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Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
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Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Mon 2 March 2026
Bivalvia | ... | Unionidae | Unioninae | Unionini | Oxynaiina | Nodularia douglasiae
nominal species Unio douglasiae Griffith & Pidgeon, 1833
Unio douglasiae Griffith & Pidgeon, 1833 = Nodularia douglasiae
Griffith & Pidgeon, 1833. [in] Cuvier, Animal Kingdom 12(38): pl. 21, fig. 2 [as ‘douglasiæ’].
‘Gray’ Griffith & Pidgeon, 1834. [in] Cuvier, Animal Kingdom 12(39-40): 601 [as ‘douglasiæ’].
Taxonomic Opinions
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Campbell & Lydeard, 2026: 294.
= Nodularia douglasiae — Hou et al., 2025: Tables S1-S2.
= Nodularia douglasiae — Hou et al., 2025: Table S2.
= Nodularia douglasiae (Gray, 1833) — Dai et al., 2025: Tables 1-2.
= Nodularia douglasiae (Gray, 1833) — Chen et al., 2025: Tables S1-S3.
= Nodularia douglasiae (Gray, 1833) — Chen et al., 2025: Tables S2.
= Nodularia douglasiae — Hou et al., 2025: Table S3.
= Nodularia douglasiae — Wang et al., 2025: Table S2.
= Nodularia douglasiae (Gray, 1833) — Chen et al., 2025: Table 1.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Chen et al., 2025: 965.
= Nodularia douglasiae — Wu et al., 2024: 6.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Dai et al., 2024: Tables 1, S2.
= Nodularia douglasiae — Lopes-Lima et al., 2024: figs. 4-5.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Wu et al., 2024: fig. 3a, suppl. material, p. 4, figs. S1, S9-S11a, Tables S1-S3.
= Nodularia douglasiae — Lopes-Lima et al., 2024: figs. 4-5.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Dai et al., 2024: Tables 2-3, S1.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Dai et al., 2024: Tables 2-3, S1.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Dai et al., 2024: Tables 2-3, S1.
= Nodularia sp. 2 — Dai et al., 2024: Table 2, S1.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Dai et al., 2024: Tables 2, S1.
= Nodularia douglasiae — Wu et al., 2024: Tables S1, S3.
= Nodularia douglasiae — Wu et al., 2024: Table S1.
= Nodularia douglasiae — Hou et al., 2024: Table 2.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Konopleva et al., 2023: Table S1.
= Nodularia douglasiae (Griffith & Pidge) [sic] — Dai et al., 2023: 157, Table 2.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Wu et al., 2023: 263, Table 1.
= Nodularia douglasiae (Gray, 1833) — Chen et al., 2023: 133, Table 1.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Liu et al., 2023: Table 1.
= Nodularia douglasiae — Wu et al., 2023: Table 2.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Wu et al., 2022: Table S2.
= Nodularia douglasiae — Liu et al., 2022: Table 1.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Jeratthitikul et al., 2022: Table S1.
= Nodularia douglasiae (Gray, 1833) — Pavan-Kumar et al., 2022: Table S1.
= Nodularia douglasiae douglasiae (Griffith et Pidgeon, 1833) — Bogatov, 2022: 59, 50, 57, pl. 26, figs. 15-20.
= Nodularia douglasiae (Griffith & Pidgeon) — Hata et al., 2022: 2636, Table S1.
= Nodularia douglasiae — Sayenko et al., 2021: 58.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Graf & Cummings, 2021: 14.
= Nodularia douglasiae — Kim et al., 2021: 80.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Wu et al., 2021: 87.
= Nodularia douglasiae — Lopes-Lima et al., 2021: Table S1.
= Nodularia douglasiae — Zieritz et al., 2021: 1092.
= Nodularia douglasiae — Mabuchi et al., 2021: 1536.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Wu et al., 2021: 1799, fig. 7G, attached Tables 1-2.
= Nodularia douglasiae — Sayenko et al., 2021: 58, Table 1.
= Nodularia douglasiae — Mabuchi et al., 2021: 1537.
= Nodularia douglasiae — Kim et al., 2021: 80.
= Nodularia douglasiae — Kim et al., 2021: 80.
= Nodularia douglasiae — Sayenko et al., 2021: 58, Table 1.
= Nodularia douglasiae — Mabuchi et al., 2021: 1537.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Lopes-Lima et al., 2020: 14, 23, Tables 1, 3, Appendix 1.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Bolotov et al., 2020: 4, S33, figs. 5E-H.
= Nodularia douglasiae — Liu et al., 2020: [6].
= Nodularia douglasiae — Fukata & Iigo, 2020: 1216.
= Nodularia douglasiae — Wu et al., 2020: 196.
= Nodularia douglasiae — Sano et al., 2020: 99.
= Nodularia douglasiae (Gray in Griffith & Pidgeon, 1833) — Wu et al., 2020: 54.
= Nodularia douglasiae — Sano et al., 2020: 99.
= Nodularia douglasiae Clade C — Choi et al., 2020: 5.
= Nodularia douglasiae Clade B — Choi et al., 2020: 5.
= Nodularia douglasiae Clade A — Choi et al., 2020: 5.
= Nodularia douglasiae Clade A — Choi et al., 2020: 5.
= Nodularia douglasiae Clade B — Choi et al., 2020: 6, fig. 3a.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Huang et al., 2019: Tables S1, S4.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Wu et al., 2019: 29.
= Nodularia douglasiae (Griffith et Pidgeon, 1833) — Klishko et al., 2019: 57, Table 1.
= Nodularia douglasiae — Froufe et al., 2019: 185.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Pfeiffer et al., 2019: 117.
= Nodularia douglasiae — Fukata & Iigo, 2019: 3488.
= Nodularia douglasiae — Fukata & Iigo, 2019: 3488.
= Nodularia douglasiae — Fukata & Iigo, 2019: 3488.
= Nodularia douglasiae (Griffith et Pidgeon, 1833) — Klishko et al., 2019: 67, fig. 9.
= Nodularia cf. douglasiae (Griffith & Pidgeon, 1833) — Huang et al., 2019: Tables S1, S4.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Zieritz et al., 2018: online Table 1.
= Unio douglasiae — Araujo et al., 2018: [5].
= Nodularia douglasiae (Gray, 1833) — Do et al., 2018: 10, Table 2.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Bolotov et al., 2018: Table S1.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Pfeiffer et al., 2018: [7].
= Nodularia douglasiae — Huang et al., 2018: 304.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Wu et al., 2018: 6.
= Unio douglasiae — Wang et al., 2018: 611.
= Unio douglasiae — Aunins et al., 2018: 1125.
= Unio douglasiae — Wang et al., 2018: 140.
= Nodularia douglasiae — Cha et al., 2018: 946.
= Nodularia douglasiae — Klishko et al., 2018: 291, figs. 2.16-2.23, 3.10-3.16, 3.19, 4.6-4.8, 4.14-4.16.
= Nodularia douglasiae — Wu et al., 2018: 732.
= Nodularia douglasiae — Cha et al., 2018: 946.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Lopes-Lima et al., 2017: 177.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Bolotov et al., 2017: [14], Table S1.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Bolotov et al., 2017: Table S1.
= Unio douglasiae — Zhou et al., 2017: 235.
= Nodularia douglasiae — Lopes-Lima et al., 2017: 250.
= Nodularia douglasiae Gray in Griffith & Pidgeon, 1833 — Sano et al., 2017: 494.
= Nodularia douglasiae — Soroka & Burzynski, 2017: 863.
= Nodularia douglasiae (Griffith et Pidgeon, 1833) — Bogatov & Prozorova, 2017: 931.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Liu et al., 2017: 2.
= Nodularia douglasiae — Klishko et al., 2017: 99.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Liu et al., 2017: 2.
= Nodularia douglasiae (Griffith & Pidgeon, 1833) — Liu et al., 2017: 2.
= Unio douglasiae — Xue et al., 2016: 2699.
= Unio douglasiae — Wu et al., 2016: 4521.
= Nodularia douglasiae — Froufe et al., 2016: 606.
= Nodularia douglasiae — Fonseca et al., 2016: 955.
= Unio douglasiae — Soliman et al., 2016: 122.
= Unio douglasiae — Wang et al., 2015: 4021.
= Unio douglasiae — Ouyang et al., 2015: Table 1.
= Unio douglasiae — Ouyang et al., 2015: Table 1.
= Unio douglasiae (Gray) — IUCN, 2014: [web].
= Nodularia douglasiae (Gray, 1833) — Pfeiffer & Graf, 2013: 251.
= Nodularidia douglasiae (Griffith & Pidgeon, 1833) — He & Zhuang, 2013: 82, figs. 185-204.
= Nodularia douglasiae (Griffith & Pidgeon, 1834) — Schneider et al., 2013: 341, fig. 2A.
= Unio douglasiae — Xiong et al., 2012: 128, Table 2.
= Unio douglasiae — Ouyang et al., 2011: 544.
= Nodularia douglasiae Griffith & Pidgeon, 1834 — Bogan, 2010: [23].
= Unio douglasiae — Jadhav & Jamkhedkar, 2009: 39.
= Unio douglasiae Gray in Griffith & Pidgeon, 1833 — Araujo et al., 2009: 132.
+ — Petit & Coan, 2008: 229.
= Unio douglasiae Griffith & Pidgeon, 1834 — Kondo, 2008: 18.
= Nodularia douglasiae (Griffith & Pidgeon, 1834) — Graf, 2007: 85.
= Nodularia douglasiae (Griffith & Pidgeon, 1834) — Graf & Cummings, 2007: 304.
= Unio douglasiae — Zhou et al., 2007: 1025.
= Nodularia douglasiae (Griffith et Pidgeon, 1843) — Prozorova et al., 2005: 53.
= Unio douglasiae — Zhang et al., 2004: 295.
= Unio douglasiae Griffith & Pidgeon, 1834 — Huang et al., 2002: 360.
= Unio douglasiae Griffit at Pitaeon, 1833 [sic] — Bába, 2000: 133, fig. 3.
= Unio douglasiae — Wu et al., 1999: 140.
= Unio douglasiae Gray — Wu et al., 1999: 141.
= Nodularia (s.s.) douglasiae (Griffith & Pidgeon, 1834) — Korniushin, 1998: 78.
= Unio douglasiae Gray — Wei et al., 1994: 304.
= Unio douglasiae — Park & Kwon, 1993: 46.
= Unio douglasiae Gray in Griffith & Pidgeon, 1834 — Habe, 1991: 3.
= Unio douglasiae (Griffith & Pidgeon, 1834) — Wu, 1982: 147, pls. 1-2.
= Unio douglasiae Griffith et Pidgeon — Liu et al., 1980: 366.
= Unio douglasiae Gray in Griffith & Pidgeon, 1834 — Liu et al., 1979: 74, fig. 65.
= Unio douglasiae Griffith & Pidgeon, 1834 — Habe, 1977: 114, pl. 21, fig. 2.
= Unio douglasiae — Kil, 1976: 29.
= Unio douglasiae douglasiae Gray in Griffith & Pidgeon, 1834 — Haas, 1969: 56.
= Unio douglasiae Griffith & Pidgeon, 1833 — Zhadin, 1965: 245, fig. 262.
= Unio (Nodularia) douglasiae Gray in Griffith & Pidgeon, 1834 — Zhadin, 1938: 98, sp. 10, fig. 30.
= Nodularia (s.s.) douglasiae (Griffith & Pidgeon, 1834) — Haas, 1924: 190.
= Nodularia douglasiae (Griff. & Pidg., 1834) — Haas, 1920: 149.
= Nodularia (s.s.) douglasiae (Gray in Griffith & Pidgeon, 1834) — Simpson, 1914: 959.
= Nodularia (s.s.) douglasiae (Gray) — Preston, 1912: 288.
= Nodularia (s.s.) douglasiae douglasiae (Griffith & Pidgeon, 1834) — Haas, 1911: 68, pl. 6, figs. 1-10.
= Unio (Nodularia) douglasiae Gray — Martens, 1905: 55, pl. 3, fig. 1.
= Nodularia (s.s.) douglasiae (Gray in Griffith & Pidgeon, 1834) — Simpson, 1900: 808.
= Unio douglasiae Gray — Neumayr, 1899: 646.
= Unio douglasiae Gr. — Heude, 1885: pls. 65-65bis, figs. 128a-i.
+ Margaron (Unio) murchisonianus (Lea) — Lea, 1870: 32.
+ Unio osbecki Philippi, 1844 — Sowerby, 1868: pl. 69, sp. 353.
= Unio douglasiae Gray in Griffith & Pidgeon, 1834 — Sowerby, 1866: pl. 38, sp. 207, errata.
= Nodularia douglasiæ (Gray) — Conrad, 1853: 267, 268.
+ Margaron (Unio) murchisonianus (Lea) — Lea, 1852: 21.
+ Unio murchisonianus Lea — Hanley, 1843: 177.
+ Margarita (Unio) murchisonianus Lea — Lea, 1838: 14.
+ Margarita (Unio) merchisonianus [sic] (Lea, 1834) — Lea, 1836: 14.
