Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Anodontina

genus Anodonta Lamarck, 1799: cladomics

5 Recent valid species (18 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (18 total)
analyzed in 213 cladograms in 115 publications
maximum number of terminals in an analysis = 4 (mean = 1.56, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 90
maximum number of included terminals = 4, (mean = 2.32 median = 2)
proportion of cladograms that support basic monophyly = 0.49

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Anodonta cladogram distances
PCoA Ordination

NJ Tree of Anodonta cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 10 9 4 627 6 2 0 
  +----------------------- Anodonta_anatina **           
  |                                                   
--+  +-------------------- Pseudanodonta              
  |  |                                                
  +--+  +----------------- Brachyanodon_oregonensis__2
     |  |                                             
     |  |           +----- Anodonta_cygnea **            
     +--+  +--------+                                 
        |  |        |  +-- Anodonta_seddonae **          
        |  |        +--+                              
        +--+           +-- Anodonta_exulcerata **        
           |                                          
           |  +----------- Brachyanodon_impura        
           |  |                                       
           +--+  +-------- Brachyanodon               
              |  |                                    
              +--+  +----- Brachyanodon_nuttalliana__1
                 |  |                                 
                 +--+----- Brachyanodon_nuttalliana__2
                    |                                 
                    |  +-- Brachyanodon_californiensis
                    +--+                              
                       +-- Brachyanodon_oregonensis__1

Lopes-Lima et al., 2021 [Publ 4217], fig. 11

iTtTbiriSoTroTroSiD
4 4 2 4 0 0 0 10 
  +-------- Anodonta_anatina **   
--+                            
  |  +----- Anodonta_exulcerata **
  +--+                         
     |  +-- Anodonta_seddonae **  
     +--+                      
        +-- Anodonta_cygnea **

Lyubas et al., 2022 [Publ 4255], fig. S1

iTtTbiriSoTroTroSiD
4 5 3 4 27 1 1 9 
  +----------- Anodonta_anatina **   
--+                               
  |  +-------- Pseudanodonta      
  +--+                            
     |  +----- Anodonta_exulcerata **
     +--+                         
        |  +-- Anodonta_seddonae **  
        +--+                      
           +-- Anodonta_cygnea **

Lyubas et al., 2022 [Publ 4255], fig. 7

iTtTbiriSoTroTroSiD
4 4 2 4 27 1 1 8 
  +----------- Pseudanodonta      
--+                               
  |  +-------- Anodonta_anatina **   
  +--+                            
     |  +----- Anodonta_cygnea **    
     +--+                         
        |  +-- Anodonta_exulcerata **
        +--+                      
           +-- Anodonta_seddonae **

Nagel et al., 1998 [Publ 1946], fig. 7

iTtTbiriSoTroTroSiD
3 4 3 4 2 1 1 12 
  +-------------- Pseudanodonta   
--+                               
  |  +----------- Anodonta_cygnea ** 
  +--+                            
     |  +-------- Sinanodonta     
     +--+                         
        |  +----- Anodonta__2 **     
        +--+                      
           |  +-- Anodonta_anatina **
           +--+                   
              +-- Anodonta__1 **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
3 13 11 3 138 2 2 18 
                 +-- Anodonta_cygnea **    
  +--------------+                      
  |              +-- Anodonta_exulcerata **
  |                                     
--+  +-------------- Anodonta_anatina **   
  |  |                                  
  |  |           +-- Brachyanodon       
  +--+  +--------+                      
     |  |        +-- Pseudanodonta      
     |  |                               
     +--+        +-- Pyganodon          
        |  +-----+                      
        |  |     +-- Utterbackia        
        |  |                            
        +--+     +-- Pterosyna          
           |  +--+                      
           |  |  +-- Prolasmidonta      
           +--+                         
              |  +-- Alasmidonta        
              +--+                      
                 +-- Platynaias

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
3 3 1 3 93 2 2 9 
        +-- Brachyanodon       
  +-----+                      
  |     +-- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
3 3 1 3 9 0 0 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_exulcerata **
  +--+                      
     +-- Anodonta_cygnea **

Froufe et al., 2017 [Publ 3802], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 1 0 0 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_cygnea **    
  +--+                      
     +-- Anodonta_exulcerata **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
3 9 7 3 25 4 1 15 
           +-- Anodonta_exulcerata **        
  +--------+                              
  |        +-- Anodonta_cygnea **            
  |                                       
--+        +-- Brachyanodon_californiensis
  |  +-----+                              
  |  |     +-- Brachyanodon_nuttalliana   
  |  |                                    
  +--+     +-- Brachyanodon_kennerlyi     
     |  +--+                              
     |  |  +-- Brachyanodon_oregonensis   
     +--+                                 
        |  +-- Anodonta_anatina **           
        +--+                              
           +-- Utterbackiana

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
3 3 1 3 89 2 2 9 
  +-------- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 92 1 1 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_cygnea **    
  +--+                      
     +-- Anodonta_exulcerata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
3 10 7 3 93 2 2 17 
  +----------------- Brachyanodon       
  |                                     
  +----------------- Pseudanodonta      
  |                                     
--+  +-------------- Anodonta_cygnea **    
  |  |                                  
  |  |           +-- Anodonta_exulcerata **
  |  +-----------+                      
  +--+           +-- Anodonta_anatina **   
     |                                  
     |           +-- Pyganodon          
     |  +--------+                      
     |  |        +-- Utterbackia        
     +--+                               
        |  +-------- Pterosyna          
        +--+                            
           |  +----- Prolasmidonta      
           +--+                         
              |  +-- Alasmidonta        
              +--+                      
                 +-- Platynaias

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
3 3 1 3 7 0 0 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_exulcerata **
  +--+                      
     +-- Anodonta_cygnea **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 3 50 2 2 9 
  +-------- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
3 4 2 3 3 1 1 10 
     +-- Anodonta_anatina **   
  +--+                      
  |  +-- Pseudanodonta      
--+                         
  |  +-- Anodonta_exulcerata **
  +--+                      
     +-- Anodonta_cygnea **

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iTtTbiriSoTroTroSiD
3 3 1 3 3 0 0 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_exulcerata **
  +--+                      
     +-- Anodonta_cygnea **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 73 2 2 9 
  +-------- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
3 3 1 3 89 2 2 9 
  +-------- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 3 51 2 2 9 
  +-------- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 22 0 0 9 
  +----- Anodonta_anatina ** 
--+                       
  |  +-- Anodonta_cygnea **  
  +--+                    
     +-- Anodonta_seddonae **

Nagel et al., 1996 [Publ 3400], fig. 351

iTtTbiriSoTroTroSiD
3 4 3 4 2 1 1 12 
  +-------------- Pseudanodonta   
--+                               
  |  +----------- Anodonta_cygnea ** 
  +--+                            
     |  +-------- Sinanodonta     
     +--+                         
        |  +----- Anodonta__2 **     
        +--+                      
           |  +-- Anodonta_anatina **
           +--+                   
              +-- Anodonta__1 **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
3 9 7 3 16 2 2 13 
  +-------------- Anodonta_anatina **   
  |                                  
  |        +----- Pseudanodonta      
--+  +-----+                         
  |  |     |  +-- Anodonta_exulcerata **
  |  |     +--+                      
  +--+        +-- Anodonta_cygnea **    
     |                               
     |  +-------- Brachyanodon       
     +--+                            
        |  +----- Platynaias         
        +--+                         
           |  +-- Pyganodon          
           +--+                      
              +-- Utterbackia

Tomilova et al., 2020 [Publ 4097], fig. 3

iTtTbiriSoTroTroSiD
3 4 2 3 5 1 1 8 
  +-------- Anodonta_anatina **   
--+                            
  |  +----- Pseudanodonta      
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
3 3 1 3 62 1 1 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_exulcerata **
  +--+                      
     +-- Anodonta_cygnea **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 3 0 2 74 2 2 9 
  +-------- Pseudanodonta   
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Brachyanodon    
     +--+                   
        +-- Anodonta_cygnea **

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
2 3 0 2 17 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
2 2 0 2 11 0 0 -- 
  +-- Anodonta_cygnea **    
--+                      
  +-- Anodonta_exulcerata **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
2 7 0 2 51 0 0 14 
  +----------- Anodonta_cygnea ** 
  |                            
--+  +-------- Anodonta_anatina **
  |  |                         
  +--+  +----- Strophitus      
     |  |                      
     +--+  +-- Platynaias      
        |  |                   
        +--+-- Alasmidonta     
           |                   
           +-- Pyganodon

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
2 37 0 2 39 2 2 47 
  +----------------------- Anodonta_cygnea ** 
  |                                        
  |                    +-- Brachyanodon    
--+  +-----------------+                   
  |  |                 +-- Pseudanodonta   
  |  |                                     
  +--+  +----------------- Anodonta_anatina **
     |  |                                  
     |  |              +-- Utterbackia     
     |  |  +-----------+                   
     +--+  |           +-- Pyganodon       
        |  |                               
        |  |     +-------- Utterbackiana   
        |  |     |                         
        +--+  +--+     +-- Lasmigona       
           |  |  |  +--+                   
           |  |  |  |  +-- Pseudodontoideus
           |  |  +--+                      
           |  |     |  +-- Pterosyna       
           |  |     +--+                   
           +--+        +-- Arcidens        
              |                            
              |     +----- Sulcularia      
              |  +--+                      
              |  |  |  +-- Anodontoides    
              |  |  +--+                   
              |  |     +-- Strophitus      
              +--+                         
                 |     +-- Prolasmidonta   
                 |  +--+                   
                 |  |  +-- Platynaias      
                 +--+                      
                    |  +-- Alasminota      
                    +--+                   
                       +-- Alasmidonta

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
2 18 0 2 16 5 2 24 
  +----------------- Anodonta_cygnea **            
  |                                             
  +----------------- Pyganodon                  
  |                                             
  +----------------- Alasminota                 
  |                                             
  +----------------- Anodonta_anatina **           
  |                                             
  +----------------- Unio                       
  |                                             
  |              +-- Brachyanodon_nuttalliana   
  +--------------+                              
  |              +-- Brachyanodon_californiensis
  |                                             
--+              +-- Beringiana                 
  +--------------+                              
  |              +-- Sinanodonta                
  |                                             
  |              +-- Pseudodontoideus           
  |              |                              
  +--------------+-- Alasmidonta                
  |              |                              
  |              +-- Pterosyna                  
  |                                             
  |  +-------------- Pseudanodonta              
  |  |                                          
  |  |        +----- Strophitus                 
  |  +--------+                                 
  +--+        |  +-- Prolasmidonta              
     |        +--+                              
     |           +-- Platynaias                 
     |                                          
     |  +----------- Brachyanodon_oregonensis__4
     +--+                                       
        |  +-------- Brachyanodon_oregonensis__3
        +--+                                    
           |  +----- Brachyanodon_oregonensis__2
           +--+                                 
              |  +-- Brachyanodon_oregonensis__1
              +--+                              
                 +-- Brachyanodon_kennerlyi

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 16 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
2 2 0 2 20 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 66 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 5a

iTtTbiriSoTroTroSiD
2 5 0 2 6 1 1 11 
  +-------- Anodonta_anatina **
--+                         
  |  +----- Sinanodonta     
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 11 1 1 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
2 3 0 2 54 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 5 1 1 11 
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Pseudanodonta

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
2 9 0 2 72 0 0 16 
  +-------------- Anodonta_cygnea ** 
  |                               
--+  +----------- Anodonta_anatina **
  |  |                            
  +--+        +-- Pyganodon       
     |  +-----+                   
     |  |     +-- Utterbackia     
     +--+                         
        |  +----- Prolasmidonta   
        +--+                      
           |  +-- Alasmidonta     
           +--+                   
              +-- Platynaias

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 21 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 3a

iTtTbiriSoTroTroSiD
2 3 0 2 7 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Pseudanodonta

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 198 2 2 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Brachyanodon

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 9 0 2 136 2 2 14 
  +-------- Anodonta_anatina **
  |                         
  +-------- Pseudanodonta   
  |                         
  |     +-- Anodonta_cygnea ** 
--+-----+                   
  |     +-- Brachyanodon    
  |                         
  |  +----- Pyganodon       
  |  |                      
  |  +----- Simpsonaias     
  +--+                      
     +----- Strophitus      
     |                      
     |  +-- Alasmidonta     
     +--+                   
        +-- Lasmigona

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 72 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 7 1 1 -- 
  +----- Pseudanodonta   
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Anodonta_cygnea **

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 10 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 66 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 12 1 1 11 
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Pseudanodonta

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 17 0 2 122 2 2 23 
  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina **     
  |                        
  +-- Anodonta_cygnea **      
  |                        
  +-- Brachyanodon         
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta    
  |                        
  +-- Cristaria_plicata    
  |                        
  +-- Cristaria_radiata    
  |                        
  +-- Pletholophus         
  |                        
  +-- Sinanodonta_lucida   
  |                        
  +-- Sinanodonta_jourdyi  
  |                        
  +-- Lanceolaria_gladiola 
  |                        
  +-- Lanceolaria_grayii__1
  |                        
  +-- Lanceolaria_grayii__2

Soroka, 2010 [Publ 3404], fig. 8b

iTtTbiriSoTroTroSiD
2 2 0 2 4 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
2 2 0 2 20 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
2 6 0 2 54 0 0 13 
  +----------- Anodonta_cygnea ** 
--+                            
  |  +-------- Anodonta_anatina **
  +--+                         
     |  +----- Lasmigona       
     +--+                      
        |  +-- Pyganodon       
        +--+                   
           +-- Utterbackia

Soroka, 2010 [Publ 3404], fig. 3b

iTtTbiriSoTroTroSiD
2 9 0 2 7 1 1 15 
  +----- Anodonta_anatina **
  |                      
  +----- Anodonta_cygnea ** 
--+                      
  +----- Pseudanodonta   
  |                      
  |  +-- Sinanodonta     
  +--+                   
     +-- Unio

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
2 3 0 2 196 2 2 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Brachyanodon

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
2 3 0 2 120 2 2 11 
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Brachyanodon

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 52 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
2 3 0 2 34 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Mezhzherin et al., 2014 [Publ 3705], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 2 1 1 -- 
  +----- Pseudanodonta   
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 8a

iTtTbiriSoTroTroSiD
2 2 0 2 4 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
2 6 0 2 89 1 1 14 
  +----------- Anodonta_cygnea ** 
  |                            
--+        +-- Lasmigona       
  |  +-----+                   
  |  |     +-- Anodonta_anatina **
  +--+                         
     |  +----- Pseudanodonta   
     +--+                      
        |  +-- Pyganodon       
        +--+                   
           +-- Strophitus

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
2 8 0 2 23 0 0 17 
  +----------------- Anodonta_cygnea ** 
  |                                  
--+  +-------------- Unio_pictorum__2
  |  |                               
  +--+           +-- Unio_pictorum__1
     |  +--------+                   
     |  |        +-- Unio_caffer     
     +--+                            
        |  +-------- Anodonta **        
        |  |                         
        +--+  +----- Lasmigona       
           |  |                      
           +--+  +-- Strophitus      
              |  |                   
              +--+-- Alasmidonta     
                 |                   
                 +-- Pyganodon

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 38 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

iTtTbiriSoTroTroSiD
2 2 0 2 34 1 1 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 1 1 1 -- 
  +----- Pseudanodonta   
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Anodonta_cygnea **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
2 47 0 2 198 2 2 52 
  +-------- Acuticosta           
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina **     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea **      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Brachyanodon         
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesma          
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Lanceolaria          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unio                 
  |                              
  +-------- Scriptolamprotula    
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Froufe et al., 2014 [Publ 3592], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 1 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
2 9 0 2 72 0 0 16 
  +-------------- Anodonta_cygnea ** 
  |                               
--+  +----------- Anodonta_anatina **
  |  |                            
  +--+        +-- Pyganodon       
     |  +-----+                   
     |  |     +-- Utterbackia     
     +--+                         
        |  +----- Prolasmidonta   
        +--+                      
           |  +-- Alasmidonta     
           +--+                   
              +-- Platynaias

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iTtTbiriSoTroTroSiD
2 3 0 2 5 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 2 74 2 2 -- 
  +-------- Pseudanodonta   
--+                         
  |  +----- Brachyanodon    
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Anodonta_cygnea **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 5b

iTtTbiriSoTroTroSiD
2 8 0 2 6 1 1 13 
  +-------- Anodonta_anatina **
  |                         
--+  +----- Sinanodonta     
  |  |                      
  +--+----- Unio            
     |                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 66 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 37 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Källersjö et al., 2005 [Publ 2684], fig. 1A

iTtTbiriSoTroTroSiD
2 3 0 2 5 1 1 10 
  +-- Anodonta_anatina **
  |                   
--+-- Pseudanodonta   
  |                   
  +-- Anodonta_cygnea **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
2 3 0 2 78 2 2 10 
  +----- Pseudanodonta   
  |                      
--+  +-- Anodonta_anatina **
  |  |                   
  +--+-- Anodonta_cygnea ** 
     |                   
     +-- Brachyanodon

Whelan et al., 2023 [Publ 4287], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 25 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
2 8 0 2 67 0 0 15 
  +-------------- Anodonta_cygnea ** 
  |                               
--+  +----------- Anodonta_anatina **
  |  |                            
  +--+        +-- Pyganodon       
     |  +-----+                   
     |  |     +-- Utterbackia     
     +--+                         
        |  +----- Prolasmidonta   
        +--+                      
           |  +-- Platynaias      
           +--+                   
              +-- Alasmidonta

Soroka, 2010 [Publ 3404], fig. 4

iTtTbiriSoTroTroSiD
2 7 0 2 5 1 1 12 
        +-- Anodonta_anatina **
  +-----+                   
  |     +-- Unio            
--+                         
  |  +----- Sinanodonta     
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 86 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Mezhzherin et al., 2014 [Publ 3705], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 2 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Pseudanodonta

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
2 3 0 2 15 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 9 0 2 136 2 2 13 
  +-------------- Anodonta_anatina **
  |                               
  |        +----- Pseudanodonta   
--+  +-----+                      
  |  |     |  +-- Anodonta_cygnea ** 
  |  |     +--+                   
  |  |        +-- Brachyanodon    
  +--+                            
     |        +-- Lasmigona       
     |  +-----+                   
     |  |     +-- Alasmidonta     
     +--+                         
        |  +----- Simpsonaias     
        +--+                      
           |  +-- Pyganodon       
           +--+                   
              +-- Strophitus

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 84 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Klishko et al., 2019 [Publ 3886], fig. 9

iTtTbiriSoTroTroSiD
2 2 0 2 9 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Hinzmann et al., 2013 [Publ 3541], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 1 1 1 -- 
  +----- Pseudanodonta   
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Anodonta_cygnea **

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 15 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
2 3 0 2 13 1 1 10 
  +-- Anodonta_anatina **
  |                   
--+-- Anodonta_cygnea ** 
  |                   
  +-- Pseudanodonta

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
2 2 0 2 89 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Zieritz et al., 2012 [Publ 3673], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 6 1 1 10 
  +-- Pseudanodonta   
  |                   
--+-- Anodonta_anatina **
  |                   
  +-- Anodonta_cygnea **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
2 2 0 2 37 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Anodonta_anatina **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 0 0 -- 
-- Anodonta_anatina **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 43 0 0 -- 
-- Anodonta_anatina **

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Anodonta **     
--+                
  +-- Pseudanodonta

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Anodonta_cygnea **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Huff et al., 2004 [Publ 2697], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta **

Soroka, 2010 [Publ 3404], fig. 3b-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Anodonta_anatina **

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Anodonta_anatina **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta_cygnea **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_cygnea **

Huff et al., 2004 [Publ 2697], fig. 5-6

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Anodonta **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 81 0 0 -- 
-- Anodonta_anatina **

Lydeard et al., 2000 [Publ 2654], fig. 2d

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_cygnea **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Anodonta_anatina **

Lydeard et al., 2000 [Publ 2654], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Anodonta_anatina **

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Anodonta_anatina **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta_cygnea **

Mock et al., 2004 [Publ 2686], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Anodonta **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta_cygnea **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 0 0 -- 
-- Anodonta_anatina **

Soroka, 2008 [Publ 3695], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Anodonta_anatina **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 43 0 0 -- 
-- Anodonta_anatina **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 5a-M

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Anodonta_anatina **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_anatina **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Anodonta_anatina **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_anatina **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Anodonta_anatina **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Anodonta_anatina **

Lydeard et al., 2000 [Publ 2654], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Chase et al., 2018 [Publ 3847], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_anatina **

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
-- Anodonta_cygnea **

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_anatina **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 101 1 1 -- 
  +-- Anodonta_cygnea **
--+                  
  +-- Brachyanodon

Wen et al., 2017 [Publ 3795], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Anodonta_anatina **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 0 0 -- 
-- Anodonta_cygnea **

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_anatina **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta_anatina **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Anodonta_anatina **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 0 0 -- 
-- Anodonta_anatina **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 3a-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Anodonta_anatina **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Anodonta_anatina **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 0 0 -- 
-- Anodonta_anatina **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_anatina **

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Anodonta_cygnea **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Anodonta_anatina **

Lydeard et al., 2000 [Publ 2654], fig. 2c

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_anatina **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta_cygnea **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 4-M

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Anodonta_anatina **

Huff et al., 2004 [Publ 2697], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Anodonta **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Anodonta_anatina **

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Anodonta_anatina **

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Anodonta_cygnea **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 31 0 0 -- 
-- Anodonta_anatina **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Anodonta_anatina **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Anodonta_anatina **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_anatina **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_anatina **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta **

Taylor et al., 2007 [Publ 2696], fig. 1-3

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Anodonta_cygnea **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Anodonta_anatina **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Anodonta_cygnea **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Anodonta_cygnea **

Bogatov et al., 2018 [Publ 3859], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Huff et al., 2004 [Publ 2697], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Anodonta **

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 32 0 0 -- 
-- Anodonta **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Anodonta_cygnea **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Anodonta_cygnea **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta_anatina **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Anodonta_anatina **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_anatina **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_anatina **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Anodonta_anatina **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta **

Mock et al., 2004 [Publ 2686], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Anodonta **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Anodonta_anatina **

Soroka, 2008 [Publ 3695], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Anodonta_anatina **

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

iTtTbiriSoTroTroSiD
1 1 0 1 4 1 1 -- 
  +-- Pseudanodonta   
--+                   
  +-- Anodonta_anatina **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 101 1 1 -- 
-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 5b-M

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Anodonta_anatina **

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Anodonta_cygnea **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Anodonta_anatina **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Anodonta_anatina **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Anodonta_anatina **

Whelan et al., 2023 [Publ 4287], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Anodonta_anatina **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_cygnea **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_anatina **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Anodonta_anatina **

Bogatov et al., 2018 [Publ 3859], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Anodonta_cygnea **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 0 0 -- 
-- Anodonta_anatina **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
-- Anodonta_cygnea **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Anodonta_anatina **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_anatina **

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Anodonta **

Araujo et al., 2018 [Publ 3779], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Anodonta_anatina **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Anodonta_cygnea **

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Anodonta_cygnea **