The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
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Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini genus Unio Philipsson in Retzius, 1788: cladomics
29 Recent valid species (45 total species) Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to
provide statistics and references indicating the extent to which published phylogenetic
work supports the classification used here. The Cladomics Summary provides an
overview of the number of studies that have included terminal species representing this
taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently
asked rhetorical questions) page. Abbreviations for the various statistics are explained on
the FARQ! Cladomics Summarynumber of species = 29 (45 total)
analyzed in 218 cladograms in 128 publications
maximum number of terminals in an analysis =
23 (mean = 3.3, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 150
maximum number of included terminals = 23, (mean =
4.34 median = 3)
proportion of cladograms that support basic monophyly
= 0.93
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup
distances within the cladome. See the Cladmoics
FARQ for more information.
Lyubas et al., 2022 [Publ 4255], fig. 7 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 23 | 23 | 20 | 23 | 8 | 0 | 0 | 0 | +----------------------------- Unio_tumidus **
|
| +-- Unio_durieui **
| +-----------------------+
| | +-- Unio_gibbus **
--+ |
| | +-------------- Unio_damascensis **
| | |
| | +-------------- Unio_sesirmensis **
| | +--------+
+--+ | | +----------- Unio_ionicus **
| | | |
| | +--+ +-- Unio_bruguierianus **
| | | +-----+
| | | | +-- Unio_tumidiformis **
| | +--+
| | | +----- Unio_vicarius **
+--+ +--+
| | +-- Unio_carneus **
| +--+
| +-- Unio_crassus **
|
| +----- Unio_caffer **
| +--------------+
| | | +-- Unio_terminalis **
| | +--+
+--+ +-- Unio_delicatus **
|
| +----------------- Unio_pictorum **
| |
+--+ +-------------- Unio_eucirrus **
| |
+--+ +-- Unio_elongatulus **
| +--------+
| | +-- Unio_mancus **
+--+
| +-------- Unio_ravoisieri **
| |
+--+ +-- Unio_delphinus **
| +--+
| | +-- Unio_foucauldianus **
+--+
| +-- Unio_hueti **
+--+
+-- Unio_tigridis **
Dai et al., 2025 [Publ 4440], fig. 1 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 20 | 10 | -- | +----------------------- Globunio
|
--+ +-------------------- Aculamprotula
| |
+--+ +----------------- Middendorffinaia
| |
+--+ +-------------- Nodularia
| |
| | +----------- Cuneopsis
+--+ |
| | +-- Diaurora
| +--------+
+--+ +-- Schistodesmus
|
| +-------- Pseudocuneopsis
| |
+--+ +-- Unio_pictorum **
| +--+
| | +-- Unio_nanus **
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Lyubas et al., 2022 [Publ 4255], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 23 | 23 | 14 | 23 | 8 | 0 | 0 | 10 | +----------------- Unio_tumidus **
|
| +-- Unio_durieui **
+--------------+
| +-- Unio_gibbus **
|
| +----------- Unio_sesirmensis **
| |
| +--+ +-------- Unio_damascensis **
--+ | | |
| | | | +----- Unio_ionicus **
| | +--+ |
| | | | +-- Unio_bruguierianus **
| | | +--+
| | +--+ +-- Unio_tumidiformis **
| | |
| | | +-- Unio_carneus **
| | | |
| | +--+-- Unio_crassus **
+--+ |
| +-- Unio_vicarius **
|
| +-------- Unio_elongatulus **
| |
| +-------- Unio_mancus **
| |
| | +-- Unio_pictorum **
| +-----+
| | +-- Unio_eucirrus **
| |
+-----+ +----- Unio_caffer **
+--+
| | +-- Unio_terminalis **
| +--+
| +-- Unio_delicatus **
|
| +----- Unio_ravoisieri **
| |
| | +-- Unio_delphinus **
+--+--+
| +-- Unio_foucauldianus **
|
| +-- Unio_hueti **
+--+
+-- Unio_tigridis **
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 19 | 17 | 19 | 57 | 30 | 7 | 29 | +-------------------------------------- Aculamprotula
|
| +-------------- Middendorffinaia
| +--------------------+
| | | +----------- Pseudocuneopsis
--+ | +--+
| | | +-------- Nodularia
| | +--+
| | | +----- Schistodesmus
| | +--+
| | | +-- Tchangsinaia
+--+ +--+
| +-- Cuneopsis
|
| +-- Unio_gibbus **
| +-----------------------------+
| | +-- Unio_tumidus **
| |
| | +----------- Unio_terminalis **
+--+ +-----------------+
| | | +-------- Unio_elongatulus **
| | +--+
| | | +----- Unio_pictorum **
| | +--+
+--+ | +-- Unio_delphinus **
| +--+
| +-- Unio_mancus **
|
| +-------------------------- Unio_tumidiformis **
| |
+--+ +----------------------- Unio_desectus **
| |
+--+ +-------------------- Unio_sesirmensis **
| |
+--+ +----------------- Unio_ionicus **
| |
+--+ +-------------- Unio_damascensis **
| |
| | +-- Unio_crassus **
+--+ +--------+
| | +-- Unio_gontierii **
| |
+--+ +-- Unio_bruguierianus **
| +-----+
| | +-- Unio_mardinensis **
+--+
| +----- Unio_vicarius **
+--+
| +-- Unio_carneus **
+--+
+-- Unio_nanus **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 17 | 5 | 2 | 2 | 27 | +-- Tchangsinaia
+--------------------------------+
| +-- Nodularia
|
--+ +-- Unio_gibbus **
| +-----------------------------+
| | +-- Unio_tumidus **
| |
+--+ +----- Unio_terminalis **
| +-----------------------+
| | | +-- Unio_delphinus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-------------------------- Unio_tumidiformis **
| |
+--+ +----------------------- Unio_sesirmensis **
| |
+--+ +-------------------- Unio_ionicus **
| |
+--+ +----------------- Unio_damascensis **
| |
+--+ +-------------- Unio_desectus **
| |
| | +-- Unio_crassus **
+--+ +--------+
| | +-- Unio_gontierii **
| |
+--+ +-- Unio_bruguierianus **
| +-----+
| | +-- Unio_mardinensis **
+--+
| +----- Unio_vicarius **
+--+
| +-- Unio_carneus **
+--+
+-- Unio_nanus **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 15 | 17 | 5 | 2 | 2 | 27 | +-- Tchangsinaia
+--------------------------+
| +-- Nodularia
--+
| +-------------------------- Unio_gibbus **
| |
+--+ +----------------------- Unio_tumidus **
| |
| | +----- Unio_terminalis **
+--+ +--------------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
+--+
| +----------------- Unio_tumidiformis **
| |
| | +-- Unio_ionicus **
+--+ +-----------+
| | +-- Unio_carneus **
| |
| | +-- Unio_damascensis **
+--+ +--------+
| | +-- Unio_sesirmensis **
| |
| | +----- Unio_bruguierianus **
+--+ +--+
| | | +-- Unio_gontierii **
| | +--+
| | +-- Unio_mardinensis **
+--+
| +-- Unio_crassus **
| +--+
| | +-- Unio_nanus **
+--+
| +-- Unio_vicarius **
+--+
+-- Unio_desectus **
Lopes-Lima et al., 2021 [Publ 4217], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 14 | 16 | 9 | 0 | 0 | 18 | +----------------------- Unio_tumidus **
|
| +-------- Unio_damascensis **
| +-----------+
--+ | | +----- Unio_sesirmensis **
| | +--+
| | | +-- Unio_bruguierianus **
| | +--+
+--+ +-- Unio_nanus **
|
| +-- Unio_delicatus **
| +--------------+
| | +-- Unio_terminalis **
| |
+--+ +-- Unio_tigridis **
| +-----------+
| | +-- Unio_hueti **
| |
+--+ +----------- Unio_ravoisieri **
| |
| | +-- Unio_pictorum **
+--+ +-----+
| | +-- Unio_eucirrus **
| |
+--+ +-- Unio_elongatulus **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_foucauldianus **
+--+
+-- Unio_delphinus **
Lopes-Lima et al., 2024 [Publ 4376], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 18 | 0 | 0 | 0 | 29 | +-- Unio_pictorum **
+--------------------------+
| +-- Unio_delphinus **
--+
| +-------------------------- Unio_tumidiformis **
| |
+--+ +----------------------- Unio_sesirmensis **
| |
+--+ +-------------------- Unio_desectus **
| |
+--+ +----------------- Unio_damascensis **
| |
| | +-------------- Unio_ionicus **
+--+ |
| | +-- Unio_nanus **
| | +--------+
+--+ | +-- Unio_carneus **
| |
| | +-- Unio_gontierii **
| | |
| | +-- Unio_crassus__1 **
+--+--------+
| +-- Unio_crassus__2 **
| |
| +-- Unio_crassus__3 **
|
| +-------- Unio_vicarius **
| |
+--+ +----- Unio_bruguierianus__1 **
| |
+--+----- Unio_bruguierianus__2 **
|
| +-- Unio_bruguierianus__3 **
+--+
+-- Unio_mardinensis **
Araujo et al., 2018 [Publ 3779], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 9 | 14 | 5 | 1 | 1 | 20 | +-------------------- Nodularia
|
| +-- Unio_durieui **
--+ +--------------+
| | +-- Unio_gibbus **
| |
+--+ +-------------- Unio_tumidus **
| |
| | +-------- Unio_tumidiformis **
| | +--+
+--+ | | +----- Unio_desectus **
| | +--+
| | | +-- Unio__1 **
| | +--+
+--+ +-- Unio_ionicus **
|
| +-- Unio_delphinus **
| +--+
| | +-- Unio_foucauldianus **
| |
+-----+ +-- Unio_pictorum **
| |
| +-- Unio_elongatulus **
| |
+--+-- Unio_mancus **
|
+-- Unio_ravoisieri **
|
+-- Unio__2 **
Froufe et al., 2016 [Publ 3738], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 12 | 2 | 2 | 1 | 25 | +----------------------------- Nodularia
|
--+ +-- Unio_gibbus **
| +-----------------------+
| | +-- Unio_durieui **
+--+
| +----------------------- Unio_tumidus **
| |
+--+ +-- Unio **
| +-----------------+
| | +-- Unio_tumidiformis **
+--+
| +----------------- Unio_delphinus **
+--+
| +-------------- Unio_foucauldianus **
+--+
| +----------- Unio_ravoisieri **
+--+
| +-------- Unio_mancus **
+--+
| +----- Unio_elongatulus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Prié & Puillandre, 2014 [Publ 3529], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 11 | 6 | 0 | 0 | 22 | +----- Unio_tumidus **
+-----------+
| | +-- Unio_gibbus **
| +--+
| +-- Unio_durieui **
--+
| +----- Unio_tumidiformis **
| +--------+
| | | +-- Unio **
| | +--+
+--+ +-- Unio_crassus **
|
| +----------- Unio_delphinus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_mancus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_ravoisieri **
Lopes-Lima et al., 2021 [Publ 4217], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 0 | 0 | 0 | 17 | +----------------- Unio_tumidus **
|
--+ +-------------- Unio_bruguierianus **
| |
| | +-- Unio_delicatus **
+--+ +--------+
| | +-- Unio_terminalis **
| |
+--+ +-- Unio_tigridis **
| +-----+
| | +-- Unio_hueti **
| |
+--+ +-- Unio_pictorum **
| +--+
| | +-- Unio_eucirrus **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Khalloufi et al., 2011 [Publ 2748], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 6 | 9 | 2 | 0 | 0 | 18 | +----- Unio_tumidus **
+-----+
| | +-- Unio_gibbus **
| +--+
| +-- Unio_durieui **
--+
| +-- Unio_tumidiformis **
| +-----+
| | +-- Unio_crassus **
+--+
| +----- Unio **
| |
+--+----- Unio_pictorum **
|
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 5 | 9 | 8 | 0 | 0 | 22 | +-- Unio_tumidus **
+--------+
| +-- Unio_gibbus **
|
--+ +-- Unio **
| +-----+
| | +-- Unio_tumidiformis **
+--+
| +----- Unio_ravoisieri **
| |
+--+ +-- Unio_mancus **
| |
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
|
+-- Unio_elongatulus **
Marrone et al., 2019 [Publ 3907], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 4 | 8 | 0 | 0 | 0 | 23 | +-------------- Unio_tumidus **
|
--+ +----------- Unio_delphinus **
| |
+--+ +-------- Unio_foucauldianus **
| |
+--+ +----- Unio_mancus **
| |
| +----- Unio_elongatulus **
+--+
+----- Unio_ravoisieri **
|
| +-- Unio_pictorum **
+--+
+-- Unio_terminalis **
Araujo et al., 2009 [Publ 2589], fig. 7B iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 8 | 4 | 0 | 0 | 19 | +-------------- Unio_gibbus **
|
--+ +----------- Unio_tumidus **
| |
| | +-- Unio_tumidiformis **
+--+ +-----+
| | +-- Unio_crassus **
| |
+--+ +-- Unio_pictorum__1 **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_pictorum__2 **
+--+
+-- Unio_caffer **
Reis et al., 2013 [Publ 3580], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 8 | 8 | 0 | 0 | 20 | +-------------- Unio_tumidus **
|
--+ +-- Unio_tumidiformis **
| +--------+
| | +-- Unio__2 **
+--+
| +-------- Unio__1 **
| |
+--+ +-- Unio_delphinus **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Lopes-Lima et al., 2021 [Publ 4217], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 7 | 0 | 0 | 0 | 21 | +----------------- Unio_tumidus **
--+
| +-------------- Unio_pictorum **
+--+
| +----------- Unio_sesirmensis **
+--+
| +-------- Unio_damascensis **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio **
+--+
+-- Unio_nanus **
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 34 | 14 | 6 | 21 | +-------------------- Aculamprotula
|
| +----------- Pseudocuneopsis
| +-----+
--+ | | +-------- Nodularia
| | +--+
| | | +----- Schistodesmus
| | +--+
+--+ | +-- Tchangsinaia
| +--+
| +-- Cuneopsis
|
| +-------------- Unio_tumidus **
+--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 47 | 23 | 6 | 21 | +-------------------- Aculamprotula
|
| +----------- Pseudocuneopsis
| +-----+
--+ | | +-------- Nodularia
| | +--+
| | | +----- Schistodesmus
| | +--+
+--+ | +-- Tchangsinaia
| +--+
| +-- Cuneopsis
|
| +-------------- Unio_tumidus **
+--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 85 | 16 | 6 | 23 | +-------------------- Aculamprotula
|
| +-------- Nodularia
| |
--+ +--------+-------- Pseudocuneopsis
| | |
| | | +----- Schistodesmus
| | +--+
+--+ | +-- Tchangsinaia
| +--+
| +-- Cuneopsis
|
| +-------------- Unio_tumidus **
+--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 82 | 23 | 8 | 21 | +----------------------- Globunio
|
| +-------------------- Aculamprotula
--+ |
| | +-- Nodularia
| | +-----+
+--+ | +-- Pseudocuneopsis
| +--------+
| | | +----- Schistodesmus
| | +--+
| | | +-- Tchangsinaia
+--+ +--+
| +-- Cuneopsis
|
| +-------------- Unio_tumidus **
+--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Araujo et al., 2009 [Publ 2589], fig. 7C iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 7 | 4 | 0 | 0 | 20 | +-- Unio_gibbus **
+-----+
| +-- Unio_tumidus **
|
--+ +-- Unio_tumidiformis **
| +--+
| | +-- Unio_crassus **
+--+
| +-- Unio_pictorum__1 **
| |
+--+-- Unio_pictorum__2 **
|
+-- Unio_mancus **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 6 | 89 | 16 | 6 | 22 | +-------------------- Aculamprotula
|
| +----------- Unio_tumidus **
| |
| +-----+ +-------- Unio_nanus **
--+ | | |
| | +--+ +----- Unio_elongatulus **
| | | |
| | +--+ +-- Unio_delphinus **
| | | |
+--+ +--+-- Unio_mancus **
| |
| +-- Unio_pictorum **
|
| +-------------- Pseudocuneopsis
| |
+--+ +----------- Nodularia
| |
+--+ +-- Schistodesmus_lampreyanus__1
| +-----+
| | +-- Cuneopsis
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus_lampreyanus__2
+--+
+-- Schistodesmus_spinosus
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 85 | 16 | 6 | 23 | +-------------------- Aculamprotula
|
| +-------- Nodularia
| |
--+ +--------+-------- Pseudocuneopsis
| | |
| | | +----- Schistodesmus
| | +--+
+--+ | +-- Tchangsinaia
| +--+
| +-- Cuneopsis
|
| +-------------- Unio_tumidus **
+--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Reis et al., 2013 [Publ 3580], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 8 | 0 | 0 | 19 | +----------- Unio_tumidus **
|
--+ +-- Unio_tumidiformis **
| +-----+
| | +-- Unio_crassus **
+--+
| +----- Unio_delphinus **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_pictorum **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 89 | 16 | 6 | 23 | +-------------------- Aculamprotula
|
| +-- Nodularia
| +-----+
--+ | +-- Pseudocuneopsis
| +--------+
| | | +----- Tchangsinaia
| | +--+
| | | +-- Schistodesmus
+--+ +--+
| +-- Cuneopsis
|
| +-------------- Unio_tumidus **
+--+
| +----------- Unio_nanus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Araujo et al., 2009 [Publ 2589], fig. 7A iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 7 | 3 | 7 | 5 | 1 | 1 | 22 | +----------- Nodularia
|
| +-- Unio_gibbus **
--+ |
| +-----+-- Unio_tumidus **
| | |
| | +-- Anodonta
+--+
| +----- Unio_pictorum__1 **
| |
| +----- Unio_pictorum__2 **
+--+
+----- Unio_mancus **
|
| +-- Unio_tumidiformis **
+--+
+-- Unio_crassus **
Prie et al., 2012 [Publ 3487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 1 | 0 | 0 | 18 | +----------- Unio_tumidus **
--+
| +-------- Unio_crassus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_elongatulus **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 7 | 0 | 0 | 22 | +----------- Unio_tumidus **
--+
| +-------- Unio_nanus **
+--+
| +----- Unio_elongatulus **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_pictorum **
Bashê, 2022 [Publ 4265], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 6 | 1 | 0 | 0 | 25 | +-------------- Unio_pictorum__2 **
--+
| +----------- Unio_foucauldianus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_tigridis **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_pictorum__1 **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 49 | 8 | 6 | 21 | +-------------- Aculamprotula
|
| +-------- Unio_tumidus **
| +--+
--+ | | +----- Unio_nanus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_delphinus **
|
| +-- Schistodesmus
| +-----+
| | +-- Cuneopsis
+--+
| +----- Pseudocuneopsis
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 27 | 3 | 3 | 21 | +-------------- Aculamprotula
|
--+ +-- Tchangsinaia
| +--------+
| | +-- Nodularia
+--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Soroka, 2010 [Publ 3404], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 5 | 0 | 0 | 21 | +-------- Unio_tumidus **
--+
| +----- Unio **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 69 | 11 | 6 | 21 | +----------------- Aculamprotula
|
| +-------- Unio_tumidus **
--+ +-----+
| | | +----- Unio_nanus **
| | +--+
| | | +-- Unio_pictorum **
+--+ +--+
| +-- Unio_delphinus **
|
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 34 | 6 | 4 | 21 | +-------------- Aculamprotula
|
| +----- Schistodesmus
--+ +-----+
| | | +-- Nodularia
| | +--+
+--+ +-- Tchangsinaia
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 6 | 1 | 1 | 21 | +----------- Aculamprotula
--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 36 | 7 | 5 | 21 | +-------------- Aculamprotula
|
| +-------- Nodularia
--+ +--+
| | | +----- Tchangsinaia
| | +--+
| | | +-- Cuneopsis
+--+ +--+
| +-- Schistodesmus
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Kim et al., 2021 [Publ 4197], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 13 | 7 | 4 | 21 | +----- Nodularia
+-----+
| | +-- Schistodesmus
| +--+
--+ +-- Cuneopsis
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 44 | 6 | 4 | 21 | +-------------- Aculamprotula
|
| +----- Nodularia
--+ +-----+
| | | +-- Schistodesmus
| | +--+
+--+ +-- Tchangsinaia
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 64 | 11 | 6 | 21 | +----------------- Aculamprotula
|
| +-------- Unio_tumidus **
--+ +-----+
| | | +----- Unio_nanus **
| | +--+
| | | +-- Unio_pictorum **
+--+ +--+
| +-- Unio_delphinus **
|
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 27 | 4 | 4 | 21 | +-------------- Aculamprotula
|
| +----- Nodularia
--+ +-----+
| | | +-- Tchangsinaia
| | +--+
+--+ +-- Cuneopsis
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Fukata & Iigo, 2019 [Publ 4108], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 12 | 8 | 6 | 21 | +-------- Unio_tumidus **
+--+
| | +----- Unio_nanus **
| +--+
| | +-- Unio_delphinus **
--+ +--+
| +-- Unio_pictorum **
|
| +-------- Pseudocuneopsis
| |
+--+ +-- Schistodesmus
| +--+
| | +-- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 195 | 15 | 5 | 19 | +-------- Unio_tumidus **
--+
| +----- Unio_crassus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Klishko et al., 2018 [Publ 3945], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 4 | 4 | 1 | 22 | +-------- Nodularia
|
--+ +----- Unio_tumidus **
| |
+--+----- Unio_gibbus **
|
| +-- Unio_nanus **
+--+
+-- Unio_pictorum **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 69 | 11 | 6 | 21 | +----------------- Aculamprotula
|
| +-------- Unio_tumidus **
--+ +-----+
| | | +----- Unio_nanus **
| | +--+
| | | +-- Unio_pictorum **
+--+ +--+
| +-- Unio_delphinus **
|
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 81 | 23 | 7 | 21 | +----------------- Aculamprotula
|
| +-------- Unio_tumidus **
--+ +-----+
| | | +----- Unio_nanus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +----------- Pseudocuneopsis
| |
+--+ +-------- Nodularia
| |
+--+ +-- Schistodesmus
| +--+
| | +-- Tongchuania
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 34 | 19 | 7 | 21 | +-------------------- Aculamprotula
|
| +-------- Unio_tumidus **
--+ +--------+
| | | +----- Unio_nanus **
| | +--+
| | | +-- Unio_pictorum **
+--+ +--+
| +-- Unio_delphinus **
|
| +-------------- Pseudocuneopsis
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Soroka, 2010 [Publ 3404], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 5 | 0 | 0 | 21 | +-------- Unio_tumidus **
--+
| +----- Unio **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 40 | 6 | 4 | 21 | +-------------- Aculamprotula
|
| +----- Nodularia
--+ +-----+
| | | +-- Tchangsinaia
| | +--+
+--+ +-- Schistodesmus
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 40 | 6 | 4 | 21 | +-------------- Aculamprotula
|
| +----- Nodularia
--+ +-----+
| | | +-- Tchangsinaia
| | +--+
+--+ +-- Schistodesmus
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Nagel et al., 1998 [Publ 1946], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 0 | 0 | 0 | 20 | +----- Unio_tumidus **
|
--+ +-- Unio_pictorum **
| |
+--+-- Unio_crassus **
|
+-- Unio_mancus **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 117 | 23 | 7 | 20 | +----------------- Aculamprotula
|
| +-------------- Middendorffinaia
--+ |
| | +----- Unio_gibbus **
| | |
+--+ +-----+----- Unio_tumidus **
| | |
| | | +-- Unio_crassus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-------- Nodularia
| |
+--+ +----- Schistodesmus
| |
+--+----- Cuneopsis
|
| +-- Tchangsinaia
+--+
+-- Inversiunio
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 193 | 14 | 5 | 19 | +-------------- Aculamprotula
|
| +-------- Unio_tumidus **
--+ +--+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 195 | 15 | 5 | 19 | +-------------- Aculamprotula
|
| +-------- Unio_tumidus **
--+ +--+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +-------- Schistodesmus
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 42 | 9 | 6 | 21 | +----------------- Aculamprotula
|
| +-------- Unio_tumidus **
--+ +-----+
| | | +----- Unio_nanus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 19 | 6 | 4 | 21 | +-------------- Aculamprotula
|
| +----- Schistodesmus
--+ +-----+
| | | +-- Tchangsinaia
| | +--+
+--+ +-- Nodularia
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 69 | 16 | 6 | 21 | +----------------- Aculamprotula
|
| +-------- Unio_tumidus **
--+ +-----+
| | | +----- Unio_nanus **
| | +--+
| | | +-- Unio_pictorum **
+--+ +--+
| +-- Unio_delphinus **
|
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Soroka, 2005 [Publ 2726], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 1 | 0 | 0 | 21 | +-------- Unio_tumidus **
--+
| +----- Unio_nanus **
+--+
| +-- Unio_caffer **
+--+
+-- Unio_pictorum **
Klishko et al., 2019 [Publ 3886], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 7 | 4 | 2 | 22 | +-- Middendorffinaia
+-----+
| +-- Nodularia
--+
| +----- Unio_tumidus **
| |
+--+----- Unio_gibbus **
|
| +-- Unio_nanus **
+--+
+-- Unio_pictorum **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 76 | 7 | 5 | 20 | +----- Unio_gibbus **
|
+--------+----- Unio_tumidus **
| |
| | +-- Unio_crassus **
--+ +--+
| +-- Unio_pictorum **
|
| +----------- Aculamprotula
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 45 | 10 | 6 | 21 | +----------------- Aculamprotula
|
| +-------- Unio_tumidus **
--+ +-----+
| | | +----- Unio_nanus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Nagel et al., 1998 [Publ 1946], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 5 | 0 | 0 | 21 | +-------- Unio_crassus **
--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Liu et al., 2017 [Publ 3939], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 11 | 7 | 3 | 20 | +-- Schistodesmus
|
+-----+-- Cuneopsis
| |
| +-- Nodularia
--+
| +----- Unio_tumidus **
| |
+--+----- Unio_gibbus **
|
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 38 | 5 | 3 | 19 | +-------------- Aculamprotula
|
--+ +-- Tchangsinaia
| +--------+
| | +-- Nodularia
+--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 16 | 1 | 1 | 21 | +----------- Aculamprotula
--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 52 | 7 | 4 | 21 | +-------------- Aculamprotula
|
| +----- Nodularia
--+ +-----+
| | | +-- Tchangsinaia
| | +--+
+--+ +-- Schistodesmus
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_nanus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Klishko et al., 2017 [Publ 3946], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 2 | 2 | 1 | 21 | +----------- Nodularia
--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_nanus **
+--+
+-- Unio_crassus **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 10 | 0 | 3 | 122 | 7 | 5 | 28 | +-- Unio_crassus **
|
+-- Unio_pictorum **
|
+-- Unio_tumidus **
|
+-- Aculamprotula
|
+-- Cuneopsis_szechenyii
--+
+-- Tchangsinaia
|
+-- Cuneopsis_rufescens
|
+-- Nodularia_douglasiae
|
+-- Nodularia_nuxpersicae
|
+-- Schistodesmus
Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 33 | 2 | 2 | 20 | +----------- Middendorffinaia
--+
| +-------- Nodularia
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Froufe et al., 2019 [Publ 3936], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 16 | 0 | 0 | 22 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Froufe et al., 2019 [Publ 3936], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 16 | 0 | 0 | 22 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 65 | 10 | 5 | 22 | +-------------- Aculamprotula
|
| +----- Unio_nanus **
--+ +-----+
| | | +-- Unio_delphinus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-------- Nodularia
+--+
| +----- Inversiunio
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia
Källersjö et al., 2005 [Publ 2684], fig. 1A iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 0 | 3 | 4 | 0 | 0 | 21 | +-- Unio_crassus **
|
--+-- Unio_pictorum **
|
+-- Unio_tumidus **
Kilikowska et al., 2020 [Publ 4335], fig. 4a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 0 | 0 | 0 | 22 | +----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_nanus **
Soroka, 2010 [Publ 3404], fig. 8b iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 3 | 0 | 0 | 22 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 53 | 9 | 5 | 20 | +-------------- Middendorffinaia
|
--+ +----------- Aculamprotula
| |
| | +----- Unio_tumidus **
+--+ +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_crassus **
|
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 2 | 0 | 0 | 22 | +----- Unio_pictorum **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_crassus **
Soroka, 2010 [Publ 3404], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 4 | 0 | 0 | 22 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 136 | 7 | 5 | 20 | +----- Unio_tumidus **
+-----+
| | +-- Unio_crassus **
| +--+
| +-- Unio_pictorum **
--+
| +-------- Aculamprotula
| |
| +-------- Schistodesmus
+--+
+-------- Nodularia
|
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Bogatov et al., 2018 [Publ 3859], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 5 | 3 | 3 | 3 | 0 | 0 | 22 | +-------- Unio_tumidus **
|
--+ +-- Anodonta
| +--+
| | +-- Pseudanodonta
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 15 | 0 | 0 | 20 | +----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Huang et al., 2018 [Publ 3828], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 14 | 1 | 1 | 20 | +-------- Aculamprotula
--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Huang et al., 2019 [Publ 3843], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 20 | 2 | 2 | 22 | +----------- Schistodesmus
--+
| +-------- Aculamprotula
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Zieritz et al., 2012 [Publ 3673], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 5 | 0 | 0 | 22 | +----- Unio_crassus **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Guerra et al., 2019 [Publ 4168], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 38 | 2 | 2 | 22 | +----------- Aculamprotula
--+
| +-------- Schistodesmus
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Bogatov et al., 2018 [Publ 3859], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 5 | 3 | 3 | 3 | 0 | 0 | 24 | +----------- Unio_crassus **
--+
| +-------- Unio_pictorum **
+--+
| +----- Unio_tumidus **
+--+
| +-- Anodonta
+--+
+-- Pseudanodonta
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 20 | 0 | 0 | 22 | +----- Unio_tumidus **
--+
| +-- Unio_delphinus **
+--+
+-- Unio_nanus **
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 65 | 10 | 5 | 22 | +-------------- Aculamprotula
|
| +----- Unio_nanus **
--+ +-----+
| | | +-- Unio_delphinus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-------- Inversiunio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 136 | 7 | 5 | 20 | +----- Unio_tumidus **
+--------+
| | +-- Unio_pictorum **
| +--+
--+ +-- Unio_crassus **
|
| +----------- Aculamprotula
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Soroka, 2010 [Publ 3404], fig. 3a-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 3 | 0 | 0 | 22 | +----- Unio_nanus **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Froufe et al., 2017 [Publ 3802], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 7 | 0 | 0 | 22 | +----- Unio_mancus **
--+
| +-- Unio_elongatulus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 8a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 3 | 0 | 0 | 22 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Guerra et al., 2019 [Publ 4168], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 38 | 2 | 2 | 22 | +-------- Schistodesmus
--+
| +----- Unio_tumidus **
+--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 65 | 10 | 5 | 22 | +-------------- Aculamprotula
|
| +----- Unio_nanus **
--+ +-----+
| | | +-- Unio_delphinus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-------- Inversiunio
+--+
| +----- Nodularia
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia
Soroka, 2010 [Publ 3404], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 0 | 3 | 4 | 0 | 0 | 22 | +-- Unio_tumidus **
|
--+-- Unio_pictorum **
|
+-- Unio_nanus **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 4 | 0 | 0 | 20 | +----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 80 | 2 | 1 | 20 | +-------- Nodularia
--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Manendo et al., 2008 [Publ 2669], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 10 | 0 | 0 | 22 | +----- Unio_crassus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_tumidus **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 73 | 7 | 5 | 20 | +----- Unio_tumidus **
+-----------+
| | +-- Unio_pictorum **
| +--+
--+ +-- Unio_crassus **
|
| +-------------- Aculamprotula
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Soroka, 2010 [Publ 3404], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 5 | 0 | 0 | 22 | +----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Guerra et al., 2019 [Publ 4168], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 19 | 2 | 2 | 22 | +----------- Schistodesmus
--+
| +-------- Aculamprotula
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Soroka, 2010 [Publ 3404], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 5 | 0 | 0 | 22 | +----- Unio_nanus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_tumidus **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 73 | 7 | 5 | 20 | +----- Unio_tumidus **
+-----------+
| | +-- Unio_pictorum **
| +--+
--+ +-- Unio_crassus **
|
| +-------------- Aculamprotula
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Guerra et al., 2019 [Publ 4168], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 19 | 2 | 2 | 22 | +----------- Aculamprotula
--+
| +-------- Schistodesmus
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_nanus **
+--+
+-- Unio_delphinus **
Soroka, 2010 [Publ 3404], fig. 3b-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 3 | 0 | 0 | 22 | +----- Unio_tumidus **
--+
| +-- Unio_nanus **
+--+
+-- Unio_pictorum **
Wu et al., 2022 [Publ 4238], fig. 2e iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 14 | 9 | 6 | 22 | +----------------- Aculamprotula
|
| +----- Unio_tumidus **
--+ +--------+
| | | +-- Unio_nanus **
| | +--+
+--+ +-- Unio_delphinus **
|
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Soroka, 2010 [Publ 3404], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 2 | 0 | 0 | 22 | +----- Unio_nanus **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Sonowal et al., 2021 [Publ 4221], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 9 | 0 | 2 | 13 | 1 | 1 | 28 | +-- Aculamprotula
|
+-- Lamprotula
|
+-- Lampsilis
|
+-- Potomida
|
--+-- Unio_pictorum **
|
+-- Anodonta
|
+-- Unio_ravoisieri **
|
+-- Lamellidens
|
+-- Elliptio
Hoeh et al., 2001 [Publ 2652], fig. 14f4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 5 | 0 | 2 | 29 | 0 | 0 | 24 | +-------- Unio_pictorum **
|
--+-------- Unio_tumidus **
|
| +----- Anodonta
+--+
| +-- Strophitus
+--+
+-- Pyganodon
Chase et al., 2018 [Publ 3861], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 9 | 0 | 0 | -- | +-- Unio_delphinus **
--+
+-- Unio_tumidus **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 30 | 5 | 4 | -- | +----------- Aculamprotula
|
--+ +-- Unio_delphinus **
| +-----+
| | +-- Unio_pictorum **
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Chase et al., 2018 [Publ 3847], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 36 | 0 | 0 | -- | +-- Unio_delphinus **
--+
+-- Unio_tumidus **
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 44 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Hoeh et al., 2002 [Publ 2653], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 5 | 0 | 2 | 29 | 0 | 0 | 24 | +----------- Unio_pictorum **
--+
| +-------- Unio_tumidus **
+--+
| +----- Anodonta
+--+
| +-- Pyganodon
+--+
+-- Strophitus
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 38 | 15 | 9 | -- | +-------------------- Aculamprotula
|
| +-- Middendorffinaia
--+ +--------------+
| | +-- Nodularia
| |
+--+ +-------------- Cuneopsis
| |
| | +-- Diaurora
+--+-----------+
| +-- Schistodesmus
|
| +----------- Pseudocuneopsis
+--+
| +-------- Inversiunio
+--+
| +----- Tchangsinaia
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Sonowal et al., 2021 [Publ 4221], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 4 | 0 | 2 | 14 | 1 | 1 | 23 | +----------- Aculamprotula
--+
| +-------- Unio_pictorum **
+--+
| +----- Anodonta
+--+
| +-- Unio_ravoisieri **
+--+
+-- Lamellidens
Bolotov et al., 2016 [Publ 3743], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_ravoisieri **
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 20 | 10 | -- | +----------------------- Globunio
|
--+ +-------------------- Aculamprotula
| |
| | +-- Middendorffinaia
+--+ +--------------+
| | +-- Nodularia
| |
+--+ +-------------- Cuneopsis
| |
| | +-- Diaurora
+--+-----------+
| +-- Schistodesmus
|
| +----------- Pseudocuneopsis
+--+
| +-------- Inversiunio
+--+
| +----- Tchangsinaia
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 45 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Riccardi et al., 2020 [Publ 4089], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 17 | 0 | 0 | -- | +-- Unio_mancus **
--+
+-- Unio_elongatulus **
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 51 | 12 | 5 | -- | +----------- Aculamprotula
|
| +-- Unio_crassus **
--+ +-----+
| | +-- Unio_pictorum **
| |
+--+ +-- Nodularia
| +--+
| | +-- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 7 | 0 | 3 | 23 | 1 | 1 | 27 | +-------------- Unio_pictorum__2 **
|
--+ +-- Unio_pictorum__1 **
| +--------+
| | +-- Unio_caffer **
+--+
| +-------- Anodonta
| |
+--+ +----- Lasmigona
| |
+--+ +-- Strophitus
| |
+--+-- Alasmidonta
|
+-- Pyganodon
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 38 | 15 | 9 | -- | +-------------------- Aculamprotula
|
| +-- Middendorffinaia
--+ +--------------+
| | +-- Nodularia
| |
+--+ +-------------- Cuneopsis
| |
| | +-- Diaurora
+--+-----------+
| +-- Schistodesmus
|
| +----------- Pseudocuneopsis
+--+
| +-------- Inversiunio
+--+
| +----- Tchangsinaia
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_nanus **
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 50 | 15 | 6 | -- | +----------------- Aculamprotula
|
--+ +-- Unio_nanus **
| +-----------+
| | +-- Unio_delphinus **
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 44 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 16 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Chase et al., 2018 [Publ 3861], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 9 | 0 | 0 | -- | +-- Unio_delphinus **
--+
+-- Unio_tumidus **
Soroka, 2010 [Publ 3404], fig. 8a-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 1 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Sonowal et al., 2021 [Publ 4221], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 4 | 0 | 2 | 13 | 1 | 1 | 23 | +-- Anodonta
|
+-- Unio_ravoisieri **
--+
+-- Unio_pictorum **
|
+-- Aculamprotula
Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Burzynski et al., 2017 [Publ 3774], fig. 1-top iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 1 | 1 | -- | +----- Aculamprotula
--+
| +-- Unio_nanus **
+--+
+-- Unio_tumidus **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Soroka, 2010 [Publ 3404], fig. 8b-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 1 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 5 | 4 | -- | +----------- Aculamprotula
|
--+ +-- Unio_pictorum **
| +-----+
| | +-- Unio_delphinus **
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Tomilova et al., 2020 [Publ 4097], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 6 | 0 | 0 | -- | +-- Unio_tumidus **
--+
+-- Unio_pictorum **
Bolotov et al., 2016 [Publ 3743], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_ravoisieri **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 32 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Wu et al., 2025 [Publ 4442], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 15 | 0 | 0 | -- | +-- Unio_crassus **
--+
+-- Unio_delphinus **
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 89 | 9 | 5 | -- | +-------------- Aculamprotula
|
--+ +-- Unio_pictorum **
| +--------+
| | +-- Unio_tumidus **
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Sayenko et al., 2021 [Publ 4189], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 11 | 3 | -- | +-- Unio_pictorum **
+-----+
| +-- Unio_nanus **
--+
| +----- Inversiunio
+--+
| +-- Middendorffinaia
+--+
+-- Nodularia
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 1 | 1 | -- | +----- Aculamprotula
--+
| +-- Unio_nanus **
+--+
+-- Unio_tumidus **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 89 | 9 | 5 | -- | +-------------- Aculamprotula
|
--+ +-- Unio_pictorum **
| +--------+
| | +-- Unio_tumidus **
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Hoeh et al., 2001 [Publ 2652], fig. 14f2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_tumidus **
Chase et al., 2018 [Publ 3847], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 8 | 0 | 0 | -- | +-- Unio_delphinus **
--+
+-- Unio_tumidus **
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 45 | 0 | 0 | -- | +-- Unio_pictorum **
--+
+-- Unio_caffer **
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 0 | 0 | -- | Wu et al., 2018 [Publ 4143], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 5 | 2 | -- | +----- Aculamprotula
--+
| +-- Unio_pictorum **
+--+
+-- Nodularia
Lopes-Lima et al., 2018 [Publ 3829], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- | Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 4 | 2 | -- | +----- Aculamprotula
--+
| +-- Unio_pictorum **
+--+
+-- Nodularia
Fonseca et al., 2016 [Publ 3775], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Marshall et al., 2014 [Publ 3593], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 3 | 3 | -- | +-------- Aculamprotula
--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Unio_pictorum **
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 8 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio_tumidus **
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Combosch et al., 2017 [Publ 3806], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- | Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 0 | 0 | -- | Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 13 | 7 | -- | +----------------- Unio_tumidus **
|
--+ +-------------- Aculamprotula
| |
+--+ +----------- Cuneopsis
| |
+--+ +-- Nodularia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Soroka & Burzynski, 2018 [Publ 3762], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- | +-- Unio_tumidus **
--+
+-- Aculamprotula
Araujo et al., 2009 [Publ 2395], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- | Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 9 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio_pictorum **
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Guerra et al., 2017 [Publ 3772], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 1 | 1 | -- | Wang et al., 2016 [Publ 3714], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | Graf, 2000 [Publ 2647], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- | Combosch et al., 2017 [Publ 3806], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | Dai et al., 2024 [Publ 4386], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 9 | 5 | -- | Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 2 | 2 | -- | +----- Unio_gibbus **
--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Huff et al., 2004 [Publ 2697], fig. 5-6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- | Graf et al., 2015 [Publ 3619], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | González & Giribet, 2015 [Publ 3632], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Wu et al., 2020 [Publ 4147], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 4 | 1 | -- | +-- Unio_pictorum **
--+
+-- Aculamprotula
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- | Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 12 | 7 | -- | +----------------- Unio_tumidus **
|
--+ +-------------- Aculamprotula
| |
+--+ +----------- Cuneopsis
| |
+--+ +-- Nodularia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Wu et al., 2020 [Publ 4121], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 4 | 1 | -- | +-- Unio_pictorum **
--+
+-- Aculamprotula
Huang et al., 2013 [Publ 3629], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 0 | 0 | -- | Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- | Bolotov et al., 2015 [Publ 3646], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Taylor et al., 2007 [Publ 2696], fig. 1-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | Lopes-Lima et al., 2018 [Publ 3829], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 0 | 0 | -- | Bieler et al., 2014 [Publ 3622], fig. 28 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Graf et al., 2014 [Publ 3591], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- | +-- Unio_tumidus **
--+
+-- Aculamprotula
Wu et al., 2016 [Publ 3711], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 4 | 2 | -- | +----- Aculamprotula
--+
| +-- Unio_pictorum **
+--+
+-- Nodularia
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 13 | 7 | -- | +-------- Unio_tumidus **
|
+-------- Aculamprotula
|
--+ +----- Cuneopsis
| |
| | +-- Nodularia
| +--+
+--+ +-- Schistodesmus
|
| +-- Tchangsinaia
| |
+--+-- Inversiunio
|
+-- Pseudocuneopsis
Wen et al., 2017 [Publ 3795], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 2 | 1 | -- | Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 2 | 2 | -- | +----- Unio_gibbus **
--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Nagel et al., 1998 [Publ 1946], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Bolotov et al., 2015 [Publ 3646], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Vikhrev et al., 2018 [Publ 3812], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 0 | 0 | -- | Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 6 | 6 | -- | +-------------- Aculamprotula
|
--+ +----------- Unio_nanus **
| |
+--+ +-- Pseudocuneopsis
| +-----+
| | +-- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Choi et al., 2020 [Publ 4169], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 11 | 4 | -- | +----------- Nodularia
--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Unio_pictorum **
+--+
+-- Tchangsinaia
González & Giribet, 2015 [Publ 3632], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Huang et al., 2018 [Publ 3828], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 1 | 1 | -- | +-- Unio_pictorum **
--+
+-- Aculamprotula
Araujo et al., 2017 [Publ 3801], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 2 | 2 | -- | Guerra et al., 2017 [Publ 3772], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 1 | 1 | -- | +-- Aculamprotula
--+
+-- Unio_pictorum **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 5 | 4 | -- | +-------- Unio_pictorum **
--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 10 | 6 | -- | +----------- Unio_crassus **
|
--+ +-------- Middendorffinaia
| |
+--+ +----- Cuneopsis
| |
+--+----- Nodularia
|
| +-- Inversiunio
+--+
+-- Schistodesmus
Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 10 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio_crassus **
+--+
| +----------- Middendorffinaia
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Inversiunio
+--+
+-- Schistodesmus
Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 0 | 0 | -- | González & Giribet, 2015 [Publ 3632], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Chong et al., 2008 [Publ 2668], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 0 | 0 | -- | González & Giribet, 2015 [Publ 3632], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Choi et al., 2020 [Publ 4169], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 5 | 2 | -- | +----- Nodularia
--+
| +-- Cuneopsis
+--+
+-- Unio_pictorum **
Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Klunzinger et al., 2021 [Publ 4086], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- | Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- | +-- Schistodesmus
--+
+-- Unio_pictorum **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 5 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio_pictorum **
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Giribet & Distel, 2003 [Publ 3670], fig. 3f5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Wu et al., 2020 [Publ 4121], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 4 | 1 | -- | +-- Unio_pictorum **
--+
+-- Aculamprotula
|
