Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Oxynaiina

genus Nodularia Conrad, 1853: cladomics

15 Recent valid species (18 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 15 (18 total)
analyzed in 136 cladograms in 82 publications
maximum number of terminals in an analysis = 11 (mean = 2.68, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 76
maximum number of included terminals = 11, (mean = 4.01 median = 3)
proportion of cladograms that support basic monophyly = 0.99

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Nodularia cladogram distances
PCoA Ordination

NJ Tree of Nodularia cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 11 11 10 11 622 21 6 **
Cladomics cladogram Tchangsinaia Inversiunio Pseudocuneopsis Diaurora Cuneopsis Schistodesmus Nodularia dorri 2 Nodularia dorri 1 Nodularia hanensis Nodularia micheloti Nodularia breviconcha Nodularia guiensis Nodularia nuxpersicae Nodularia fusiformans Nodularia huana Nodularia nipponensis Nodularia douglasiae Nodularia dualobtusa

Hou et al., 2025 [Publ 4401], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 11 17 14 4 9.0
Cladomics cladogram Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis Nodularia dorri Nodularia hanensis Nodularia micheloti Nodularia breviconcha Nodularia huana Nodularia fusiformans Nodularia guiensis Nodularia douglasiae Nodularia dualobtusa Nodularia nipponensis Nodularia nuxpersicae

Hou et al., 2025 [Publ 4401], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 11 0 0 0 11.0
Cladomics cladogram Nodularia dorri Nodularia hanensis Nodularia micheloti Nodularia guiensis Nodularia douglasiae Nodularia dualobtusa Nodularia nipponensis Nodularia nuxpersicae Nodularia huana Nodularia breviconcha Nodularia fusiformans

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 10 43 9 4 8.0
Cladomics cladogram Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis Nodularia dorri Nodularia hanensis Nodularia micheloti Nodularia breviconcha Nodularia huana Nodularia fusiformans Nodularia douglasiae Nodularia dualobtusa Nodularia nipponensis Nodularia nuxpersicae

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 10 65 14 4 8.0
Cladomics cladogram Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis Nodularia dorri Nodularia hanensis Nodularia micheloti Nodularia breviconcha Nodularia huana Nodularia fusiformans Nodularia douglasiae Nodularia dualobtusa Nodularia nipponensis Nodularia nuxpersicae

Hou et al., 2025 [Publ 4432], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 10 18 15 4 8.0
Cladomics cladogram Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis Nodularia dorri Nodularia hanensis Nodularia micheloti Nodularia breviconcha Nodularia huana Nodularia fusiformans Nodularia douglasiae Nodularia dualobtusa Nodularia nipponensis Nodularia nuxpersicae

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 10 42 12 4 8.0
Cladomics cladogram Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis Nodularia dorri Nodularia micheloti Nodularia hanensis Nodularia breviconcha Nodularia huana Nodularia fusiformans Nodularia nipponensis Nodularia nuxpersicae Nodularia douglasiae Nodularia dualobtusa

Wu et al., 2024 [Publ 4379], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 10 3 3 3 6.0
Cladomics cladogram Cuneopsis Tchangsinaia Pseudocuneopsis Nodularia dorri Nodularia hanensis Nodularia micheloti Nodularia breviconcha Nodularia fusiformans Nodularia huana Nodularia douglasiae Nodularia dualobtusa Nodularia nuxpersicae Nodularia nipponensis

Wu et al., 2024 [Publ 4379], fig. 4_6

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 10 13 2 1 8.0
Cladomics cladogram Schistodesmus Nodularia dorri Nodularia micheloti Nodularia hanensis Nodularia breviconcha Nodularia huana Nodularia fusiformans Nodularia douglasiae Nodularia dualobtusa Nodularia nuxpersicae Nodularia nipponensis

Choi et al., 2020 [Publ 4169], fig. 2a

cst iT tT bi riS oT roT roS iD
0.800 7 7 6 7 5 4 3 12.0
Cladomics cladogram Schistodesmus Cuneopsis Unio Tchangsinaia Nodularia breviconcha Nodularia huana Nodularia fusiformans Nodularia nipponensis Nodularia nuxpersicae Nodularia dualobtusa 2 Nodularia douglasiae Nodularia dualobtusa 1

Wu et al., 2026 [Publ 4852], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 7 13 4 2 9.0
Cladomics cladogram Pseudocuneopsis Schistodesmus Nodularia dorri Nodularia liuovatus Nodularia hanensis Nodularia breviconcha Nodularia douglasiae Nodularia huana Nodularia nuxpersicae

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 7 3 0 0 9.0
Cladomics cladogram Middendorffinaia Nodularia breviconcha Nodularia fusiformans Nodularia huana Nodularia nuxpersicae Nodularia nipponensis Nodularia douglasiae Nodularia dualobtusa

Choi et al., 2020 [Publ 4169], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 7 3 0 0 9.0
Cladomics cladogram Radiatula Lamellidens Leoparreysia Nodularia breviconcha Nodularia fusiformans Nodularia huana Nodularia dualobtusa Nodularia douglasiae Nodularia nipponensis Nodularia nuxpersicae

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 116 12 4 8.0
Cladomics cladogram Schistodesmus Cuneopsis Tchangsinaia Inversiunio Nodularia breviconcha Nodularia huana Nodularia nuxpersicae Nodularia nipponensis Nodularia douglasiae Nodularia dualobtusa

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 16 3 3 12.0
Cladomics cladogram Cuneopsis Inversiunio Schistodesmus Nodularia huana Nodularia breviconcha Nodularia nuxpersicae Nodularia nipponensis Nodularia douglasiae Nodularia dualobtusa

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
1.000 6 6 3 6 16 3 3 11.0
Cladomics cladogram Cuneopsis Inversiunio Schistodesmus Nodularia nuxpersicae Nodularia nipponensis Nodularia huana Nodularia breviconcha Nodularia douglasiae Nodularia dualobtusa

Dai et al., 2024 [Publ 4386], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 8 2 2 8.0
Cladomics cladogram Inversiunio Cuneopsis Nodularia breviconcha Nodularia fusiformans Nodularia huana Nodularia nuxpersicae Nodularia douglasiae Nodularia nipponensis

Sayenko et al., 2021 [Publ 4189], fig. 6

cst iT tT bi riS oT roT roS iD
0.667 6 6 4 6 7 4 1 8.0
Cladomics cladogram Middendorffinaia Nodularia breviconcha Nodularia huana Nodularia nuxpersicae Nodularia nipponensis Nodularia douglasiae Nodularia dualobtusa

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 36 6 4 11.0
Cladomics cladogram Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis Nodularia breviconcha Nodularia fusiformans Nodularia douglasiae Nodularia nuxpersicae Nodularia nipponensis

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 135 16 4 9.0
Cladomics cladogram Tchangsinaia Cuneopsis Pseudocuneopsis Schistodesmus Nodularia breviconcha Nodularia fusiformans Nodularia nuxpersicae Nodularia douglasiae Nodularia dualobtusa

Wu et al., 2024 [Publ 4388], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 4 0 0 9.0
Cladomics cladogram Cosmopseudodon Ptychorhynchus Postolata Obovalis Nodularia breviconcha Nodularia douglasiae Nodularia dualobtusa Nodularia nipponensis Nodularia nuxpersicae

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 83 12 4 10.0
Cladomics cladogram Schistodesmus Tchangsinaia Cuneopsis Pseudocuneopsis Nodularia breviconcha Nodularia fusiformans Nodularia nuxpersicae Nodularia douglasiae

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 60 19 4 10.0
Cladomics cladogram Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis Nodularia hanensis Nodularia fusiformans Nodularia douglasiae Nodularia guiensis

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 91 16 4 10.0
Cladomics cladogram Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis Nodularia hanensis Nodularia fusiformans Nodularia douglasiae Nodularia guiensis

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 61 16 4 10.0
Cladomics cladogram Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis Nodularia hanensis Nodularia fusiformans Nodularia douglasiae Nodularia guiensis

Liu et al., 2017 [Publ 3939], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 3 4 11 3 2 10.0
Cladomics cladogram Schistodesmus Cuneopsis Nodularia breviconcha Nodularia douglasiae 3 Nodularia douglasiae 2 Nodularia douglasiae 1 Nodularia nuxpersicae Nodularia nipponensis

Kim et al., 2021 [Publ 4197], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 14 3 2 8.0
Cladomics cladogram Cuneopsis Schistodesmus Nodularia breviconcha Nodularia nipponensis Nodularia dualobtusa Nodularia douglasiae

Choi et al., 2020 [Publ 4169], fig. 3a

cst iT tT bi riS oT roT roS iD
0.667 4 4 3 4 2 1 1 11.0
Cladomics cladogram Cuneopsis Unio Nodularia breviconcha Nodularia nipponensis 3 Nodularia douglasiae 3 Nodularia dualobtusa Nodularia nipponensis 1 Nodularia nipponensis 2 Nodularia douglasiae 1 Nodularia douglasiae 2

Mabuchi et al., 2021 [Publ 4233], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 1 1 1 12.0
Cladomics cladogram Tchangsinaia Nodularia Nodularia nipponensis Nodularia fusiformans Nodularia breviconcha

Klishko et al., 2018 [Publ 3945], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 4 0 0 8.0
Cladomics cladogram Unio Nodularia breviconcha Nodularia huana Nodularia douglasiae Nodularia nipponensis

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 82 16 5 9.0
Cladomics cladogram Pseudocuneopsis Nodularia breviconcha Nodularia douglasiae Nodularia nipponensis Tongchuania Schistodesmus Tchangsinaia Cuneopsis

Fukata & Iigo, 2019 [Publ 4108], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 14 5 4 9.0
Cladomics cladogram Pseudocuneopsis Cuneopsis Schistodesmus Tchangsinaia Nodularia nipponensis Nodularia douglasiae Nodularia dualobtusa

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 50 4 4 9.0
Cladomics cladogram Pseudocuneopsis Nodularia breviconcha Nodularia douglasiae Nodularia nipponensis Tchangsinaia Cuneopsis Schistodesmus

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 3 3 2 3 30 5 3 10.0
Cladomics cladogram Tchangsinaia Nodularia nipponensis 1 Nodularia breviconcha Nodularia nipponensis 2 Nodularia douglasiae

Huang et al., 2019 [Publ 3843], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 3 3 1 3 43 9 3 11.0
Cladomics cladogram Schistodesmus Nodularia douglasiae Nodularia huana Nodularia nuxpersicae Cuneopsis rufescens Tchangsinaia Cuneopsis celtiformis Cuneopsis szechenyii Cuneopsis heudei

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 198 9 3 11.0
Cladomics cladogram Schistodesmus Tchangsinaia Cuneopsis Nodularia douglasiae Nodularia huana Nodularia nuxpersicae

Huang et al., 2019 [Publ 3843], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 44 9 3 11.0
Cladomics cladogram Nodularia huana Nodularia nuxpersicae Nodularia douglasiae Tchangsinaia Cuneopsis Schistodesmus

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 196 8 3 11.0
Cladomics cladogram Nodularia douglasiae Nodularia huana Nodularia nuxpersicae Schistodesmus Tchangsinaia Cuneopsis

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.600 3 3 1 3 198 9 3 11.0
Cladomics cladogram Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Cuneopsis szechenyii Cuneopsis heudei Nodularia huana Nodularia douglasiae Nodularia nuxpersicae

Wu et al., 2026 [Publ 4825], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 16 2 1 9.0
Cladomics cladogram Schistodesmus Nodularia fusiformans Nodularia douglasiae Nodularia dualobtusa

Klishko et al., 2019 [Publ 3886], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 8 0 0 9.0
Cladomics cladogram Middendorffinaia Nodularia breviconcha Nodularia douglasiae Nodularia huana

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 2 2 0 2 39 12 6 --
Cladomics cladogram Middendorffinaia Nodularia douglasiae Nodularia breviconcha

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 2 2 0 2 39 12 6 --
Cladomics cladogram Middendorffinaia Nodularia douglasiae Nodularia breviconcha

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 2 2 0 2 42 11 6 --
Cladomics cladogram Nodularia douglasiae Nodularia breviconcha Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Unio Tchangsinaia Inversiunio

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 2 2 0 2 42 11 6 --
Cladomics cladogram Middendorffinaia Nodularia douglasiae Nodularia breviconcha

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 39 11 6 --
Cladomics cladogram Tchangsinaia Inversiunio Cuneopsis Pseudocuneopsis Nodularia douglasiae Nodularia breviconcha Diaurora Schistodesmus

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 39 11 6 --
Cladomics cladogram Tchangsinaia Inversiunio Pseudocuneopsis Cuneopsis Nodularia douglasiae Nodularia breviconcha Diaurora Schistodesmus

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 36 13 5 --
Cladomics cladogram Pseudocuneopsis Nodularia breviconcha Nodularia douglasiae Schistodesmus Cuneopsis Tchangsinaia Inversiunio

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 71 5 4 --
Cladomics cladogram Pseudocuneopsis Nodularia douglasiae Nodularia breviconcha Schistodesmus Cuneopsis Tchangsinaia

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 71 5 4 --
Cladomics cladogram Pseudocuneopsis Nodularia douglasiae Nodularia breviconcha Tchangsinaia Cuneopsis Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 66 5 4 --
Cladomics cladogram Pseudocuneopsis Nodularia douglasiae Nodularia breviconcha Schistodesmus Tchangsinaia Cuneopsis

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 71 10 4 --
Cladomics cladogram Pseudocuneopsis Nodularia douglasiae Nodularia breviconcha Schistodesmus Tchangsinaia Cuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 93 10 4 --
Cladomics cladogram Pseudocuneopsis Nodularia breviconcha Nodularia douglasiae Tchangsinaia Schistodesmus Cuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.833 2 2 0 2 93 10 4 --
Cladomics cladogram Pseudocuneopsis Nodularia breviconcha Nodularia douglasiae Schistodesmus lampreyanus Cuneopsis Tchangsinaia Schistodesmus xinyuensis Schistodesmus spinosus

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 89 10 4 --
Cladomics cladogram Pseudocuneopsis Nodularia breviconcha Nodularia douglasiae Schistodesmus Tchangsinaia Cuneopsis

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 89 10 4 --
Cladomics cladogram Pseudocuneopsis Nodularia breviconcha Nodularia douglasiae Schistodesmus Tchangsinaia Cuneopsis

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 51 12 4 --
Cladomics cladogram Pseudocuneopsis Nodularia breviconcha Nodularia douglasiae Schistodesmus Tchangsinaia Cuneopsis

Ouyang et al., 2015 [Publ 4493], fig. 1

cst iT tT bi riS oT roT roS iD
0.600 2 2 0 2 29 5 3 --
Cladomics cladogram Tchangsinaia Cuneopsis rufescens Cuneopsis szechenyii Cuneopsis celtiformis Nodularia douglasiae Nodularia huana

Ouyang et al., 2015 [Publ 4493], fig. 2

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 2 29 5 3 --
Cladomics cladogram Nodularia douglasiae Nodularia huana Cuneopsis rufescens 2 Schistodesmus Cuneopsis celtiformis 2 Tchangsinaia Cuneopsis celtiformis 1 Cuneopsis szechenyii 2 Cuneopsis szechenyii 1 Cuneopsis rufescens 1

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 79 4 3 --
Cladomics cladogram Nodularia nuxpersicae Nodularia douglasiae Schistodesmus Tchangsinaia Cuneopsis

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 137 4 3 --
Cladomics cladogram Nodularia douglasiae Nodularia nuxpersicae Cuneopsis Tchangsinaia Schistodesmus

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.600 2 2 0 2 137 4 3 --
Cladomics cladogram Scriptolamprotula Schistodesmus Nodularia douglasiae Nodularia nuxpersicae Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.400 2 10 0 2 123 4 3 18.0
Cladomics cladogram Unio crassus Unio pictorum Unio tumidus Scriptolamprotula Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 2 32 4 3 --
Cladomics cladogram Cuneopsis Nodularia nipponensis 1 Nodularia douglasiae Nodularia nipponensis 2

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 1 2 67 5 3 --
Cladomics cladogram Inversiunio Tchangsinaia Schistodesmus Nodularia nipponensis 1 Nodularia nipponensis 2 Nodularia douglasiae

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 2 2 67 5 3 --
Cladomics cladogram Inversiunio Schistodesmus Tchangsinaia Nodularia nipponensis 1 Nodularia nipponensis 2 Nodularia nipponensis 3 Nodularia douglasiae

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 2 2 67 5 3 --
Cladomics cladogram Inversiunio Schistodesmus Tchangsinaia Nodularia nipponensis 3 Nodularia nipponensis 1 Nodularia douglasiae Nodularia nipponensis 2

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 2 74 4 3 --
Cladomics cladogram Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 2 74 4 3 --
Cladomics cladogram Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens

Lopes-Lima et al., 2020 [Publ 4080], fig. 12

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 6 4 1 --
Cladomics cladogram Inversiunio Nodularia breviconcha Nodularia douglasiae

Chen et al., 2025 [Publ 4448], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 7 6 1 --
Cladomics cladogram Pseudocuneopsis Nodularia nipponensis Nodularia douglasiae

Froufe et al., 2016 [Publ 3738], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 12 0 0 --
Cladomics cladogram Unio Nodularia douglasiae Nodularia breviconcha

Klishko et al., 2017 [Publ 3946], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 4 0 0 --
Cladomics cladogram Unio Nodularia douglasiae Nodularia breviconcha

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 1 2 80 0 0 --
Cladomics cladogram Unio Nodularia douglasiae 1 Nodularia nuxpersicae Nodularia douglasiae 2

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 10 0 0 --
Cladomics cladogram Lanceolaria Nodularia breviconcha Nodularia douglasiae

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 7 0 0 --
Cladomics cladogram Scriptolamprotula Nodularia breviconcha Nodularia douglasiae

Wu et al., 2022 [Publ 4238], fig. 2a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 10 5 --
Cladomics cladogram Nodularia Pseudocuneopsis Schistodesmus Cuneopsis Tchangsinaia Inversiunio

Wu et al., 2023 [Publ 4296], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 9 5 --
Cladomics cladogram Cuneopsis Nodularia douglasiae Schistodesmus Tchangsinaia Inversiunio Pseudocuneopsis

Liu et al., 2023 [Publ 4310], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 23 10 5 --
Cladomics cladogram Cuneopsis Nodularia douglasiae Schistodesmus Tchangsinaia Inversiunio Pseudocuneopsis

Liu et al., 2023 [Publ 4310], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 23 10 5 --
Cladomics cladogram Cuneopsis Nodularia douglasiae Schistodesmus Tchangsinaia Inversiunio Pseudocuneopsis

Zhang et al., 2004 [Publ 2735], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 13 6 4 --
Cladomics cladogram Tchangsinaia Cuneopsis heudei Nodularia douglasiae Pseudocuneopsis Cuneopsis rufescens Schistodesmus

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 4 4 --
Cladomics cladogram Pseudocuneopsis Nodularia douglasiae Tchangsinaia Cuneopsis Schistodesmus

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 53 5 4 --
Cladomics cladogram Cuneopsis Schistodesmus Pseudocuneopsis Nodularia douglasiae Tchangsinaia

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 4 4 --
Cladomics cladogram Pseudocuneopsis Nodularia douglasiae Tchangsinaia Cuneopsis Schistodesmus

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 5 4 --
Cladomics cladogram Pseudocuneopsis Nodularia douglasiae Schistodesmus Cuneopsis Tchangsinaia

Wu et al., 2022 [Publ 4238], fig. 2e

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 8 4 --
Cladomics cladogram Pseudocuneopsis Nodularia douglasiae Tchangsinaia Cuneopsis Schistodesmus

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 4 4 --
Cladomics cladogram Pseudocuneopsis Nodularia douglasiae Schistodesmus Cuneopsis Tchangsinaia

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 4 4 --
Cladomics cladogram Pseudocuneopsis Schistodesmus Nodularia breviconcha Tchangsinaia Cuneopsis

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 4 4 --
Cladomics cladogram Nodularia douglasiae Pseudocuneopsis Schistodesmus Tchangsinaia Cuneopsis

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 8 4 --
Cladomics cladogram Pseudocuneopsis Nodularia douglasiae Tchangsinaia Cuneopsis Schistodesmus

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 6 3 --
Cladomics cladogram Nodularia douglasiae Schistodesmus Tchangsinaia Cuneopsis

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 3 3 --
Cladomics cladogram Nodularia douglasiae Tchangsinaia Cuneopsis Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 5 3 --
Cladomics cladogram Nodularia douglasiae Cuneopsis Tchangsinaia Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 5 3 --
Cladomics cladogram Nodularia douglasiae Cuneopsis Tchangsinaia Schistodesmus

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 33 2 2 --
Cladomics cladogram Nodularia douglasiae Tchangsinaia

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 33 2 2 --
Cladomics cladogram Nodularia douglasiae Tchangsinaia

Zhou et al., 2007 [Publ 2658], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 20 3 2 --
Cladomics cladogram Tchangsinaia Nodularia douglasiae

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 2 2 --
Cladomics cladogram Schistodesmus Nodularia douglasiae Tchangsinaia

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 31 2 2 --
Cladomics cladogram Schistodesmus Tchangsinaia Nodularia douglasiae

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 2 2 --
Cladomics cladogram Schistodesmus Tchangsinaia Nodularia douglasiae

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 2 2 --
Cladomics cladogram Nodularia douglasiae Tchangsinaia Cuneopsis

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 2 2 --
Cladomics cladogram Schistodesmus Nodularia Tchangsinaia

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 2 2 --
Cladomics cladogram Nodularia douglasiae Tchangsinaia Schistodesmus

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 2 2 --
Cladomics cladogram Nodularia douglasiae Tchangsinaia Schistodesmus

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 2 2 --
Cladomics cladogram Nodularia douglasiae Tchangsinaia Schistodesmus

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 2 2 --
Cladomics cladogram Nodularia douglasiae Schistodesmus Tchangsinaia

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 36 3 2 --
Cladomics cladogram Schistodesmus Nodularia nipponensis Tchangsinaia

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 3 2 --
Cladomics cladogram Schistodesmus Nodularia nipponensis Tchangsinaia

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 3 2 --
Cladomics cladogram Nodularia douglasiae Tchangsinaia Cuneopsis

Dai et al., 2024 [Publ 4358], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 5 2 --
Cladomics cladogram Pseudocuneopsis Schistodesmus Nodularia douglasiae

Zhou et al., 2007 [Publ 2658], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 1 1 --
Cladomics cladogram Nodularia douglasiae Cuneopsis

Ouyang et al., 2011 [Publ 3408], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 1 1 --
Cladomics cladogram Nodularia douglasiae Cuneopsis

Xue et al., 2016 [Publ 3659], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 14 1 1 --
Cladomics cladogram Nodularia douglasiae Unio

Zhou et al., 2017 [Publ 3788], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 1 1 --
Cladomics cladogram Nodularia douglasiae Tchangsinaia

Sano et al., 2017 [Publ 3799], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 3 1 --
Cladomics cladogram Nodularia nipponensis Inversiunio

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 1 1 --
Cladomics cladogram Tchangsinaia Nodularia douglasiae

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Tchangsinaia Nodularia douglasiae

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 1 1 --
Cladomics cladogram Nodularia douglasiae Tchangsinaia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 1 1 --
Cladomics cladogram Tchangsinaia Nodularia douglasiae

Kano et al., 2019 [Publ 4194], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 1 1 --
Cladomics cladogram Inversiunio Nodularia nipponensis

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 11 1 1 --
Cladomics cladogram Nodularia douglasiae Lanceolaria Inversiunio

Wu et al., 2021 [Publ 4215], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 23 2 1 --
Cladomics cladogram Nodularia douglasiae Inversiunio

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 --
Cladomics cladogram Tchangsinaia Nodularia douglasiae

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 1 1 --
Cladomics cladogram Tchangsinaia Nodularia dualobtusa

Araujo et al., 2009 [Publ 2589], fig. 7A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Nodularia douglasiae Unio gibbus Unio tumidus Anodonta Unio pictorum 1 Unio pictorum 2 Unio mancus Unio tumidiformis Unio crassus

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram Nodularia douglasiae Anodonta cygnea Unio pictorum 2 Unio pictorum 1 Unio caffer Anodonta Lasmigona Strophitus Alasmidonta Pyganodon

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Unio Nodularia douglasiae

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 0 0 --
Cladomics cladogram Unio Nodularia douglasiae

Araujo et al., 2018 [Publ 3779], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Nodularia douglasiae Unio

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Unio Nodularia douglasiae

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Nodularia douglasiae Unio

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 0 0 --
Cladomics cladogram Anodonta Nodularia douglasiae

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Anodonta Nodularia douglasiae

Altaba et al., 2025 [Publ 4849], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Nodularia douglasiae Unio

Altaba et al., 2025 [Publ 4849], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Nodularia douglasiae Unio

Altaba et al., 2025 [Publ 4849], fig. 12

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram Unio Nodularia douglasiae