Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Anodontina

genus Anodonta Lamarck, 1799: cladomics

5 Recent valid species (20 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (20 total)
analyzed in 221 cladograms in 119 publications
maximum number of terminals in an analysis = 4 (mean = 1.57, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 94
maximum number of included terminals = 4, (mean = 2.33 median = 2)
proportion of cladograms that support basic monophyly = 0.49

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Anodonta cladogram distances
PCoA Ordination

NJ Tree of Anodonta cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
0.500 4 10 9 4 629 6 2 **
Cladomics cladogram Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus Anodonta anatina Pseudanodonta Brachyanodon oregonensis 2 Anodonta cygnea Anodonta seddonae Anodonta exulcerata Brachyanodon impura Brachyanodon Brachyanodon nuttalliana 1 Brachyanodon nuttalliana 2 Brachyanodon californiensis Brachyanodon oregonensis 1

Lyubas et al., 2022 [Publ 4255], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 27 1 1 8.0
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea Anodonta exulcerata Anodonta seddonae

Lyubas et al., 2022 [Publ 4255], fig. S1

cst iT tT bi riS oT roT roS iD
0.667 4 5 3 4 27 1 1 9.0
Cladomics cladogram Unio Anodonta anatina Pseudanodonta Anodonta exulcerata Anodonta seddonae Anodonta cygnea

Lopes-Lima et al., 2021 [Publ 4217], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 0 0 0 10.0
Cladomics cladogram Anodonta anatina Anodonta exulcerata Anodonta seddonae Anodonta cygnea

Riccardi et al., 2020 [Publ 4089], fig. 2

cst iT tT bi riS oT roT roS iD
0.500 3 9 7 3 16 2 2 13.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta anatina Pseudanodonta Anodonta exulcerata Anodonta cygnea Brachyanodon Platynaias Pyganodon Utterbackia

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 92 2 2 9.0
Cladomics cladogram Brachyanodon Pseudanodonta Anodonta anatina Anodonta cygnea Anodonta exulcerata

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
0.500 3 10 7 3 92 2 2 17.0
Cladomics cladogram Brachyanodon Pseudanodonta Anodonta cygnea Anodonta exulcerata Anodonta anatina Pyganodon Utterbackia Pterosyna Prolasmidonta Alasmidonta Platynaias

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 3 88 2 2 9.0
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea Anodonta exulcerata

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 3 88 2 2 9.0
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea Anodonta exulcerata

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 3 50 2 2 9.0
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea Anodonta exulcerata

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 3 72 2 2 9.0
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea Anodonta exulcerata

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 3 3 1 3 49 2 2 9.0
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea Anodonta exulcerata

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
0.500 3 13 11 3 137 2 2 18.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta exulcerata Anodonta anatina Brachyanodon Pseudanodonta Pyganodon Utterbackia Pterosyna Prolasmidonta Alasmidonta Platynaias

Nagel et al., 1996 [Publ 3400], fig. 351

cst iT tT bi riS oT roT roS iD
0.333 3 4 3 3 2 1 1 12.0
Cladomics cladogram Pseudanodonta Anodonta cygnea Sinanodonta Anodonta 2 Anodonta anatina Anodonta 1

Nagel et al., 1998 [Publ 1946], fig. 7

cst iT tT bi riS oT roT roS iD
0.333 3 4 3 3 2 1 1 12.0
Cladomics cladogram Pseudanodonta Anodonta cygnea Sinanodonta Anodonta 2 Anodonta anatina Anodonta 1

Riccardi et al., 2020 [Publ 4089], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 3 4 2 3 3 1 1 10.0
Cladomics cladogram Sinanodonta Anemina Anodonta anatina Pseudanodonta Anodonta exulcerata Anodonta cygnea

Tomilova et al., 2020 [Publ 4097], fig. 3

cst iT tT bi riS oT roT roS iD
0.667 3 4 2 3 5 1 1 8.0
Cladomics cladogram Unio Sinanodonta Anodonta anatina Pseudanodonta Anodonta cygnea Anodonta exulcerata

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

cst iT tT bi riS oT roT roS iD
0.000 3 9 7 3 25 4 1 15.0
Cladomics cladogram Beringiana Sinanodonta Anodonta exulcerata Anodonta cygnea Brachyanodon californiensis Brachyanodon nuttalliana Brachyanodon kennerlyi Brachyanodon oregonensis Anodonta anatina Utterbackiana

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
0.667 3 3 1 3 61 1 1 9.0
Cladomics cladogram Anodonta anatina Anodonta exulcerata Anodonta cygnea Pseudanodonta Pyganodon Utterbackia Pterosyna Prolasmidonta Alasmidonta Platynaias

Altaba et al., 2025 [Publ 4849], fig. 11

cst iT tT bi riS oT roT roS iD
0.333 3 4 2 3 11 3 1 10.0
Cladomics cladogram Brachyanodon kennerlyi Brachyanodon oregonensis Anodonta cygnea Anodonta exulcerata Anodonta anatina Brachyanodon californiensis

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 3 3 1 3 92 1 1 9.0
Cladomics cladogram Anodonta anatina Anodonta cygnea Anodonta exulcerata Pseudanodonta Pyganodon Utterbackia Pterosyna Prolasmidonta Alasmidonta Platynaias

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
0.667 3 3 1 3 62 1 1 9.0
Cladomics cladogram Anodonta anatina Anodonta cygnea Anodonta exulcerata Pseudanodonta Pyganodon Utterbackia Pterosyna Prolasmidonta Alasmidonta Platynaias

Froufe et al., 2017 [Publ 3802], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 1 0 0 9.0
Cladomics cladogram Cristaria Anodonta anatina Anodonta cygnea Anodonta exulcerata

Froufe et al., 2017 [Publ 3802], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 7 0 0 9.0
Cladomics cladogram Sinanodonta Anodonta anatina Anodonta exulcerata Anodonta cygnea

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 3 0 0 9.0
Cladomics cladogram Anodonta anatina Anodonta exulcerata Anodonta cygnea Cristaria Anemina Sinanodonta

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 22 0 0 9.0
Cladomics cladogram Unio Anodonta anatina Anodonta cygnea Anodonta seddonae

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 9 0 0 9.0
Cladomics cladogram Cristaria Sinanodonta Anodonta anatina Anodonta exulcerata Anodonta cygnea

Chong et al., 2008 [Publ 2668], fig. 3

cst iT tT bi riS oT roT roS iD
0.250 2 18 0 2 16 5 2 24.0
Cladomics cladogram Anodonta cygnea Pyganodon Alasminota Anodonta anatina Unio Brachyanodon nuttalliana Brachyanodon californiensis Beringiana Sinanodonta Pseudodontoideus Alasmidonta Pterosyna Pseudanodonta Strophitus Prolasmidonta Platynaias Brachyanodon oregonensis 4 Brachyanodon oregonensis 3 Brachyanodon oregonensis 2 Brachyanodon oregonensis 1 Brachyanodon kennerlyi

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.750 2 3 0 2 79 2 2 10.0
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea Brachyanodon

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.500 2 9 0 2 137 2 2 13.0
Cladomics cladogram Cristaria Pletholophus Sinanodonta Anodonta anatina Pseudanodonta Anodonta cygnea Brachyanodon Lasmigona Alasmidonta Simpsonaias Pyganodon Strophitus

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.500 2 9 0 2 137 2 2 14.0
Cladomics cladogram Lanceolaria Cristaria Pletholophus Sinanodonta Anodonta anatina Pseudanodonta Anodonta cygnea Brachyanodon Pyganodon Simpsonaias Strophitus Alasmidonta Lasmigona

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.500 2 18 0 2 123 2 2 23.0
Cladomics cladogram Unio crassus Unio pictorum Unio tumidus Scriptolamprotula Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.500 2 3 0 2 199 2 2 9.0
Cladomics cladogram Alasmidonta Lasmigona Pseudanodonta Simpsonaias Strophitus Pyganodon Utterbackia Anodonta anatina Anodonta cygnea Brachyanodon

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.500 2 3 0 2 197 2 2 9.0
Cladomics cladogram Alasmidonta Lasmigona Anodonta anatina Anodonta cygnea Brachyanodon Pyganodon Utterbackia Pseudanodonta Simpsonaias Strophitus

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.500 2 48 0 2 199 2 2 53.0
Cladomics cladogram Chamberlainia Megalonaias Uniomerus Quadrula Sinohyriopsis Lamprotula Potomida Pronodularia Indopseudodon Nyeinchanconcha Thaiconcha Sundadontina cumingii Sundadontina tumida 1 Sundadontina tumida 2 Pseudodon Pilsbryoconcha Bineurus Sinosolenaia carinata Gonidea Leguminaia Microcondylaea Ptychorhynchus Parvasolenaia Koreosolenaia Sinosolenaia recognita Sinosolenaia iridinea Sinosolenaia oleivora Physunio modelli Physunio Physunio superbus Yaukthwa Lens Hyriopsis desowitzi Hyriopsis myersiana Hyriopsis panhai Hyriopsis bialata Hyriopsis khoratensis Rectidens Ensidens Amblema Toxolasma Obliquaria Plectomerus Elliptoideus Elliptio Eurynia Fusconaia Pleurobema Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus Unio Scriptolamprotula Lanceolaria Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.500 2 3 0 2 120 2 2 11.0
Cladomics cladogram Pseudanodonta Anodonta cygnea Anodonta anatina Brachyanodon Pyganodon Simpsonaias Strophitus Alasmidonta Lasmigona

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 74 2 2 --
Cladomics cladogram Pseudanodonta Brachyanodon Anodonta anatina Anodonta cygnea

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
0.750 2 3 0 2 74 2 2 9.0
Cladomics cladogram Pseudanodonta Anodonta anatina Brachyanodon Anodonta cygnea

Whelan et al., 2023 [Publ 4287], fig. 2

cst iT tT bi riS oT roT roS iD
0.500 2 37 0 2 39 2 2 47.0
Cladomics cladogram Anemina Sinanodonta Anodonta cygnea Brachyanodon Pseudanodonta Anodonta anatina Utterbackia Pyganodon Utterbackiana Lasmigona Pseudodontoideus Pterosyna Arcidens Sulcularia Anodontoides Strophitus Prolasmidonta Platynaias Alasminota Alasmidonta

Nagel et al., 1998 [Publ 1946], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 7 1 1 --
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea

Källersjö et al., 2005 [Publ 2684], fig. 1A

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 5 1 1 10.0
Cladomics cladogram Margaritifera Unio Anodonta anatina Pseudanodonta Anodonta cygnea

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 5 1 1 9.0
Cladomics cladogram Margaritifera Unio Anodonta anatina Pseudanodonta Anodonta cygnea

Manendo et al., 2008 [Publ 2669], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 2 2 0 2 11 1 1 --
Cladomics cladogram Anodonta cygnea Anodonta anatina Pseudanodonta Platynaias Pyganodon

Soroka, 2010 [Publ 3404], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 5 1 1 11.0
Cladomics cladogram Unio Anodonta cygnea Anodonta anatina Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 3a

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 7 1 1 9.0
Cladomics cladogram Unio Anodonta anatina Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 3b

cst iT tT bi riS oT roT roS iD
0.333 2 9 0 2 7 1 1 15.0
Cladomics cladogram Anodonta anatina Anodonta cygnea Pseudanodonta Sinanodonta Unio

Soroka, 2010 [Publ 3404], fig. 4

cst iT tT bi riS oT roT roS iD
0.333 2 7 0 2 5 1 1 12.0
Cladomics cladogram Anodonta anatina Unio Sinanodonta Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 5a

cst iT tT bi riS oT roT roS iD
0.333 2 5 0 2 6 1 1 11.0
Cladomics cladogram Unio Anodonta anatina Sinanodonta Anodonta cygnea Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 5b

cst iT tT bi riS oT roT roS iD
0.333 2 8 0 2 6 1 1 13.0
Cladomics cladogram Anodonta anatina Sinanodonta Unio Anodonta cygnea Pseudanodonta

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
0.667 2 2 0 2 16 1 1 --
Cladomics cladogram Lanceolaria Unio Pyganodon Pseudanodonta Anodonta anatina Anodonta cygnea Alathyria Pronodularia Sinohyriopsis Quadrula Parvaspina Lampsilis Epioblasma Venustaconcha

Zieritz et al., 2012 [Publ 3673], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 6 1 1 10.0
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea Sinanodonta Margaritifera Unio

Hinzmann et al., 2013 [Publ 3541], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 1 1 1 --
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea

Reis et al., 2013 [Publ 3580], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 12 1 1 11.0
Cladomics cladogram Unio Anodonta cygnea Anodonta anatina Pseudanodonta

Reis et al., 2013 [Publ 3580], fig. 5

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 13 1 1 10.0
Cladomics cladogram Unio Anodonta anatina Anodonta cygnea Pseudanodonta

Prié & Puillandre, 2014 [Publ 3529], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 15 1 1 9.0
Cladomics cladogram Unio Anodonta anatina Pseudanodonta Anodonta cygnea

Froufe et al., 2014 [Publ 3592], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 1 1 1 9.0
Cladomics cladogram Anodonta anatina Pseudanodonta Anodonta cygnea

Mezhzherin et al., 2014 [Publ 3705], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 2 1 1 --
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea

Mezhzherin et al., 2014 [Publ 3705], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 2 1 1 9.0
Cladomics cladogram Sinanodonta Anodonta anatina Anodonta cygnea Pseudanodonta

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 1 1 1 --
Cladomics cladogram Pseudanodonta Anodonta anatina Anodonta cygnea

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 15 1 1 9.0
Cladomics cladogram Unio Anodonta anatina Pseudanodonta Anodonta cygnea

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 34 1 1 9.0
Cladomics cladogram Anodonta anatina Pseudanodonta Anodonta cygnea Amuranodonta Anemina Buldowskia Cristaria Beringiana Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
0.667 2 2 0 2 89 1 1 --
Cladomics cladogram Anodonta anatina Anodonta cygnea Pseudanodonta Pyganodon Strophitus

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
0.333 2 6 0 2 89 1 1 14.0
Cladomics cladogram Anemina Cristaria Sinanodonta Anodonta cygnea Lasmigona Anodonta anatina Pseudanodonta Pyganodon Strophitus

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 54 1 1 9.0
Cladomics cladogram Anodonta anatina Pseudanodonta Anodonta cygnea Anemina Amuranodonta Buldowskia Cristaria Beringiana Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 5

cst iT tT bi riS oT roT roS iD
0.667 2 3 0 2 17 1 1 9.0
Cladomics cladogram Platynaias Anodonta anatina Pseudanodonta Anodonta cygnea

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

cst iT tT bi riS oT roT roS iD
0.667 2 2 0 2 34 1 1 --
Cladomics cladogram Anodonta cygnea Anodonta anatina Pseudanodonta Utterbackiana Alasminota Pseudodontoideus Lasmigona Pterosyna Arcidens Prolasmidonta Platynaias Ligodonta Alasmidonta Anodontoides Strophitus Sulcularia Pyganodon Utterbackia

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

cst iT tT bi riS oT roT roS iD
0.000 2 8 0 2 23 0 0 17.0
Cladomics cladogram Nodularia Anodonta cygnea Unio pictorum 2 Unio pictorum 1 Unio caffer Anodonta Lasmigona Strophitus Alasmidonta Pyganodon

Soroka, 2010 [Publ 3404], fig. 8a

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 4 0 0 --
Cladomics cladogram Sinanodonta Anodonta anatina Anodonta cygnea

Soroka, 2010 [Publ 3404], fig. 8b

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 4 0 0 --
Cladomics cladogram Unio Anodonta anatina Anodonta cygnea

Froufe et al., 2017 [Publ 3802], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 10 0 0 --
Cladomics cladogram Anodonta anatina Anodonta cygnea Anemina Cristaria Beringiana Sinanodonta

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 21 0 0 --
Cladomics cladogram Anodonta cygnea Anodonta anatina Platynaias Pyganodon Utterbackia

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
0.000 2 7 0 2 51 0 0 14.0
Cladomics cladogram Anemina Cristaria Sinanodonta Anodonta cygnea Anodonta anatina Strophitus Platynaias Alasmidonta Pyganodon

Chase et al., 2018 [Publ 3847], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 37 0 0 --
Cladomics cladogram Anodonta cygnea Anodonta anatina Pyganodon Utterbackia Lasmigona Pterosyna Alasmidonta Strophitus Platynaias

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 38 0 0 --
Cladomics cladogram Anodonta anatina Anodonta cygnea Prolasmidonta Alasmidonta Pyganodon Utterbackia

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
0.000 2 6 0 2 54 0 0 13.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta anatina Lasmigona Pyganodon Utterbackia

Klishko et al., 2019 [Publ 3886], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 9 0 0 --
Cladomics cladogram Anodonta anatina Anodonta cygnea Unio Middendorffinaia Nodularia

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 52 0 0 --
Cladomics cladogram Anodonta anatina Anodonta cygnea Pyganodon Utterbackia Alasmidonta Prolasmidonta

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 67 0 0 --
Cladomics cladogram Anodonta cygnea Anodonta anatina Pyganodon Utterbackia Alasmidonta Prolasmidonta Platynaias

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 67 0 0 --
Cladomics cladogram Anodonta anatina Anodonta cygnea Pyganodon Utterbackia Alasmidonta Platynaias Prolasmidonta

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 67 0 0 --
Cladomics cladogram Anodonta cygnea Anodonta anatina Pyganodon Utterbackia Alasmidonta Prolasmidonta Platynaias

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 32 0 0 --
Cladomics cladogram Cristaria Anodonta cygnea Anodonta anatina

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
0.000 2 9 0 2 71 0 0 16.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta anatina Pyganodon Utterbackia Prolasmidonta Alasmidonta Platynaias

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
0.000 2 9 0 2 71 0 0 16.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta anatina Pyganodon Utterbackia Prolasmidonta Alasmidonta Platynaias

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
0.000 2 8 0 2 66 0 0 15.0
Cladomics cladogram Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta anatina Pyganodon Utterbackia Prolasmidonta Platynaias Alasmidonta

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 36 0 0 --
Cladomics cladogram Anodonta cygnea Anodonta anatina Platynaias Pyganodon Utterbackia

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 71 0 0 --
Cladomics cladogram Anodonta cygnea Anodonta anatina Platynaias Pyganodon Utterbackia

Whelan et al., 2023 [Publ 4287], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 25 0 0 --
Cladomics cladogram Anodonta anatina Anodonta cygnea Sulcularia Utterbackiana Alasminota Arcidens Pseudodontoideus Alasmidonta Platynaias Strophitus Pyganodon Utterbackia Anodontoides

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 0 0 --
Cladomics cladogram Lanceolaria Anodonta anatina Anodonta cygnea

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 20 0 0 --
Cladomics cladogram Lanceolaria Anodonta anatina Anodonta cygnea

Xiang et al., 2025 [Publ 4433], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 11 0 0 --
Cladomics cladogram Anodonta cygnea Anodonta exulcerata Cristaria Anemina Beringiana Mimo Sinanodonta

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 85 0 0 --
Cladomics cladogram Anodonta anatina Anodonta cygnea Platynaias Pyganodon Utterbackia

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 83 0 0 --
Cladomics cladogram Anodonta cygnea Anodonta anatina Platynaias Pyganodon Utterbackia

Altaba et al., 2025 [Publ 4849], fig. 10

cst iT tT bi riS oT roT roS iD
0.000 2 8 0 2 10 0 0 15.0
Cladomics cladogram Nodularia Unio Anodonta anatina Anodonta cygnea Anemina Amuranodonta Cristaria Beringiana Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 12

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 11 0 0 --
Cladomics cladogram Sinanodonta Anodonta anatina Anodonta cygnea

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 1 1 --
Cladomics cladogram Pseudanodonta Anodonta anatina

Hoeh, 1990 [Publ 2717], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 14 1 1 --
Cladomics cladogram Anodonta cygnea Brachyanodon Utterbackiana

Hoeh, 1990 [Publ 2717], fig. 3

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 14 1 1 --
Cladomics cladogram Pyganodon fragilis Utterbackia imbecillis Utterbackia peggyae Anodonta cygnea Brachyanodon Utterbackiana couperiana 1 Utterbackiana couperiana 2 Utterbackiana suborbiculata Lasmigona Utterbackiana implicata Pterosyna Platynaias Pyganodon cataracta Pyganodon gibbosa Pyganodon grandis Pyganodon lacustris

Hoeh, 1990 [Publ 2717], fig. 4

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 14 1 1 --
Cladomics cladogram Anodonta cygnea Brachyanodon Utterbackiana

Nagel et al., 1998 [Publ 1946], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 1 1 --
Cladomics cladogram Anodonta Pseudanodonta

Campbell et al., 2005 [Publ 2660], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 101 1 1 --
Cladomics cladogram Anodonta cygnea Alasminota

Campbell et al., 2005 [Publ 2660], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 101 1 1 --
Cladomics cladogram Anodonta cygnea Brachyanodon

Zanatta et al., 2007 [Publ 2698], fig. 1A

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 15 1 1 --
Cladomics cladogram Anodonta cygnea Brachyanodon Utterbackiana Utterbackia Pyganodon

Zanatta et al., 2007 [Publ 2698], fig. 1B

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 15 1 1 --
Cladomics cladogram Anodonta cygnea Brachyanodon Utterbackiana

Bogatov et al., 2018 [Publ 3859], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 1 1 --
Cladomics cladogram Anodonta anatina Pseudanodonta

Bogatov et al., 2018 [Publ 3859], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 1 1 --
Cladomics cladogram Anodonta anatina Pseudanodonta

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram Anodonta anatina Pseudanodonta

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram Anodonta anatina Pseudanodonta

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 1 1 --
Cladomics cladogram Anodonta anatina Pseudanodonta

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 1 1 --
Cladomics cladogram Anodonta anatina Pseudanodonta

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram Anodonta anatina Pseudanodonta

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 1 1 --
Cladomics cladogram Anodonta anatina Pseudanodonta

Lydeard et al., 1996 [Publ 2656], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Utterbackiana Anodonta

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Anodonta anatina Gonidea Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus Pleurobema Unio

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Anodonta Gonidea Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus Amblema Elliptio Fusconaia Plectomerus Scriptolamprotula Unio

Hoeh et al., 1998 [Publ 2695], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram Anodonta cygnea Margaritifera Amblema

Hoeh et al., 1998 [Publ 2695], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram Margaritifera Anodonta cygnea

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Anodonta cygnea Alasmidonta Lasmigona Strophitus Pyganodon

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Anodonta cygnea Alasmidonta Lasmigona Strophitus Pyganodon

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 0 0 --
Cladomics cladogram Anodonta cygnea Strophitus Pyganodon

Lydeard et al., 2000 [Publ 2654], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta cygnea Cumberlandia Reginaia Lampsilis Obovaria Quadrula Pustulosa Pleurobema Fusconaia

Lydeard et al., 2000 [Publ 2654], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta cygnea Cumberlandia Lampsilis Obovaria Reginaia Pleurobema Fusconaia Quadrula Pustulosa infucata Pustulosa succissa

Lydeard et al., 2000 [Publ 2654], fig. 2c

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta cygnea Cumberlandia Quadrula Pustulosa Obovaria unicolor Obovaria olivaria Lampsilis Reginaia Pleurobema Fusconaia

Lydeard et al., 2000 [Publ 2654], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta cygnea Cumberlandia Reginaia Lampsilis Obovaria Pleurobema Fusconaia Quadrula Pustulosa infucata Pustulosa succissa

Hoeh et al., 2001 [Publ 2652], fig. 14f2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Anodonta cygnea Strophitus

Hoeh et al., 2001 [Publ 2652], fig. 14f4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Anodonta cygnea Strophitus Pyganodon

Hoeh et al., 2002 [Publ 2653], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Anodonta cygnea Pyganodon Strophitus

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Anodonta cygnea Strophitus

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Anodonta cygnea Pyganodon Strophitus

Krebs et al., 2003 [Publ 2691], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 32 0 0 --
Cladomics cladogram Pyganodon Anodonta

Mock et al., 2004 [Publ 2686], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 0 0 --
Cladomics cladogram Anodonta Sinanodonta

Mock et al., 2004 [Publ 2686], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 0 0 --
Cladomics cladogram Anodonta Sinanodonta

Huff et al., 2004 [Publ 2697], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Unio Anodonta

Huff et al., 2004 [Publ 2697], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Anodonta Margaritifera margaritifera Cumberlandia Pseudunio Margaritifera dahurica Margaritifera middendorffi 1 Margaritifera middendorffi 2 Margaritifera falcata 1 Margaritifera falcata 2 Margaritifera falcata 3 Margaritifera falcata 4

Huff et al., 2004 [Publ 2697], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Unio Anodonta

Huff et al., 2004 [Publ 2697], fig. 5-6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Unio Anodonta Lampsilis

Walker et al., 2006 [Publ 2666], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Anodonta Pyganodon

Walker et al., 2006 [Publ 2666], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Anodonta Pyganodon

Walker et al., 2006 [Publ 2666], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Anodonta Pyganodon

Taylor et al., 2007 [Publ 2696], fig. 1-3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram Anodonta cygnea Unio

Soroka, 2008 [Publ 3695], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon

Soroka, 2008 [Publ 3695], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon

Araujo et al., 2009 [Publ 2589], fig. 7A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Unio gibbus Unio tumidus Anodonta cygnea

Araujo et al., 2009 [Publ 2589], fig. 7B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Anodonta cygnea Unio

Araujo et al., 2009 [Publ 2589], fig. 7C

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Anodonta cygnea Unio

Soroka, 2010 [Publ 3404], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 4 0 0 --
Cladomics cladogram Anodonta anatina Unio

Soroka, 2010 [Publ 3404], fig. 3a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Anodonta anatina Sinanodonta Unio

Soroka, 2010 [Publ 3404], fig. 3b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Anodonta anatina Sinanodonta Unio

Soroka, 2010 [Publ 3404], fig. 4-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 0 0 --
Cladomics cladogram Anodonta anatina Sinanodonta

Soroka, 2010 [Publ 3404], fig. 5a-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 0 0 --
Cladomics cladogram Anodonta anatina Sinanodonta

Soroka, 2010 [Publ 3404], fig. 5b-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 1 0 0 --
Cladomics cladogram Anodonta anatina Sinanodonta

Froufe et al., 2015 [Publ 3664], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Froufe et al., 2015 [Publ 3664], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Utterbackia

Bolotov et al., 2016 [Publ 3743], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Anodonta anatina Unio

Bolotov et al., 2016 [Publ 3743], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Anodonta anatina Unio

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Fonseca et al., 2016 [Publ 3775], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Feng et al., 2016 [Publ 3800], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Burzynski et al., 2017 [Publ 3774], fig. 1-top

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 31 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Anodonta anatina Sinanodonta Cristaria

Wen et al., 2017 [Publ 3795], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Araujo et al., 2017 [Publ 3801], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta anatina Unio

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Utterbackia

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Utterbackia

Araujo et al., 2018 [Publ 3779], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Alasmidonta

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 81 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Simpsonaias Strophitus Alasmidonta Lasmigona

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Huang et al., 2018 [Publ 3828], fig. 6-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 16 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Utterbackia

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Platynaias

Chase et al., 2018 [Publ 3847], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia

Chase et al., 2018 [Publ 3861], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Chase et al., 2018 [Publ 3861], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 10 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Sinanodonta Anodonta anatina Utterbackia Pyganodon

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Platynaias

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Utterbackia

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Utterbackia

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 0 0 --
Cladomics cladogram Anodonta anatina Pterosyna Pyganodon Utterbackia

Froufe et al., 2019 [Publ 3936], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Froufe et al., 2019 [Publ 3936], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 0 0 --
Cladomics cladogram Anodonta anatina Pterosyna Pyganodon Utterbackia

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram Anodonta cygnea Pyganodon

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram Anodonta cygnea Pyganodon

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Unio Anodonta anatina

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Unio Anodonta anatina

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Anodonta anatina Sinanodonta

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Utterbackia

Sonowal et al., 2021 [Publ 4221], fig. 3a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Anodonta cygnea Unio ravoisieri Unio pictorum Scriptolamprotula

Sonowal et al., 2021 [Publ 4221], fig. 3b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Scriptolamprotula Lamprotula Lampsilis Potomida Unio pictorum Anodonta cygnea Unio ravoisieri Lamellidens Elliptio

Sonowal et al., 2021 [Publ 4221], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Anodonta cygnea Unio Lamellidens

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Wu et al., 2022 [Publ 4238], fig. 2e

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 36 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 0 0 --
Cladomics cladogram Anodonta cygnea Platynaias Pyganodon Utterbackia

Whelan et al., 2023 [Publ 4287], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 27 0 0 --
Cladomics cladogram Anodonta anatina Utterbackiana Utterbackia Pyganodon Alasmidonta Arcidens Pseudodontoideus Alasminota Anodontoides Strophitus Sulcularia Platynaias Prolasmidonta

Bogan et al., 2023 [Publ 4295], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Anodonta cygnea Cristaria Amuranodonta Anemina Buldowskia Pletholophus Beringiana Sinanodonta

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 0 0 --
Cladomics cladogram Anodonta anatina Nodularia

Gomes-dos-Santos et al., 2023 [Publ 4463], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Anodonta anatina Nodularia

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 0 0 --
Cladomics cladogram Anodonta anatina Platynaias Pyganodon Utterbackia

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 0 0 --
Cladomics cladogram Anodonta cygnea Unio

Dai et al., 2024 [Publ 4386], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 0 0 --
Cladomics cladogram Anodonta anatina Lepidodesma

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Acuticosta Anemina Anodonta anatina Lanceolaria Scriptolamprotula Lepidodesma Unio Middendorffinaia Cuneopsis Nodularia Inversiunio Schistodesmus

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Anodonta anatina Acuticosta Anemina Lepidodesma Lanceolaria Scriptolamprotula Unio Middendorffinaia Nodularia Cuneopsis Inversiunio Schistodesmus

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 0 0 --
Cladomics cladogram Anodonta cygnea Alasmidonta Utterbackia

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Anodonta anatina Utterbackia Pyganodon

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 0 0 --
Cladomics cladogram Anodonta anatina Pyganodon Utterbackia