yangshuoensis **
     +--+                                   
        |  +-- Pseudocuneopsis_sichuanensis ** 
        +--+                                
           +-- Pseudocuneopsis_yemaoi **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 20 10 7 4 
  +----------------------- Unio                         
  |                                                     
--+  +-------------------- Aculamprotula                
  |  |                                                  
  +--+  +----------------- Cuneopsis                    
     |  |                                               
     +--+              +-- Nodularia                    
        |  +-----------+                                
        |  |           +-- Schistodesmus                
        +--+                                            
           |  +----------- Tchangsinaia                 
           +--+                                         
              |  +-------- Inversiunio                  
              +--+                                      
                 |  +----- Pseudocuneopsis_capitata **     
                 +--+                                   
                    |  +-- Pseudocuneopsis_yangshuoensis **
                    +--+                                
                       +-- Pseudocuneopsis_sichuanensis **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
2 2 0 2 90 21 6 -- 
  +-------------- Aculamprotula               
  |                                           
--+  +----------- Unio                        
  |  |                                        
  +--+  +-------- Nodularia                   
     |  |                                     
     |  |     +-- Pseudocuneopsis_capitata **    
     +--+-----+                               
        |     +-- Pseudocuneopsis_sichuanensis **
        |                                     
        |  +----- Schistodesmus               
        +--+                                  
           |  +-- Tchangsinaia                
           +--+                               
              +-- Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 72 19 6 -- 
  +----------------- Aculamprotula               
  |                                              
--+  +-------------- Unio                        
  |  |                                           
  +--+           +-- Pseudocuneopsis_capitata **    
     |  +--------+                               
     |  |        +-- Pseudocuneopsis_sichuanensis **
     +--+                                        
        |  +-------- Nodularia                   
        +--+                                     
           |  +----- Schistodesmus               
           +--+                                  
              |  +-- Tchangsinaia                
              +--+                               
                 +-- Cuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
2 2 0 2 94 21 6 -- 
  +-------------------- Aculamprotula               
  |                                                 
--+  +----------------- Unio                        
  |  |                                              
  +--+              +-- Pseudocuneopsis_capitata **    
     |  +-----------+                               
     |  |           +-- Pseudocuneopsis_sichuanensis **
     +--+                                           
        |  +----------- Nodularia                   
        |  |                                        
        +--+        +-- Schistodesmus_lampreyanus   
           |  +-----+                               
           |  |     +-- Cuneopsis                   
           +--+                                     
              |  +----- Tchangsinaia                
              +--+                                  
                 |  +-- Schistodesmus_xinyuensis    
                 +--+                               
                    +-- Schistodesmus_spinosus

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 39 16 9 -- 
  +----------------- Aculamprotula               
  |                                              
  |              +-- Middendorffinaia            
--+  +-----------+                               
  |  |           +-- Nodularia                   
  |  |                                           
  +--+  +----------- Cuneopsis                   
     |  |                                        
     |  |        +-- Diaurora                    
     |  +--------+                               
     +--+        +-- Schistodesmus               
        |                                        
        |        +-- Pseudocuneopsis_sichuanensis **
        |  +-----+                               
        |  |     +-- Pseudocuneopsis_capitata **    
        +--+                                     
           |  +----- Inversiunio                 
           +--+                                  
              |  +-- Tchangsinaia                
              +--+                               
                 +-- Unio

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 43 20 10 -- 
  +-------------------- Globunio                    
  |                                                 
--+  +----------------- Aculamprotula               
  |  |                                              
  |  |              +-- Middendorffinaia            
  +--+  +-----------+                               
     |  |           +-- Nodularia                   
     |  |                                           
     +--+  +----------- Cuneopsis                   
        |  |                                        
        |  |        +-- Diaurora                    
        |  +--------+                               
        +--+        +-- Schistodesmus               
           |                                        
           |        +-- Pseudocuneopsis_sichuanensis **
           |  +-----+                               
           |  |     +-- Pseudocuneopsis_capitata **    
           +--+                                     
              |  +----- Inversiunio                 
              +--+                                  
                 |  +-- Tchangsinaia                
                 +--+                               
                    +-- Unio

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
2 2 0 2 37 22 7 -- 
  +-------------------- Aculamprotula               
  |                                                 
--+  +----------------- Unio                        
  |  |                                              
  +--+              +-- Pseudocuneopsis_capitata **    
     |  +-----------+                               
     |  |           +-- Pseudocuneopsis_sichuanensis **
     +--+                                           
        |  +----------- Nodularia                   
        +--+                                        
           |  +-------- Schistodesmus               
           +--+                                     
              |  +----- Cuneopsis                   
              +--+                                  
                 |  +-- Tchangsinaia                
                 +--+                               
                    +-- Inversiunio

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
2 2 0 2 16 11 6 -- 
  +----------------- Aculamprotula               
  |                                              
--+  +-------------- Unio                        
  |  |                                           
  +--+           +-- Pseudocuneopsis_sichuanensis **
     |  +--------+                               
     |  |        +-- Pseudocuneopsis_capitata **    
     +--+                                        
        |  +-------- Nodularia                   
        +--+                                     
           |  +----- Tchangsinaia                
           +--+                                  
              |  +-- Cuneopsis                   
              +--+                               
                 +-- Schistodesmus

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
2 2 0 2 94 21 6 -- 
  +-------------- Aculamprotula               
  |                                           
--+  +----------- Unio                        
  |  |                                        
  |  |     +----- Nodularia                   
  +--+  +--+                                  
     |  |  |  +-- Pseudocuneopsis_capitata **    
     |  |  +--+                               
     +--+     +-- Pseudocuneopsis_sichuanensis **
        |                                     
        |  +----- Tchangsinaia                
        +--+                                  
           |  +-- Schistodesmus               
           +--+                               
              +-- Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
2 2 0 2 10 10 6 -- 
  +----------------- Aculamprotula               
  |                                              
--+  +-------------- Nodularia                   
  |  |                                           
  +--+           +-- Pseudocuneopsis_capitata **    
     |  +--------+                               
     |  |        +-- Pseudocuneopsis_sichuanensis **
     +--+                                        
        |  +-------- Schistodesmus               
        +--+                                     
           |  +----- Cuneopsis                   
           +--+                                  
              |  +-- Tchangsinaia                
              +--+                               
                 +-- Inversiunio

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 51 31 6 -- 
  +----------------- Aculamprotula               
  |                                              
--+  +-------------- Unio                        
  |  |                                           
  +--+           +-- Pseudocuneopsis_sichuanensis **
     |  +--------+                               
     |  |        +-- Pseudocuneopsis_capitata **    
     +--+                                        
        |  +-------- Nodularia                   
        +--+                                     
           |  +----- Schistodesmus               
           +--+                                  
              |  +-- Tchangsinaia                
              +--+                               
                 +-- Cuneopsis

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 51 27 6 -- 
  +----------------- Aculamprotula               
  |                                              
--+  +-------------- Unio                        
  |  |                                           
  +--+           +-- Pseudocuneopsis_capitata **    
     |  +--------+                               
     |  |        +-- Pseudocuneopsis_sichuanensis **
     +--+                                        
        |  +-------- Nodularia                   
        +--+                                     
           |  +----- Schistodesmus               
           +--+                                  
              |  +-- Tchangsinaia                
              +--+                               
                 +-- Cuneopsis

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
2 2 0 2 90 21 6 -- 
  +-------------- Aculamprotula               
  |                                           
--+  +----------- Unio                        
  |  |                                        
  +--+  +-------- Nodularia                   
     |  |                                     
     |  |     +-- Pseudocuneopsis_capitata **    
     +--+-----+                               
        |     +-- Pseudocuneopsis_sichuanensis **
        |                                     
        |  +----- Schistodesmus               
        +--+                                  
           |  +-- Tchangsinaia                
           +--+                               
              +-- Cuneopsis

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 16 9 -- 
  +----------------- Aculamprotula               
  |                                              
  |              +-- Middendorffinaia            
--+  +-----------+                               
  |  |           +-- Nodularia                   
  |  |                                           
  +--+  +----------- Cuneopsis                   
     |  |                                        
     |  |        +-- Diaurora                    
     |  +--------+                               
     +--+        +-- Schistodesmus               
        |                                        
        |        +-- Pseudocuneopsis_sichuanensis **
        |  +-----+                               
        |  |     +-- Pseudocuneopsis_capitata **    
        +--+                                     
           |  +----- Inversiunio                 
           +--+                                  
              |  +-- Tchangsinaia                
              +--+                               
                 +-- Unio

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 16 10 -- 
  +----------------- Aculamprotula               
  |                                              
--+  +-------------- Unio                        
  |  |                                           
  |  |           +-- Tchangsinaia                
  +--+  +--------+                               
     |  |        +-- Inversiunio                 
     |  |                                        
     +--+     +----- Cuneopsis                   
        |  +--+                                  
        |  |  |  +-- Pseudocuneopsis_sichuanensis **
        |  |  +--+                               
        +--+     +-- Pseudocuneopsis_capitata **    
           |                                     
           |  +----- Nodularia                   
           +--+                                  
              |  +-- Diaurora                    
              +--+                               
                 +-- Schistodesmus

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 39 16 10 -- 
                       +-- Globunio                    
  +--------------------+                               
  |                    +-- Huizhou                     
--+                                                    
  |  +-------------------- Aculamprotula               
  |  |                                                 
  +--+  +----------------- Unio                        
     |  |                                              
     |  |              +-- Tchangsinaia                
     +--+  +-----------+                               
        |  |           +-- Inversiunio                 
        |  |                                           
        +--+           +-- Pseudocuneopsis_sichuanensis **
           |  +--------+                               
           |  |        +-- Pseudocuneopsis_capitata **    
           +--+                                        
              |  +-------- Cuneopsis                   
              +--+                                     
                 |  +----- Nodularia                   
                 +--+                                  
                    |  +-- Diaurora                    
                    +--+                               
                       +-- Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 15 6 -- 
  +----------------- Aculamprotula           
--+                                          
  |  +-------------- Unio                    
  +--+                                       
     |  +----------- Pseudocuneopsis_capitata **
     +--+                                    
        |  +-------- Nodularia               
        +--+                                 
           |  +----- Tchangsinaia            
           +--+                              
              |  +-- Cuneopsis               
              +--+                           
                 +-- Schistodesmus

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 40 20 6 -- 
  +----------------- Aculamprotula           
--+                                          
  |  +-------------- Unio                    
  +--+                                       
     |  +----------- Pseudocuneopsis_capitata **
     +--+                                    
        |  +-------- Nodularia               
        +--+                                 
           |  +----- Schistodesmus           
           +--+                              
              |  +-- Tchangsinaia            
              +--+                           
                 +-- Cuneopsis

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 13 6 -- 
  +----------------- Aculamprotula           
--+                                          
  |  +-------------- Unio                    
  +--+                                       
     |  +----------- Pseudocuneopsis_capitata **
     +--+                                    
        |  +-------- Nodularia               
        +--+                                 
           |  +----- Tchangsinaia            
           +--+                              
              |  +-- Cuneopsis               
              +--+                           
                 +-- Schistodesmus

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 6 6 -- 
  +-------------- Aculamprotula           
  |                                       
--+  +----------- Unio                    
  |  |                                    
  +--+        +-- Pseudocuneopsis_capitata **
     |  +-----+                           
     |  |     +-- Nodularia               
     +--+                                 
        |  +----- Schistodesmus           
        +--+                              
           |  +-- Cuneopsis               
           +--+                           
              +-- Tchangsinaia

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 53 8 6 -- 
  +----------------- Aculamprotula               
--+                                              
  |  +-------------- Unio                        
  +--+                                           
     |  +----------- Pseudocuneopsis_sichuanensis **
     +--+                                        
        |  +-------- Nodularia                   
        +--+                                     
           |  +----- Tchangsinaia                
           +--+                                  
              |  +-- Cuneopsis                   
              +--+                               
                 +-- Schistodesmus

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 13 6 4 -- 
  +-------------- Tchangsinaia            
--+                                       
  |  +----------- Cuneopsis_heudei        
  +--+                                    
     |  +-------- Nodularia               
     +--+                                 
        |  +----- Pseudocuneopsis_capitata **
        +--+                              
           |  +-- Cuneopsis_rufescens     
           +--+                           
              +-- Schistodesmus

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 53 12 6 -- 
  +-------------- Aculamprotula           
  |                                       
--+  +----------- Unio                    
  |  |                                    
  +--+        +-- Cuneopsis               
     |  +-----+                           
     |  |     +-- Schistodesmus           
     +--+                                 
        |  +----- Pseudocuneopsis_capitata **
        +--+                              
           |  +-- Nodularia               
           +--+                           
              +-- Tchangsinaia

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 16 12 6 -- 
  +----------- Unio                    
  |                                    
--+  +-------- Pseudocuneopsis_capitata **
  |  |                                 
  +--+     +-- Cuneopsis               
     |  +--+                           
     |  |  +-- Schistodesmus           
     +--+                              
        |  +-- Tchangsinaia            
        +--+                           
           +-- Nodularia

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 29 10 6 -- 
  +----------------- Aculamprotula           
--+                                          
  |  +-------------- Unio                    
  +--+                                       
     |  +----------- Pseudocuneopsis_capitata **
     +--+                                    
        |  +-------- Nodularia               
        +--+                                 
           |  +----- Schistodesmus           
           +--+                              
              |  +-- Cuneopsis               
              +--+                           
                 +-- Tchangsinaia

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 15 6 -- 
  +----------------- Aculamprotula           
--+                                          
  |  +-------------- Unio                    
  +--+                                       
     |  +----------- Pseudocuneopsis_capitata **
     +--+                                    
        |  +-------- Nodularia               
        +--+                                 
           |  +----- Schistodesmus           
           +--+                              
              |  +-- Tchangsinaia            
              +--+                           
                 +-- Cuneopsis

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 15 6 -- 
  +----------------- Aculamprotula           
--+                                          
  |  +-------------- Unio                    
  +--+                                       
     |  +----------- Pseudocuneopsis_capitata **
     +--+                                    
        |  +-------- Nodularia               
        +--+                                 
           |  +----- Schistodesmus           
           +--+                              
              |  +-- Cuneopsis               
              +--+                           
                 +-- Tchangsinaia

Hou et al., 2025 [Publ 4432], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 10 9 3 -- 
  +-------- Tchangsinaia            
--+                                 
  |  +----- Pseudocuneopsis_capitata **
  +--+                              
     |  +-- Middendorffinaia        
     +--+                           
        +-- Cuneopsis

Wu et al., 2024 [Publ 4379], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 11 11 3 -- 
  +----- Nodularia               
  |                              
--+  +-- Cuneopsis               
  |  |                           
  +--+-- Tchangsinaia            
     |                           
     +-- Pseudocuneopsis_capitata **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 12 6 -- 
  +----------------- Aculamprotula           
--+                                          
  |  +-------------- Unio                    
  +--+                                       
     |  +----------- Pseudocuneopsis_capitata **
     +--+                                    
        |  +-------- Nodularia               
        +--+                                 
           |  +----- Tchangsinaia            
           +--+                              
              |  +-- Cuneopsis               
              +--+                           
                 +-- Schistodesmus
 
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