Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Lanceolariini

genus Lanceolaria Conrad, 1853: cladomics

14 Recent valid species (16 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 14 (16 total)
analyzed in 109 cladograms in 65 publications
maximum number of terminals in an analysis = 10 (mean = 2.55, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 80
maximum number of included terminals = 10, (mean = 3.11 median = 3)
proportion of cladograms that support basic monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lanceolaria cladogram distances
PCoA Ordination

NJ Tree of Lanceolaria cladogram distances
NJ Tree

Cladograms

Wu et al., 2026 [Publ 4825], fig. 3

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
10 10 8 10 2 0 0 0 
  +----------------- Lanceolaria_oxyrhyncha **   
  |                                           
  |              +-- Lanceolaria_gladiola **     
--+  +-----------+                            
  |  |           +-- Lanceolaria_bilirata **     
  |  |                                        
  +--+        +----- Lanceolaria_acrorrhyncha ** 
     |  +-----+                               
     |  |     |  +-- Lanceolaria_grayii **       
     |  |     +--+                            
     +--+        +-- Lanceolaria_danzhouensis ** 
        |                                     
        |  +-------- Lanceolaria_kihirai **      
        +--+                                  
           |  +----- Lanceolaria_triformis **    
           +--+                               
              |  +-- Lanceolaria_lanceolata **   
              +--+                            
                 +-- Lanceolaria_helianthoides **

Wu et al., 2026 [Publ 4825], fig. 5

iTtTbiriSoTroTroSiD
10 10 8 10 9 0 0 2 
  +-------------------- Lanceolaria_oxyrhyncha **   
  |                                              
  |                 +-- Lanceolaria_bilirata **     
--+  +--------------+                            
  |  |              +-- Lanceolaria_gladiola **     
  |  |                                           
  +--+              +-- Lanceolaria_grayii **       
     |  +-----------+                            
     |  |           +-- Lanceolaria_danzhouensis ** 
     +--+                                        
        |  +----------- Lanceolaria_acrorrhyncha ** 
        +--+                                     
           |  +-------- Lanceolaria_kihirai **      
           +--+                                  
              |  +----- Lanceolaria_triformis **    
              +--+                               
                 |  +-- Lanceolaria_helianthoides **
                 +--+                            
                    +-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
8 8 6 8 114 0 0 4 
  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  +--+           +-- Lanceolaria_danzhouensis **
     |  +--------+                           
     |  |        +-- Lanceolaria_grayii **      
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_lanceolata **  
              +--+                           
                 +-- Lanceolaria_triformis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iTtTbiriSoTroTroSiD
8 8 6 8 4 0 0 4 
  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  +--+           +-- Lanceolaria_grayii **      
     |  +--------+                           
     |  |        +-- Lanceolaria_danzhouensis **
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_lanceolata **  
              +--+                           
                 +-- Lanceolaria_triformis **

Wu et al., 2026 [Publ 4825], fig. 6B

iTtTbiriSoTroTroSiD
7 7 5 7 58 0 0 3 
           +-- Lanceolaria_bilirata **     
  +--------+                            
  |        +-- Lanceolaria_gladiola **     
  |                                     
--+        +-- Lanceolaria_grayii **       
  |  +-----+                            
  |  |     +-- Lanceolaria_danzhouensis ** 
  +--+                                  
     |  +----- Lanceolaria_triformis **    
     +--+                               
        |  +-- Lanceolaria_helianthoides **
        +--+                            
           +-- Lanceolaria_lanceolata **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
7 7 5 7 623 0 0 5 
  +----------------- Lanceolaria_oxyrhyncha **  
--+                                          
  |  +-------------- Lanceolaria_gladiola **    
  +--+                                       
     |  +----------- Lanceolaria_grayii **      
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_triformis **   
              +--+                           
                 +-- Lanceolaria_lanceolata **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 3 5 196 0 0 5 
  +-------- Lanceolaria_gladiola **    
  |                                 
--+     +-- Lanceolaria_lanceolata **  
  |  +--+                           
  |  |  +-- Lanceolaria_triformis **   
  +--+                              
     |  +-- Lanceolaria_grayii **      
     +--+                           
        +-- Lanceolaria_danzhouensis **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 5 194 0 0 5 
  +-------- Lanceolaria_gladiola **    
  |                                 
--+     +-- Lanceolaria_lanceolata **  
  |  +--+                           
  |  |  +-- Lanceolaria_triformis **   
  +--+                              
     |  +-- Lanceolaria_grayii **      
     +--+                           
        +-- Lanceolaria_danzhouensis **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
5 5 2 5 42 0 0 6 
  +-------- Lanceolaria_gladiola **    
  |                                 
--+  +----- Lanceolaria_danzhouensis **
  |  |                              
  +--+----- Lanceolaria_grayii **      
     |                              
     |  +-- Lanceolaria_lanceolata **  
     +--+                           
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 41 0 0 5 
  +-------- Lanceolaria_gladiola **    
  |                                 
--+     +-- Lanceolaria_lanceolata **  
  |  +--+                           
  |  |  +-- Lanceolaria_triformis **   
  +--+                              
     |  +-- Lanceolaria_grayii **      
     +--+                           
        +-- Lanceolaria_danzhouensis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 2 5 196 0 0 6 
  +-------- Lanceolaria_gladiola **    
  |                                 
--+  +----- Lanceolaria_grayii **      
  |  |                              
  +--+----- Lanceolaria_danzhouensis **
     |                              
     |  +-- Lanceolaria_lanceolata **  
     +--+                           
        +-- Lanceolaria_triformis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
4 4 2 4 77 0 0 8 
  +-------- Lanceolaria_gladiola **    
--+                                 
  |  +----- Lanceolaria_danzhouensis **
  +--+                              
     |  +-- Lanceolaria_lanceolata **  
     +--+                           
        +-- Lanceolaria_grayii **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 49 0 0 6 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 65 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
3 3 1 3 35 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
3 3 1 3 10 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 84 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
3 18 0 3 122 0 0 23 
  +-- Alasmidonta             
  |                           
  +-- Anodonta_anatina        
  |                           
  +-- Anodonta_cygnea         
  |                           
  +-- Brachyanodon            
  |                           
  +-- Lasmigona               
  |                           
  +-- Pseudanodonta           
  |                           
  +-- Pyganodon               
  |                           
  +-- Simpsonaias             
  |                           
  +-- Strophitus              
--+                           
  +-- Sinanodonta_lauta       
  |                           
  +-- Cristaria_plicata       
  |                           
  +-- Cristaria_radiata       
  |                           
  +-- Pletholophus            
  |                           
  +-- Sinanodonta_lucida      
  |                           
  +-- Sinanodonta_jourdyi     
  |                           
  +-- Lanceolaria_gladiola **    
  |                           
  +-- Lanceolaria_grayii **      
  |                           
  +-- Lanceolaria_danzhouensis **

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 15 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 82 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
3 3 1 3 136 0 0 7 
  +----- Lanceolaria_gladiola **    
--+                              
  |  +-- Lanceolaria_danzhouensis **
  +--+                           
     +-- Lanceolaria_grayii **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 51 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 20 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
3 3 1 3 70 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
3 3 1 3 92 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 66 0 0 7 
  +----- Lanceolaria_oxyrhyncha **
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
3 3 1 3 70 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
3 21 14 3 21 0 0 26 
  +-------------- Lanceolaria_grayii **    
  |                                     
  +-------------- Lanceolaria_gladiola **  
  |                                     
  +-------------- Lanceolaria_lanceolata **
  |                                     
--+     +-------- Lasmigona             
  |  +--+                               
  |  |  |  +----- Pseudanodonta         
  |  |  +--+                            
  |  |     |  +-- Pyganodon             
  |  |     +--+                         
  +--+        +-- Strophitus            
     |                                  
     |  +-------- Unio                  
     |  |                               
     |  +-------- Scriptolamprotula     
     |  |                               
     +--+  +----- Cuneopsis             
        |  |                            
        |  |  +-- Nodularia             
        |  +--+                         
        +--+  +-- Schistodesmus         
           |                            
           |  +-- Tchangsinaia          
           |  |                         
           +--+-- Inversiunio           
              |                         
              +-- Pseudocuneopsis

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 14 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 66 0 0 7 
  +----- Lanceolaria_oxyrhyncha **
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 92 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 70 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 3 50 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
3 22 5 4 31 0 0 37 
  +----- Lanceolaria_oxyrhyncha__1 **
  |                               
  +----- Lanceolaria_oxyrhyncha__2 **
  |                               
  +----- Cristaria_plicata__1     
  |                               
  +----- Cristaria_plicata__2     
  |                               
  +----- Sinanodonta_lauta        
  |                               
  +----- Beringiana_japonica      
  |                               
  +----- Beringiana_fukuharai__1  
  |                               
  +----- Beringiana_beringiana    
  |                               
  +----- Beringiana_fukuharai__2  
  |                               
  +----- Beringiana_fukuharai__3  
  |                               
  +----- Buldowskia_iwakawai__1   
  |                               
  +----- Buldowskia_iwakawai__2   
  |                               
  +----- Sinanodonta_tumens__1    
--+                               
  +----- Sinanodonta_tumens__2    
  |                               
  +----- Pletholophus             
  |                               
  +----- Anemina                  
  |                               
  +----- Lanceolaria_lanceolata **   
  |                               
  +----- Cristaria_plicata__3     
  |                               
  +----- Lanceolaria_gladiola **     
  |                               
  +----- Sinanodonta_angula       
  |                               
  +----- Lepidodesma              
  |                               
  |  +-- Scriptolamprotula        
  +--+                            
  |  +-- Acuticosta               
  |                               
  |  +-- Cuneopsis                
  +--+                            
  |  +-- Nodularia                
  |                               
  |  +-- Schistodesmus            
  +--+                            
     +-- Inversiunio

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 53 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
3 3 1 3 92 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
3 3 1 3 136 0 0 7 
  +----- Lanceolaria_gladiola **    
--+                              
  |  +-- Lanceolaria_grayii **      
  +--+                           
     +-- Lanceolaria_danzhouensis **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
3 3 1 3 53 0 0 7 
  +----- Lanceolaria_gladiola ** 
--+                           
  |  +-- Lanceolaria_grayii **   
  +--+                        
     +-- Lanceolaria_triformis **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
3 21 19 3 21 0 0 25 
                          +-- Lanceolaria_lanceolata **
  +-----------------------+                         
  |                       +-- Lanceolaria_grayii **    
  |                                                 
--+  +----------------------- Lanceolaria_gladiola **  
  |  |                                              
  |  |              +-------- Pseudanodonta         
  +--+  +-----------+                               
     |  |           |  +----- Lasmigona             
     |  |           +--+                            
     |  |              |  +-- Strophitus            
     +--+              +--+                         
        |                 +-- Pyganodon             
        |                                           
        |  +----------------- Unio                  
        |  |                                        
        +--+  +-------------- Scriptolamprotula     
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              +--+        +-- Nodularia             
                 |  +-----+                         
                 |  |     +-- Schistodesmus         
                 +--+                               
                    |  +----- Tchangsinaia          
                    +--+                            
                       |  +-- Inversiunio           
                       +--+                         
                          +-- Pseudocuneopsis

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 72 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 66 0 0 7 
  +----- Lanceolaria_oxyrhyncha **
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 7 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 21 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 30 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 26 0 2 42 0 0 32 
  +-------------------------- Lanceolaria_gladiola **  
  |                                                 
  +-------------------------- Lanceolaria_oxyrhyncha **
  |                                                 
--+-------------------------- Cristaria             
  |                                                 
  |  +----------------------- Acuticosta            
  |  |                                              
  |  +----------------------- Lepidodesma           
  +--+                                              
     |  +-------------------- Globunio              
     |  |                                           
     +--+  +----------------- Scriptolamprotula     
        |  |                                        
        |  |              +-- Middendorffinaia      
        +--+  +-----------+                         
           |  |           +-- Nodularia             
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              |  |        +-- Diaurora              
              +--+--------+                         
                 |        +-- Schistodesmus         
                 |                                  
                 |  +-------- Pseudocuneopsis       
                 +--+                               
                    |  +----- Inversiunio           
                    +--+                            
                       |  +-- Tchangsinaia          
                       +--+                         
                          +-- Unio

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 34 0 0 -- 
  +-- Lanceolaria_grayii **      
--+                           
  +-- Lanceolaria_danzhouensis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 2 0 2 74 0 0 -- 
  +-- Lanceolaria **         
--+                       
  +-- Lanceolaria_gladiola **

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **   
--+                        
  +-- Lanceolaria_triformis **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
2 4 0 2 11 0 0 10 
  +----- Lanceolaria_grayii **  
  |                          
--+----- Acuticosta          
  |                          
  |  +-- Lanceolaria_gladiola **
  +--+                       
     +-- Scriptolamprotula

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 17 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 46 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 38 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 42 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 2 74 0 0 -- 
  +-- Lanceolaria **         
--+                       
  +-- Lanceolaria_gladiola **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
2 2 0 2 138 0 0 -- 
  +-- Lanceolaria **         
--+                       
  +-- Lanceolaria_gladiola **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
2 27 0 2 39 0 0 33 
  +-------------------- Lanceolaria_gladiola **  
  |                                           
  +-------------------- Acuticosta            
  |                                           
  +-------------------- Lepidodesma           
  |                                           
  +-------------------- Lanceolaria_oxyrhyncha **
  |                                           
  +-------------------- Cristaria             
  |                                           
--+                 +-- Alasmidonta           
  |              +--+                         
  |              |  +-- Lasmigona             
  +--------------+                            
  |              |  +-- Anodonta              
  |              +--+                         
  |                 +-- Pseudanodonta         
  |                                           
  |  +----------------- Scriptolamprotula     
  |  |                                        
  |  |              +-- Middendorffinaia      
  +--+  +-----------+                         
     |  |           +-- Nodularia             
     |  |                                     
     +--+  +----------- Cuneopsis             
        |  |                                  
        |  |        +-- Diaurora              
        +--+--------+                         
           |        +-- Schistodesmus         
           |                                  
           |  +-------- Pseudocuneopsis       
           +--+                               
              |  +----- Inversiunio           
              +--+                            
                 |  +-- Tchangsinaia          
                 +--+                         
                    +-- Unio

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **   
--+                        
  +-- Lanceolaria_triformis **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 44 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 42 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 80 0 0 -- 
  +-- Lanceolaria_gladiola **
--+                       
  +-- Lanceolaria_grayii **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 19 0 0 -- 
  +-- Lanceolaria_triformis ** 
--+                         
  +-- Lanceolaria_lanceolata **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 35 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_gladiola **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
2 2 0 2 37 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_gladiola **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
2 2 0 2 46 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 15 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_lanceolata **

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 12 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 13 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 3 31 0 0 -- 
  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 42 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 0 0 -- 
-- Lanceolaria_grayii **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Lanceolaria_grayii **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Lanceolaria_lanceolata **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Lanceolaria_grayii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Lanceolaria_gladiola **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_grayii **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Lanceolaria_lanceolata **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Lanceolaria_lanceolata **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Lanceolaria_grayii **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_grayii **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Lanceolaria_lanceolata **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Lanceolaria_lanceolata **

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Lanceolaria_grayii **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Lanceolaria_grayii **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Lanceolaria_lanceolata **

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Lanceolaria_lanceolata **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Lanceolaria_grayii **

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Lanceolaria_grayii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Lanceolaria_lanceolata **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Lanceolaria_lanceolata **