Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Anodontina

genus Anodonta Lamarck, 1799: cladomics

5 Recent valid species (18 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (18 total)
analyzed in 216 cladograms in 117 publications
maximum number of terminals in an analysis = 4 (mean = 1.56, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 91
maximum number of included terminals = 4, (mean = 2.33 median = 2)
proportion of cladograms that support basic monophyly = 0.49

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Anodonta cladogram distances
PCoA Ordination

NJ Tree of Anodonta cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 10 9 4 626 6 2 0 
  +----------------------- Anodonta_anatina **           
  |                                                   
--+  +-------------------- Pseudanodonta              
  |  |                                                
  +--+  +----------------- Brachyanodon_oregonensis__2
     |  |                                             
     |  |           +----- Anodonta_cygnea **            
     +--+  +--------+                                 
        |  |        |  +-- Anodonta_seddonae **          
        |  |        +--+                              
        +--+           +-- Anodonta_exulcerata **        
           |                                          
           |  +----------- Brachyanodon_impura        
           |  |                                       
           +--+  +-------- Brachyanodon               
              |  |                                    
              +--+  +----- Brachyanodon_nuttalliana__1
                 |  |                                 
                 +--+----- Brachyanodon_nuttalliana__2
                    |                                 
                    |  +-- Brachyanodon_californiensis
                    +--+                              
                       +-- Brachyanodon_oregonensis__1

Lopes-Lima et al., 2021 [Publ 4217], fig. 11

iTtTbiriSoTroTroSiD
4 4 2 4 0 0 0 10 
  +-------- Anodonta_anatina **   
--+                            
  |  +----- Anodonta_exulcerata **
  +--+                         
     |  +-- Anodonta_seddonae **  
     +--+                      
        +-- Anodonta_cygnea **

Lyubas et al., 2022 [Publ 4255], fig. S1

iTtTbiriSoTroTroSiD
4 5 3 4 27 1 1 9 
  +----------- Anodonta_anatina **   
--+                               
  |  +-------- Pseudanodonta      
  +--+                            
     |  +----- Anodonta_exulcerata **
     +--+                         
        |  +-- Anodonta_seddonae **  
        +--+                      
           +-- Anodonta_cygnea **

Lyubas et al., 2022 [Publ 4255], fig. 7

iTtTbiriSoTroTroSiD
4 4 2 4 27 1 1 8 
  +----------- Pseudanodonta      
--+                               
  |  +-------- Anodonta_anatina **   
  +--+                            
     |  +----- Anodonta_cygnea **    
     +--+                         
        |  +-- Anodonta_exulcerata **
        +--+                      
           +-- Anodonta_seddonae **

Tomilova et al., 2020 [Publ 4097], fig. 3

iTtTbiriSoTroTroSiD
3 4 2 3 5 1 1 8 
  +-------- Anodonta_anatina **   
--+                            
  |  +----- Pseudanodonta      
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iTtTbiriSoTroTroSiD
3 3 1 3 3 0 0 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_exulcerata **
  +--+                      
     +-- Anodonta_cygnea **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
3 9 7 3 16 2 2 13 
  +-------------- Anodonta_anatina **   
  |                                  
  |        +----- Pseudanodonta      
--+  +-----+                         
  |  |     |  +-- Anodonta_exulcerata **
  |  |     +--+                      
  +--+        +-- Anodonta_cygnea **    
     |                               
     |  +-------- Brachyanodon       
     +--+                            
        |  +----- Platynaias         
        +--+                         
           |  +-- Pyganodon          
           +--+                      
              +-- Utterbackia

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
3 3 1 3 92 2 2 9 
        +-- Brachyanodon       
  +-----+                      
  |     +-- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
3 4 2 3 3 1 1 10 
     +-- Anodonta_anatina **   
  +--+                      
  |  +-- Pseudanodonta      
--+                         
  |  +-- Anodonta_exulcerata **
  +--+                      
     +-- Anodonta_cygnea **

Froufe et al., 2017 [Publ 3802], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 1 0 0 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_cygnea **    
  +--+                      
     +-- Anodonta_exulcerata **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
3 13 11 3 137 2 2 18 
                 +-- Anodonta_cygnea **    
  +--------------+                      
  |              +-- Anodonta_exulcerata **
  |                                     
--+  +-------------- Anodonta_anatina **   
  |  |                                  
  |  |           +-- Brachyanodon       
  +--+  +--------+                      
     |  |        +-- Pseudanodonta      
     |  |                               
     +--+        +-- Pyganodon          
        |  +-----+                      
        |  |     +-- Utterbackia        
        |  |                            
        +--+     +-- Pterosyna          
           |  +--+                      
           |  |  +-- Prolasmidonta      
           +--+                         
              |  +-- Alasmidonta        
              +--+                      
                 +-- Platynaias

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
3 9 7 3 25 4 1 15 
           +-- Anodonta_exulcerata **        
  +--------+                              
  |        +-- Anodonta_cygnea **            
  |                                       
--+        +-- Brachyanodon_californiensis
  |  +-----+                              
  |  |     +-- Brachyanodon_nuttalliana   
  |  |                                    
  +--+     +-- Brachyanodon_kennerlyi     
     |  +--+                              
     |  |  +-- Brachyanodon_oregonensis   
     +--+                                 
        |  +-- Anodonta_anatina **           
        +--+                              
           +-- Utterbackiana

Nagel et al., 1996 [Publ 3400], fig. 351

iTtTbiriSoTroTroSiD
3 4 3 4 2 1 1 12 
  +-------------- Pseudanodonta   
--+                               
  |  +----------- Anodonta_cygnea ** 
  +--+                            
     |  +-------- Sinanodonta     
     +--+                         
        |  +----- Anodonta__2 **     
        +--+                      
           |  +-- Anodonta_anatina **
           +--+                   
              +-- Anodonta__1 **

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
3 3 1 3 7 0 0 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_exulcerata **
  +--+                      
     +-- Anodonta_cygnea **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 92 1 1 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_cygnea **    
  +--+                      
     +-- Anodonta_exulcerata **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
3 3 1 3 61 1 1 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_exulcerata **
  +--+                      
     +-- Anodonta_cygnea **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 3 50 2 2 9 
  +-------- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 3 49 2 2 9 
  +-------- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 22 0 0 9 
  +----- Anodonta_anatina ** 
--+                       
  |  +-- Anodonta_cygnea **  
  +--+                    
     +-- Anodonta_seddonae **

Wu et al., 2026 [Publ 4825], fig. 6B

iTtTbiriSoTroTroSiD
3 3 1 3 62 1 1 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_cygnea **    
  +--+                      
     +-- Anodonta_exulcerata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
3 10 7 3 92 2 2 17 
  +----------------- Brachyanodon       
  |                                     
  +----------------- Pseudanodonta      
  |                                     
--+  +-------------- Anodonta_cygnea **    
  |  |                                  
  |  |           +-- Anodonta_exulcerata **
  |  +-----------+                      
  +--+           +-- Anodonta_anatina **   
     |                                  
     |           +-- Pyganodon          
     |  +--------+                      
     |  |        +-- Utterbackia        
     +--+                               
        |  +-------- Pterosyna          
        +--+                            
           |  +----- Prolasmidonta      
           +--+                         
              |  +-- Alasmidonta        
              +--+                      
                 +-- Platynaias

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
3 3 1 3 88 2 2 9 
  +-------- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Nagel et al., 1998 [Publ 1946], fig. 7

iTtTbiriSoTroTroSiD
3 4 3 4 2 1 1 12 
  +-------------- Pseudanodonta   
--+                               
  |  +----------- Anodonta_cygnea ** 
  +--+                            
     |  +-------- Sinanodonta     
     +--+                         
        |  +----- Anodonta__2 **     
        +--+                      
           |  +-- Anodonta_anatina **
           +--+                   
              +-- Anodonta__1 **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 72 2 2 9 
  +-------- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
3 3 1 3 88 2 2 9 
  +-------- Pseudanodonta      
--+                            
  |  +----- Anodonta_anatina **   
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
3 3 1 3 9 0 0 9 
  +----- Anodonta_anatina **   
--+                         
  |  +-- Anodonta_exulcerata **
  +--+                      
     +-- Anodonta_cygnea **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 21 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
2 8 0 2 66 0 0 15 
  +-------------- Anodonta_cygnea ** 
  |                               
--+  +----------- Anodonta_anatina **
  |  |                            
  +--+        +-- Pyganodon       
     |  +-----+                   
     |  |     +-- Utterbackia     
     +--+                         
        |  +----- Prolasmidonta   
        +--+                      
           |  +-- Platynaias      
           +--+                   
              +-- Alasmidonta

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
2 2 0 2 36 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
2 3 0 2 13 1 1 10 
  +-- Anodonta_anatina **
  |                   
--+-- Anodonta_cygnea ** 
  |                   
  +-- Pseudanodonta

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
2 2 0 2 11 0 0 -- 
  +-- Anodonta_cygnea **    
--+                      
  +-- Anodonta_exulcerata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
2 3 0 2 120 2 2 11 
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Brachyanodon

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 85 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 4

iTtTbiriSoTroTroSiD
2 7 0 2 5 1 1 12 
        +-- Anodonta_anatina **
  +-----+                   
  |     +-- Unio            
--+                         
  |  +----- Sinanodonta     
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 18 0 2 123 2 2 23 
  +-- Alasmidonta             
  |                           
  +-- Anodonta_anatina **        
  |                           
  +-- Anodonta_cygnea **         
  |                           
  +-- Brachyanodon            
  |                           
  +-- Lasmigona               
  |                           
  +-- Pseudanodonta           
  |                           
  +-- Pyganodon               
  |                           
  +-- Simpsonaias             
  |                           
  +-- Strophitus              
--+                           
  +-- Sinanodonta_lauta       
  |                           
  +-- Cristaria_plicata       
  |                           
  +-- Cristaria_radiata       
  |                           
  +-- Pletholophus            
  |                           
  +-- Sinanodonta_lucida      
  |                           
  +-- Sinanodonta_jourdyi     
  |                           
  +-- Lanceolaria_gladiola    
  |                           
  +-- Lanceolaria_grayii      
  |                           
  +-- Lanceolaria_danzhouensis

Mezhzherin et al., 2014 [Publ 3705], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 2 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Pseudanodonta

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
2 48 0 2 199 2 2 53 
  +-------- Acuticosta           
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina **     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea **      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Brachyanodon         
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesma          
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unio                 
  |                              
  +-------- Scriptolamprotula    
  |                              
  +-------- Lanceolaria          
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
2 3 0 2 197 2 2 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Brachyanodon

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
2 3 0 2 17 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 5a

iTtTbiriSoTroTroSiD
2 5 0 2 6 1 1 11 
  +-------- Anodonta_anatina **
--+                         
  |  +----- Sinanodonta     
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 16 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 83 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
2 3 0 2 79 2 2 10 
  +----- Pseudanodonta   
  |                      
--+  +-- Anodonta_anatina **
  |  |                   
  +--+-- Anodonta_cygnea ** 
     |                   
     +-- Brachyanodon

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 52 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
2 3 0 2 34 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 9 0 2 137 2 2 14 
  +-------- Anodonta_anatina **
  |                         
  +-------- Pseudanodonta   
  |                         
  |     +-- Anodonta_cygnea ** 
--+-----+                   
  |     +-- Brachyanodon    
  |                         
  |  +----- Pyganodon       
  |  |                      
  |  +----- Simpsonaias     
  +--+                      
     +----- Strophitus      
     |                      
     |  +-- Alasmidonta     
     +--+                   
        +-- Lasmigona

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 3 0 2 74 2 2 9 
  +-------- Pseudanodonta   
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Brachyanodon    
     +--+                   
        +-- Anodonta_cygnea **

Zieritz et al., 2012 [Publ 3673], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 6 1 1 10 
  +-- Pseudanodonta   
  |                   
--+-- Anodonta_anatina **
  |                   
  +-- Anodonta_cygnea **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
2 7 0 2 51 0 0 14 
  +----------- Anodonta_cygnea ** 
  |                            
--+  +-------- Anodonta_anatina **
  |  |                         
  +--+  +----- Strophitus      
     |  |                      
     +--+  +-- Platynaias      
        |  |                   
        +--+-- Alasmidonta     
           |                   
           +-- Pyganodon

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
2 9 0 2 71 0 0 16 
  +-------------- Anodonta_cygnea ** 
  |                               
--+  +----------- Anodonta_anatina **
  |  |                            
  +--+        +-- Pyganodon       
     |  +-----+                   
     |  |     +-- Utterbackia     
     +--+                         
        |  +----- Prolasmidonta   
        +--+                      
           |  +-- Alasmidonta     
           +--+                   
              +-- Platynaias

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 38 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 1 1 1 -- 
  +----- Pseudanodonta   
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Anodonta_cygnea **

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
2 18 0 2 16 5 2 24 
  +----------------- Anodonta_cygnea **            
  |                                             
  +----------------- Pyganodon                  
  |                                             
  +----------------- Alasminota                 
  |                                             
  +----------------- Anodonta_anatina **           
  |                                             
  +----------------- Unio                       
  |                                             
  |              +-- Brachyanodon_nuttalliana   
  +--------------+                              
  |              +-- Brachyanodon_californiensis
  |                                             
--+              +-- Beringiana                 
  +--------------+                              
  |              +-- Sinanodonta                
  |                                             
  |              +-- Pseudodontoideus           
  |              |                              
  +--------------+-- Alasmidonta                
  |              |                              
  |              +-- Pterosyna                  
  |                                             
  |  +-------------- Pseudanodonta              
  |  |                                          
  |  |        +----- Strophitus                 
  |  +--------+                                 
  +--+        |  +-- Prolasmidonta              
     |        +--+                              
     |           +-- Platynaias                 
     |                                          
     |  +----------- Brachyanodon_oregonensis__4
     +--+                                       
        |  +-------- Brachyanodon_oregonensis__3
        +--+                                    
           |  +----- Brachyanodon_oregonensis__2
           +--+                                 
              |  +-- Brachyanodon_oregonensis__1
              +--+                              
                 +-- Brachyanodon_kennerlyi

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
2 8 0 2 23 0 0 17 
  +----------------- Anodonta_cygnea ** 
  |                                  
--+  +-------------- Unio_pictorum__2
  |  |                               
  +--+           +-- Unio_pictorum__1
     |  +--------+                   
     |  |        +-- Unio_caffer     
     +--+                            
        |  +-------- Anodonta **        
        |  |                         
        +--+  +----- Lasmigona       
           |  |                      
           +--+  +-- Strophitus      
              |  |                   
              +--+-- Alasmidonta     
                 |                   
                 +-- Pyganodon

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 2 74 2 2 -- 
  +-------- Pseudanodonta   
--+                         
  |  +----- Brachyanodon    
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Anodonta_cygnea **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 67 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 8a

iTtTbiriSoTroTroSiD
2 2 0 2 4 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
2 9 0 2 71 0 0 16 
  +-------------- Anodonta_cygnea ** 
  |                               
--+  +----------- Anodonta_anatina **
  |  |                            
  +--+        +-- Pyganodon       
     |  +-----+                   
     |  |     +-- Utterbackia     
     +--+                         
        |  +----- Prolasmidonta   
        +--+                      
           |  +-- Alasmidonta     
           +--+                   
              +-- Platynaias

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 15 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 67 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 10 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 71 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
2 3 0 2 12 1 1 11 
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Pseudanodonta

Whelan et al., 2023 [Publ 4287], fig. 2

iTtTbiriSoTroTroSiD
2 37 0 2 39 2 2 47 
  +----------------------- Anodonta_cygnea ** 
  |                                        
  |                    +-- Brachyanodon    
--+  +-----------------+                   
  |  |                 +-- Pseudanodonta   
  |  |                                     
  +--+  +----------------- Anodonta_anatina **
     |  |                                  
     |  |              +-- Utterbackia     
     |  |  +-----------+                   
     +--+  |           +-- Pyganodon       
        |  |                               
        |  |     +-------- Utterbackiana   
        |  |     |                         
        +--+  +--+     +-- Lasmigona       
           |  |  |  +--+                   
           |  |  |  |  +-- Pseudodontoideus
           |  |  +--+                      
           |  |     |  +-- Pterosyna       
           |  |     +--+                   
           +--+        +-- Arcidens        
              |                            
              |     +----- Sulcularia      
              |  +--+                      
              |  |  |  +-- Anodontoides    
              |  |  +--+                   
              |  |     +-- Strophitus      
              +--+                         
                 |     +-- Prolasmidonta   
                 |  +--+                   
                 |  |  +-- Platynaias      
                 +--+                      
                    |  +-- Alasminota      
                    +--+                   
                       +-- Alasmidonta

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
2 2 0 2 89 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Klishko et al., 2019 [Publ 3886], fig. 9

iTtTbiriSoTroTroSiD
2 2 0 2 9 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

iTtTbiriSoTroTroSiD
2 2 0 2 34 1 1 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
2 6 0 2 54 0 0 13 
  +----------- Anodonta_cygnea ** 
--+                            
  |  +-------- Anodonta_anatina **
  +--+                         
     |  +----- Lasmigona       
     +--+                      
        |  +-- Pyganodon       
        +--+                   
           +-- Utterbackia

Soroka, 2010 [Publ 3404], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 5 1 1 11 
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Pseudanodonta

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
2 2 0 2 20 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
2 6 0 2 89 1 1 14 
  +----------- Anodonta_cygnea ** 
  |                            
--+        +-- Lasmigona       
  |  +-----+                   
  |  |     +-- Anodonta_anatina **
  +--+                         
     |  +----- Pseudanodonta   
     +--+                      
        |  +-- Pyganodon       
        +--+                   
           +-- Strophitus

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 9 0 2 137 2 2 13 
  +-------------- Anodonta_anatina **
  |                               
  |        +----- Pseudanodonta   
--+  +-----+                      
  |  |     |  +-- Anodonta_cygnea ** 
  |  |     +--+                   
  |  |        +-- Brachyanodon    
  +--+                            
     |        +-- Lasmigona       
     |  +-----+                   
     |  |     +-- Alasmidonta     
     +--+                         
        |  +----- Simpsonaias     
        +--+                      
           |  +-- Pyganodon       
           +--+                   
              +-- Strophitus

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
2 2 0 2 20 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
2 3 0 2 15 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 199 2 2 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Brachyanodon

Soroka, 2010 [Publ 3404], fig. 3b

iTtTbiriSoTroTroSiD
2 9 0 2 7 1 1 15 
  +----- Anodonta_anatina **
  |                      
  +----- Anodonta_cygnea ** 
--+                      
  +----- Pseudanodonta   
  |                      
  |  +-- Sinanodonta     
  +--+                   
     +-- Unio

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iTtTbiriSoTroTroSiD
2 3 0 2 5 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Whelan et al., 2023 [Publ 4287], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 25 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Hinzmann et al., 2013 [Publ 3541], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 1 1 1 -- 
  +----- Pseudanodonta   
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Anodonta_cygnea **

Källersjö et al., 2005 [Publ 2684], fig. 1A

iTtTbiriSoTroTroSiD
2 3 0 2 5 1 1 10 
  +-- Anodonta_anatina **
  |                   
--+-- Pseudanodonta   
  |                   
  +-- Anodonta_cygnea **

Froufe et al., 2014 [Publ 3592], fig. 2

iTtTbiriSoTroTroSiD
2 3 0 2 1 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 3a

iTtTbiriSoTroTroSiD
2 3 0 2 7 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Pseudanodonta

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 7 1 1 -- 
  +----- Pseudanodonta   
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Anodonta_cygnea **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
2 3 0 2 54 1 1 9 
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta   
  +--+                   
     +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 8b

iTtTbiriSoTroTroSiD
2 2 0 2 4 0 0 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Anodonta_cygnea **

Mezhzherin et al., 2014 [Publ 3705], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 2 1 1 -- 
  +----- Pseudanodonta   
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Anodonta_cygnea **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 67 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 11 1 1 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 37 0 0 -- 
  +-- Anodonta_cygnea ** 
--+                   
  +-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 5b

iTtTbiriSoTroTroSiD
2 8 0 2 6 1 1 13 
  +-------- Anodonta_anatina **
  |                         
--+  +----- Sinanodonta     
  |  |                      
  +--+----- Unio            
     |                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta_anatina **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 0 0 -- 
-- Anodonta_anatina **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Anodonta_cygnea **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_anatina **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 31 0 0 -- 
-- Anodonta_anatina **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_cygnea **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
-- Anodonta_cygnea **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Anodonta_anatina **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Anodonta_anatina **

Wen et al., 2017 [Publ 3795], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Anodonta_anatina **

Mock et al., 2004 [Publ 2686], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Anodonta **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Anodonta_anatina **

Soroka, 2008 [Publ 3695], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Anodonta_anatina **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 4-M

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Anodonta_anatina **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_anatina **

Lydeard et al., 2000 [Publ 2654], fig. 2d

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

iTtTbiriSoTroTroSiD
1 1 0 1 4 1 1 -- 
  +-- Pseudanodonta   
--+                   
  +-- Anodonta_anatina **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 43 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Anodonta_cygnea **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Anodonta_anatina **

Whelan et al., 2023 [Publ 4287], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Anodonta_anatina **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Anodonta_cygnea **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 3b-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Anodonta_anatina **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Anodonta_anatina **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Anodonta **

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Anodonta_cygnea **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Anodonta_anatina **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 47 0 0 -- 
-- Anodonta_anatina **

Lydeard et al., 2000 [Publ 2654], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 43 0 0 -- 
-- Anodonta_anatina **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Anodonta_cygnea **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Anodonta_anatina **

Huff et al., 2004 [Publ 2697], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta **

Huff et al., 2004 [Publ 2697], fig. 5-6

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Anodonta **

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Anodonta_cygnea **

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Anodonta_anatina **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Soroka, 2010 [Publ 3404], fig. 3a-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Anodonta_anatina **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Anodonta_anatina **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_anatina **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta_cygnea **

Bogatov et al., 2018 [Publ 3859], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_anatina **

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
-- Anodonta_cygnea **

Chase et al., 2018 [Publ 3847], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_anatina **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Anodonta_anatina **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 43 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Anodonta_anatina **

Bogatov et al., 2018 [Publ 3859], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Mock et al., 2004 [Publ 2686], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Anodonta **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 43 0 0 -- 
-- Anodonta_anatina **

Taylor et al., 2007 [Publ 2696], fig. 1-3

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Anodonta_cygnea **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Anodonta_anatina **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_anatina **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Anodonta_anatina **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Anodonta_anatina **
--+                   
  +-- Pseudanodonta

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Anodonta_anatina **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Anodonta_anatina **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Anodonta_anatina **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Anodonta_anatina **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_anatina **

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 5a-M

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Anodonta_anatina **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Anodonta_cygnea **

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_cygnea **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 81 0 0 -- 
-- Anodonta_anatina **

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Anodonta_anatina **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Anodonta_anatina **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Anodonta_anatina **

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_anatina **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 101 1 1 -- 
-- Anodonta_cygnea **

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Anodonta_anatina **

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 5 1 1 -- 
  +-- Anodonta **     
--+                
  +-- Pseudanodonta

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta_cygnea **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_anatina **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Anodonta_cygnea **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 0 0 -- 
-- Anodonta_anatina **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Anodonta_anatina **

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta_cygnea **

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Anodonta_cygnea **

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Anodonta_anatina **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_anatina **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Anodonta_anatina **

Huff et al., 2004 [Publ 2697], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Anodonta **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_cygnea **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 0 0 -- 
-- Anodonta_anatina **

Araujo et al., 2018 [Publ 3779], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Anodonta_anatina **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Anodonta_cygnea **

Lydeard et al., 2000 [Publ 2654], fig. 2c

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Anodonta_cygnea **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Anodonta_anatina **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 0 0 -- 
-- Anodonta_anatina **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Anodonta_anatina **

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Anodonta_anatina **

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 101 1 1 -- 
  +-- Anodonta_cygnea **
--+                  
  +-- Brachyanodon

Soroka, 2010 [Publ 3404], fig. 5b-M

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Anodonta_anatina **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 0 0 -- 
-- Anodonta_cygnea **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Anodonta_anatina **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Anodonta_cygnea **

Huff et al., 2004 [Publ 2697], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Anodonta **

Lydeard et al., 2000 [Publ 2654], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Anodonta_cygnea **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Anodonta_cygnea **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Anodonta_anatina **

Soroka, 2008 [Publ 3695], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Anodonta_anatina **

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 32 0 0 -- 
-- Anodonta **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Anodonta_anatina **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Anodonta_anatina **