Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Lanceolariini

genus Lanceolaria Conrad, 1853: cladomics

14 Recent valid species (16 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 14 (16 total)
analyzed in 109 cladograms in 65 publications
maximum number of terminals in an analysis = 10 (mean = 2.55, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 80
maximum number of included terminals = 10, (mean = 3.11 median = 3)
proportion of cladograms that support basic monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lanceolaria cladogram distances
PCoA Ordination

NJ Tree of Lanceolaria cladogram distances
NJ Tree

Cladograms

Wu et al., 2026 [Publ 4825], fig. 3

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 10 2 0 0 **
Cladomics cladogram Schistodesmus Lanceolaria oxyrhyncha Lanceolaria gladiola Lanceolaria bilirata Lanceolaria acrorrhyncha Lanceolaria grayii Lanceolaria danzhouensis Lanceolaria kihirai Lanceolaria triformis Lanceolaria lanceolata Lanceolaria helianthoides

Wu et al., 2026 [Publ 4825], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 10 10 8 10 9 0 0 2.0
Cladomics cladogram Scriptolamprotula Lanceolaria oxyrhyncha Lanceolaria bilirata Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis Lanceolaria acrorrhyncha Lanceolaria kihirai Lanceolaria triformis Lanceolaria helianthoides Lanceolaria lanceolata

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 4 0 0 4.0
Cladomics cladogram Nodularia Lanceolaria oxyrhyncha Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis Lanceolaria acrorrhyncha Lanceolaria kihirai Lanceolaria lanceolata Lanceolaria triformis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 114 0 0 4.0
Cladomics cladogram Lanceolaria oxyrhyncha Lanceolaria gladiola Lanceolaria danzhouensis Lanceolaria grayii Lanceolaria acrorrhyncha Lanceolaria kihirai Lanceolaria lanceolata Lanceolaria triformis Cristaria Anemina Amuranodonta Buldowskia Pletholophus Beringiana Sinanodonta Pseudanodonta Anodonta cygnea Anodonta anatina Brachyanodon Pyganodon Simpsonaias Strophitus Alasmidonta Lasmigona

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 7 626 0 0 5.0
Cladomics cladogram Pseudoacuticosta retiaria 2 Acuticosta chinensis 2 Acuticosta chinensis 1 Pseudoacuticosta retiaria 1 Lanceolaria oxyrhyncha Lanceolaria gladiola Lanceolaria grayii Lanceolaria acrorrhyncha Lanceolaria kihirai Lanceolaria triformis Lanceolaria lanceolata

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 7 58 0 0 3.0
Cladomics cladogram Lanceolaria bilirata Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis Lanceolaria triformis Lanceolaria helianthoides Lanceolaria lanceolata Anodonta Pseudanodonta Pyganodon Utterbackia Pterosyna Prolasmidonta Alasmidonta Platynaias Cristaria Amuranodonta Anemina Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Huang et al., 2019 [Publ 3843], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 41 0 0 5.0
Cladomics cladogram Cristaria Lanceolaria gladiola Lanceolaria lanceolata Lanceolaria triformis Lanceolaria grayii Lanceolaria danzhouensis

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 2 5 196 0 0 6.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis Lanceolaria lanceolata Lanceolaria triformis Anemina Cristaria Pletholophus Sinanodonta Alasmidonta Lasmigona Pseudanodonta Simpsonaias Strophitus Pyganodon Utterbackia Anodonta anatina Anodonta cygnea Brachyanodon

Huang et al., 2019 [Publ 3843], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 5 5 2 5 42 0 0 6.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria danzhouensis Lanceolaria grayii Lanceolaria lanceolata Lanceolaria triformis Scriptolamprotula Nodularia Tchangsinaia Cuneopsis Schistodesmus

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 194 0 0 5.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria lanceolata Lanceolaria triformis Lanceolaria grayii Lanceolaria danzhouensis Anemina Cristaria Pletholophus Sinanodonta Alasmidonta Lasmigona Anodonta anatina Anodonta cygnea Brachyanodon Pyganodon Utterbackia Pseudanodonta Simpsonaias Strophitus

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 196 0 0 5.0
Cladomics cladogram Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus Unio Scriptolamprotula Pletholophus Sinanodonta Lanceolaria gladiola Lanceolaria lanceolata Lanceolaria triformis Lanceolaria grayii Lanceolaria danzhouensis Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 77 0 0 8.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria danzhouensis Lanceolaria lanceolata Lanceolaria grayii Cristaria Anemina Pletholophus Sinanodonta Pseudanodonta Anodonta anatina Anodonta cygnea Brachyanodon Pyganodon Simpsonaias Strophitus Alasmidonta Lasmigona

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 49 0 0 6.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata Lanceolaria triformis

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 136 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis Cristaria Pletholophus Sinanodonta Anodonta anatina Pseudanodonta Anodonta cygnea Brachyanodon Lasmigona Alasmidonta Simpsonaias Pyganodon Strophitus

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 136 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria danzhouensis Lanceolaria grayii Cristaria Pletholophus Sinanodonta Anodonta anatina Pseudanodonta Anodonta cygnea Brachyanodon Pyganodon Simpsonaias Strophitus Alasmidonta Lasmigona

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.000 3 18 0 3 122 0 0 23.0
Cladomics cladogram Unio crassus Unio pictorum Unio tumidus Scriptolamprotula Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 53 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria lanceolata Lanceolaria grayii Acuticosta Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta anatina Lasmigona Pyganodon Utterbackia

Fukata & Iigo, 2019 [Publ 4108], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 14 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 51 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata Scriptolamprotula Unio Cuneopsis Schistodesmus Pseudocuneopsis Nodularia Tchangsinaia Acuticosta Cristaria Anemina Sinanodonta Anodonta Pyganodon Utterbackia Alasmidonta Prolasmidonta

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.000 3 22 5 3 31 0 0 37.0
Cladomics cladogram Sinohyriopsis Lamprotula Pronodularia Sinosolenaia Inversidens Obovalis omiensis 1 Obovalis omiensis 2 Ptychorhynchus Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula Lepidodesma Scriptolamprotula Acuticosta Cuneopsis Nodularia Schistodesmus Inversiunio

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 66 0 0 7.0
Cladomics cladogram Lanceolaria oxyrhyncha Lanceolaria lanceolata Lanceolaria grayii Lepidodesma Scriptolamprotula Unio Inversiunio Nodularia Tchangsinaia Schistodesmus

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 66 0 0 7.0
Cladomics cladogram Lanceolaria oxyrhyncha Lanceolaria lanceolata Lanceolaria grayii Lepidodesma Scriptolamprotula Unio Inversiunio Nodularia Schistodesmus Tchangsinaia

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 66 0 0 7.0
Cladomics cladogram Lanceolaria oxyrhyncha Lanceolaria lanceolata Lanceolaria grayii Scriptolamprotula Unio Nodularia Inversiunio Schistodesmus Tchangsinaia

Kim et al., 2021 [Publ 4197], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 15 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 88 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata Anemina Cristaria Sinanodonta Lasmigona Anodonta Pseudanodonta Pyganodon Strophitus

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 70 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 70 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 65 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 88 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata Lepidodesma Anemina Cristaria Sinanodonta Anodonta cygnea Lasmigona Anodonta anatina Pseudanodonta Pyganodon Strophitus

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 35 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata Acuticosta Cristaria Anemina Sinanodonta Anodonta Platynaias Pyganodon Utterbackia

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 70 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 53 0 0 7.0
Cladomics cladogram Lepidodesma Lanceolaria gladiola Lanceolaria grayii Lanceolaria triformis

Wu et al., 2023 [Publ 4296], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 20 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria lanceolata Lanceolaria grayii Pseudanodonta Lasmigona Strophitus Pyganodon Unio Scriptolamprotula Cuneopsis Nodularia Schistodesmus Tchangsinaia Inversiunio Pseudocuneopsis

Liu et al., 2023 [Publ 4310], fig. 4

cst iT tT bi riS oT roT roS iD
0.000 3 21 19 3 21 0 0 25.0
Cladomics cladogram Acuticosta Lanceolaria lanceolata Lanceolaria grayii Lanceolaria gladiola Pseudanodonta Lasmigona Strophitus Pyganodon Unio Scriptolamprotula Cuneopsis Nodularia Schistodesmus Tchangsinaia Inversiunio Pseudocuneopsis

Liu et al., 2023 [Publ 4310], fig. 5

cst iT tT bi riS oT roT roS iD
0.000 3 21 14 3 21 0 0 26.0
Cladomics cladogram Acuticosta Lanceolaria grayii Lanceolaria gladiola Lanceolaria lanceolata Lasmigona Pseudanodonta Pyganodon Strophitus Unio Scriptolamprotula Cuneopsis Nodularia Schistodesmus Tchangsinaia Inversiunio Pseudocuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 92 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 92 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 88 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 88 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 50 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata Cristaria Anemina Beringiana Sinanodonta Brachyanodon Pseudanodonta Anodonta Pyganodon Utterbackia Pterosyna Prolasmidonta Alasmidonta Platynaias

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 72 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata Cristaria Anemina Beringiana Sinanodonta Pseudanodonta Anodonta Brachyanodon Pyganodon Utterbackia Pterosyna Prolasmidonta Alasmidonta Platynaias

Xiang et al., 2025 [Publ 4433], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 10 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria lanceolata Lanceolaria grayii Anodonta Cristaria Anemina Beringiana Mimo Sinanodonta

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 84 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria lanceolata Lanceolaria grayii

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 82 0 0 7.0
Cladomics cladogram Lanceolaria gladiola Lanceolaria lanceolata Lanceolaria grayii Acuticosta Cristaria Anemina Sinanodonta Anodonta Platynaias Pyganodon Utterbackia

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 92 0 0 7.0
Cladomics cladogram Acuticosta Lanceolaria gladiola Lanceolaria grayii Lanceolaria lanceolata

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 32 0 0 --
Cladomics cladogram Lanceolaria grayii Lanceolaria lanceolata Schistodesmus Lepidodesma

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 32 0 0 --
Cladomics cladogram Strophitus Schistodesmus Lepidodesma Acuticosta Sinanodonta Cristaria Lanceolaria grayii Lanceolaria lanceolata Nodularia Tchangsinaia Utterbackia Utterbackiana Alasmidonta Lasmigona

Zhang et al., 2004 [Publ 2735], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 12 0 0 --
Cladomics cladogram Acuticosta Lanceolaria grayii Lanceolaria lanceolata

Zhou et al., 2007 [Publ 2658], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 19 0 0 --
Cladomics cladogram Acuticosta Lanceolaria triformis Lanceolaria lanceolata

Ouyang et al., 2011 [Publ 3408], fig. 1

cst iT tT bi riS oT roT roS iD
0.000 2 4 0 2 11 0 0 10.0
Cladomics cladogram Nodularia Cuneopsis Lanceolaria grayii Acuticosta Lanceolaria gladiola Scriptolamprotula

Ouyang et al., 2015 [Publ 4493], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 0 0 --
Cladomics cladogram Lanceolaria grayii Lanceolaria triformis Acuticosta Anemina Sinanodonta

Ouyang et al., 2015 [Publ 4493], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 0 0 --
Cladomics cladogram Lanceolaria grayii Lanceolaria triformis Scriptolamprotula Acuticosta Anemina Sinanodonta Nodularia Cuneopsis rufescens 2 Schistodesmus Cuneopsis celtiformis 2 Tchangsinaia Cuneopsis celtiformis 1 Cuneopsis szechenyii 2 Cuneopsis szechenyii 1 Cuneopsis rufescens 1

Xue et al., 2016 [Publ 3659], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 13 0 0 --
Cladomics cladogram Lanceolaria lanceolata Lanceolaria grayii Scriptolamprotula Schistodesmus Nodularia Unio

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 0 0 --
Cladomics cladogram Lanceolaria lanceolata Lanceolaria grayii Scriptolamprotula Unio Schistodesmus Nodularia Tchangsinaia

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 30 0 0 --
Cladomics cladogram Lanceolaria lanceolata Lanceolaria grayii Scriptolamprotula Unio Schistodesmus Tchangsinaia Nodularia

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 1 2 31 0 0 --
Cladomics cladogram Lepidodesma Lanceolaria grayii 1 Lanceolaria lanceolata Lanceolaria grayii 2

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 21 0 0 --
Cladomics cladogram Lanceolaria lanceolata Lanceolaria grayii Lepidodesma Scriptolamprotula Unio Schistodesmus Tchangsinaia Nodularia

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 80 0 0 --
Cladomics cladogram Lanceolaria gladiola Lanceolaria grayii Cristaria Pletholophus Sinanodonta Anodonta Pyganodon Simpsonaias Strophitus Alasmidonta Lasmigona

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 0 0 --
Cladomics cladogram Lanceolaria grayii Lanceolaria lanceolata Scriptolamprotula Unio Nodularia Tchangsinaia Cuneopsis

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 38 0 0 --
Cladomics cladogram Acuticosta Lanceolaria grayii Lanceolaria lanceolata

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 0 0 --
Cladomics cladogram Acuticosta Lanceolaria grayii Lanceolaria lanceolata

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 44 0 0 --
Cladomics cladogram Acuticosta Lanceolaria grayii Lanceolaria lanceolata

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 42 0 0 --
Cladomics cladogram Lanceolaria lanceolata Lanceolaria grayii Cristaria Anemina Sinanodonta Anodonta Pterosyna Pyganodon Utterbackia

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 42 0 0 --
Cladomics cladogram Lanceolaria lanceolata Lanceolaria grayii Cristaria Anemina Sinanodonta Anodonta Pterosyna Pyganodon Utterbackia

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 34 0 0 --
Cladomics cladogram Lanceolaria grayii Lanceolaria danzhouensis Anodonta anatina Pseudanodonta Anodonta cygnea Amuranodonta Anemina Buldowskia Cristaria Beringiana Sinanodonta

Riccardi et al., 2020 [Publ 4089], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 17 0 0 --
Cladomics cladogram Unio Lanceolaria grayii Lanceolaria lanceolata

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 46 0 0 --
Cladomics cladogram Acuticosta Lanceolaria lanceolata Lanceolaria grayii

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 46 0 0 --
Cladomics cladogram Lanceolaria grayii Lanceolaria lanceolata Cristaria Anemina Sinanodonta Anodonta Platynaias Pyganodon Utterbackia

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 74 0 0 --
Cladomics cladogram Lanceolaria Lanceolaria gladiola Cristaria Pletholophus Sinanodonta Pseudanodonta Brachyanodon Anodonta Pyganodon Simpsonaias Strophitus Alasmidonta Lasmigona

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 74 0 0 --
Cladomics cladogram Lanceolaria Lanceolaria gladiola Cristaria Pletholophus Sinanodonta Pseudanodonta Anodonta anatina Brachyanodon Anodonta cygnea Alasmidonta Lasmigona Pyganodon Strophitus Simpsonaias

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 35 0 0 --
Cladomics cladogram Lanceolaria lanceolata Lanceolaria gladiola Acuticosta Anodonta Pyganodon Cristaria Sinanodonta Anemina

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 37 0 0 --
Cladomics cladogram Acuticosta Lanceolaria lanceolata Lanceolaria gladiola

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.000 2 27 0 2 39 0 0 33.0
Cladomics cladogram Pletholophus Sinanodonta Beringiana Lanceolaria gladiola Acuticosta Lepidodesma Lanceolaria oxyrhyncha Cristaria Alasmidonta Lasmigona Anodonta Pseudanodonta Scriptolamprotula Middendorffinaia Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia Unio

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 39 0 0 --
Cladomics cladogram Lanceolaria gladiola Lanceolaria oxyrhyncha Alasmidonta Lasmigona Anodonta Pseudanodonta Cristaria Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 42 0 0 --
Cladomics cladogram Lanceolaria gladiola Lanceolaria oxyrhyncha Lepidodesma Acuticosta Globunio Scriptolamprotula Middendorffinaia Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Unio Tchangsinaia Inversiunio

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.000 2 26 0 2 42 0 0 32.0
Cladomics cladogram Alasmidonta Lasmigona Anodonta Pseudanodonta Lanceolaria gladiola Lanceolaria oxyrhyncha Cristaria Acuticosta Lepidodesma Globunio Scriptolamprotula Middendorffinaia Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia Unio

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 39 0 0 --
Cladomics cladogram Lanceolaria gladiola Lanceolaria oxyrhyncha Alasmidonta Lasmigona Anodonta Pseudanodonta Cristaria Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 39 0 0 --
Cladomics cladogram Cristaria Lanceolaria gladiola Lanceolaria oxyrhyncha

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 138 0 0 --
Cladomics cladogram Lanceolaria Lanceolaria gladiola Acuticosta Cristaria Anemina Sinanodonta Anodonta cygnea Anodonta exulcerata Anodonta anatina Brachyanodon Pseudanodonta Pyganodon Utterbackia Pterosyna Prolasmidonta Alasmidonta Platynaias

Wu et al., 2026 [Publ 4811], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 15 0 0 --
Cladomics cladogram Lanceolaria gladiola Lanceolaria lanceolata Acuticosta Brachyanodon Globunio Pseudoacuticosta Huizhou

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Lanceolaria grayii Anodonta Unio Pyganodon Pseudanodonta Alathyria Pronodularia Sinohyriopsis Quadrula Parvaspina Lampsilis Epioblasma Venustaconcha

Zhou et al., 2017 [Publ 3788], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 0 0 --
Cladomics cladogram Lanceolaria lanceolata Scriptolamprotula Nodularia Tchangsinaia

Sano et al., 2017 [Publ 3799], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 0 0 --
Cladomics cladogram Lanceolaria grayii Nodularia Inversiunio

Liu et al., 2017 [Publ 3939], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 0 0 --
Cladomics cladogram Lanceolaria grayii Unio Schistodesmus Cuneopsis Nodularia

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Lanceolaria lanceolata Scriptolamprotula Unio

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Lanceolaria lanceolata Scriptolamprotula Unio Sinanodonta Anodonta Utterbackia Pyganodon

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 0 0 --
Cladomics cladogram Lanceolaria grayii Cristaria Anemina Sinanodonta Anodonta Platynaias Pyganodon Utterbackia

Huang et al., 2019 [Publ 3843], fig. 3-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 0 0 --
Cladomics cladogram Lanceolaria lanceolata Schistodesmus Scriptolamprotula Unio

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram Lanceolaria lanceolata Unio Nodularia Tchangsinaia Cristaria Sinanodonta Anodonta Pyganodon

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 0 0 --
Cladomics cladogram Lanceolaria lanceolata Unio Tchangsinaia Nodularia

Guerra et al., 2019 [Publ 4168], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Lanceolaria lanceolata Scriptolamprotula Schistodesmus Unio

Guerra et al., 2019 [Publ 4168], fig. 1-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Lanceolaria lanceolata Schistodesmus Scriptolamprotula Unio

Guerra et al., 2019 [Publ 4168], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Lanceolaria lanceolata Sinanodonta Anodonta Platynaias Pyganodon Utterbackia

Guerra et al., 2019 [Publ 4168], fig. 2-M

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Lanceolaria lanceolata Scriptolamprotula Schistodesmus Unio

Kano et al., 2019 [Publ 4194], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 9 0 0 --
Cladomics cladogram Lanceolaria grayii Inversiunio Nodularia

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 0 0 --
Cladomics cladogram Lanceolaria grayii Inversiunio

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 0 0 --
Cladomics cladogram Lanceolaria grayii Chamberlainia Lamprotula

Wu et al., 2022 [Publ 4238], fig. 2e

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Lanceolaria lanceolata Scriptolamprotula Unio Pseudocuneopsis Nodularia Tchangsinaia Cuneopsis Schistodesmus

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 0 0 --
Cladomics cladogram Acuticosta Lanceolaria grayii

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 0 0 --
Cladomics cladogram Lanceolaria gladiola Acuticosta Cristaria Anemina Sinanodonta Anodonta Platynaias Pyganodon Utterbackia

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Lanceolaria lanceolata Anodonta

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Lanceolaria lanceolata Anodonta

Dai et al., 2024 [Publ 4386], fig. 5-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Lanceolaria grayii Scriptolamprotula Lepidodesma Unio Middendorffinaia Cuneopsis Nodularia Inversiunio Schistodesmus

Dai et al., 2024 [Publ 4386], fig. 5-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 21 0 0 --
Cladomics cladogram Lanceolaria grayii Scriptolamprotula Unio Middendorffinaia Nodularia Cuneopsis Inversiunio Schistodesmus

Dai et al., 2024 [Publ 4386], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 0 0 --
Cladomics cladogram Lanceolaria lanceolata Acuticosta Sinanodonta Pyganodon Utterbackia

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 0 0 --
Cladomics cladogram Lanceolaria lanceolata Anodonta Alasmidonta Utterbackia Cristaria Anemina Beringiana Sinanodonta

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Lanceolaria lanceolata Sinanodonta Anodonta Utterbackia Pyganodon Scriptolamprotula Unio Pseudocuneopsis Schistodesmus Nodularia Tchangsinaia Cuneopsis

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Scriptolamprotula Lanceolaria lanceolata

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 0 0 --
Cladomics cladogram Lanceolaria lanceolata Scriptolamprotula Unio Pseudocuneopsis Nodularia Tchangsinaia Cuneopsis Schistodesmus