Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Lanceolariini

genus Lanceolaria Conrad, 1853: cladomics

15 Recent valid species (17 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 15 (17 total)
analyzed in 106 cladograms in 64 publications
maximum number of terminals in an analysis = 8 (mean = 2.27, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 77
maximum number of included terminals = 8, (mean = 2.75 median = 2)
proportion of cladograms that support basic monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lanceolaria cladogram distances
PCoA Ordination

NJ Tree of Lanceolaria cladogram distances
NJ Tree

Cladograms

Huang et al., 2019 [Publ 3843], fig. S1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
5 5 4 12 42 0 0 0 
  +----------- Lanceolaria_gladiola **       
  |                                       
  |  +-------- Lanceolaria_grayii__5 **      
--+  |                                    
  |  |     +-- Lanceolaria_lanceolata **     
  |  +-----+                              
  |  |     +-- Lanceolaria_triformis **      
  +--+                                    
     |  +----- Lanceolaria_grayii__4 **      
     |  |                                 
     |  |  +-- Lanceolaria_eucylindrica__1 **
     |  |  |                              
     +--+  +-- Lanceolaria_eucylindrica__2 **
        |  |                              
        |  +-- Lanceolaria_eucylindrica__3 **
        |  |                              
        +--+-- Lanceolaria_eucylindrica__4 **
           |                              
           +-- Lanceolaria_grayii__1 **      
           |                              
           +-- Lanceolaria_grayii__2 **      
           |                              
           +-- Lanceolaria_grayii__3 **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
8 8 6 8 114 0 0 9 
  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  +--+           +-- Lanceolaria_bilirata **    
     |  +--------+                           
     |  |        +-- Lanceolaria_grayii **      
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_lanceolata **  
              +--+                           
                 +-- Lanceolaria_triformis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
8 8 7 9 624 0 0 8 
  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  |  |        +----- Lanceolaria_grayii__2 **   
  +--+  +-----+                              
     |  |     |  +-- Lanceolaria_eucylindrica **
     |  |     +--+                           
     +--+        +-- Lanceolaria_grayii__1 **   
        |                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_triformis **   
              +--+                           
                 +-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iTtTbiriSoTroTroSiD
8 8 6 8 4 0 0 9 
  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  +--+           +-- Lanceolaria_grayii **      
     |  +--------+                           
     |  |        +-- Lanceolaria_bilirata **    
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_lanceolata **  
              +--+                           
                 +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 4 41 0 0 8 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 5 196 0 0 8 
  +-------- Lanceolaria_gladiola **  
  |                               
--+  +----- Lanceolaria_grayii__1 ** 
  |  |                            
  +--+----- Lanceolaria_grayii__2 ** 
     |                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
4 4 2 4 194 0 0 8 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
4 4 2 4 196 0 0 8 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 49 0 0 8 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 71 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 53 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
3 3 2 4 77 0 0 9 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii__2 ** 
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_grayii__1 **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 73 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 20 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 66 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
3 3 1 3 89 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
3 3 1 3 93 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
3 3 1 3 10 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
3 3 1 3 53 0 0 9 
  +----- Lanceolaria_gladiola ** 
--+                           
  |  +-- Lanceolaria_grayii **   
  +--+                        
     +-- Lanceolaria_triformis **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 51 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 93 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
3 3 1 3 71 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
3 3 1 3 89 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 85 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 15 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
3 3 1 3 36 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 14 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 83 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
3 21 14 3 21 0 0 22 
  +-------------- Lanceolaria_grayii **    
  |                                     
  +-------------- Lanceolaria_gladiola **  
  |                                     
  +-------------- Lanceolaria_lanceolata **
  |                                     
--+     +-------- Lasmigona             
  |  +--+                               
  |  |  |  +----- Pseudanodonta         
  |  |  +--+                            
  |  |     |  +-- Pyganodon             
  |  |     +--+                         
  +--+        +-- Strophitus            
     |                                  
     |  +-------- Unio                  
     |  |                               
     |  +-------- Scriptolamprotula     
     |  |                               
     +--+  +----- Cuneopsis             
        |  |                            
        |  |  +-- Nodularia             
        |  +--+                         
        +--+  +-- Schistodesmus         
           |                            
           |  +-- Tchangsinaia          
           |  |                         
           +--+-- Inversiunio           
              |                         
              +-- Pseudocuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
3 3 1 3 93 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
3 22 5 4 31 0 0 30 
  +----- Lanceolaria_grayii__1 **  
  |                             
  +----- Lanceolaria_grayii__2 **  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata ** 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola **   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Scriptolamprotula      
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
3 3 1 3 71 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 3 51 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
3 21 19 3 21 0 0 21 
                          +-- Lanceolaria_lanceolata **
  +-----------------------+                         
  |                       +-- Lanceolaria_grayii **    
  |                                                 
--+  +----------------------- Lanceolaria_gladiola **  
  |  |                                              
  |  |              +-------- Pseudanodonta         
  +--+  +-----------+                               
     |  |           |  +----- Lasmigona             
     |  |           +--+                            
     |  |              |  +-- Strophitus            
     +--+              +--+                         
        |                 +-- Pyganodon             
        |                                           
        |  +----------------- Unio                  
        |  |                                        
        +--+  +-------------- Scriptolamprotula     
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              +--+        +-- Nodularia             
                 |  +-----+                         
                 |  |     +-- Schistodesmus         
                 +--+                               
                    |  +----- Tchangsinaia          
                    +--+                            
                       |  +-- Inversiunio           
                       +--+                         
                          +-- Pseudocuneopsis

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 44 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 80 0 0 -- 
  +-- Lanceolaria_gladiola **
--+                       
  +-- Lanceolaria_grayii **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 30 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 3 66 0 0 -- 
  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 12 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
2 2 0 2 37 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_gladiola **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 21 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
2 2 0 2 139 0 0 -- 
  +-- Lanceolaria **         
--+                       
  +-- Lanceolaria_gladiola **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 2 136 0 0 -- 
  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 3 31 0 0 -- 
  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 34 0 0 -- 
  +-- Lanceolaria_grayii **
--+                     
  +-- Lanceolaria **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 13 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 15 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_lanceolata **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 43 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 27 0 2 43 0 0 30 
  +-------------------------- Lanceolaria_gladiola **  
  |                                                 
  +-------------------------- Lanceolaria_oxyrhyncha **
  |                                                 
--+-------------------------- Cristaria             
  |                                                 
  |  +----------------------- Acuticosta            
  |  |                                              
  |  +----------------------- Lepidodesma           
  +--+                                              
     |  +-------------------- Globunio              
     |  |                                           
     +--+  +----------------- Scriptolamprotula     
        |  |                                        
        |  |              +-- Middendorffinaia      
        +--+  +-----------+                         
           |  |           +-- Nodularia             
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              |  |        +-- Diaurora              
              +--+--------+                         
                 |        +-- Schistodesmus         
                 |                                  
                 |  +-------- Pseudocuneopsis       
                 +--+                               
                    |  +----- Inversiunio           
                    +--+                            
                       |  +-- Tchangsinaia          
                       +--+                         
                          +-- Unio

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 136 0 0 -- 
  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 38 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 17 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
2 27 0 2 39 0 0 30 
  +-------------------- Lanceolaria_gladiola **  
  |                                           
  +-------------------- Acuticosta            
  |                                           
  +-------------------- Lepidodesma           
  |                                           
  +-------------------- Lanceolaria_oxyrhyncha **
  |                                           
  +-------------------- Cristaria             
  |                                           
--+                 +-- Alasmidonta           
  |              +--+                         
  |              |  +-- Lasmigona             
  +--------------+                            
  |              |  +-- Anodonta              
  |              +--+                         
  |                 +-- Pseudanodonta         
  |                                           
  |  +----------------- Scriptolamprotula     
  |  |                                        
  |  |              +-- Middendorffinaia      
  +--+  +-----------+                         
     |  |           +-- Nodularia             
     |  |                                     
     +--+  +----------- Cuneopsis             
        |  |                                  
        |  |        +-- Diaurora              
        +--+--------+                         
           |        +-- Schistodesmus         
           |                                  
           |  +-------- Pseudocuneopsis       
           +--+                               
              |  +----- Inversiunio           
              +--+                            
                 |  +-- Tchangsinaia          
                 +--+                         
                    +-- Unio

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 17 0 3 122 0 0 19 
  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina     
  |                        
  +-- Anodonta_cygnea      
  |                        
  +-- Brachyanodon         
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta    
  |                        
  +-- Cristaria_plicata    
  |                        
  +-- Cristaria_radiata    
  |                        
  +-- Pletholophus         
  |                        
  +-- Sinanodonta_lucida   
  |                        
  +-- Sinanodonta_jourdyi  
  |                        
  +-- Lanceolaria_gladiola ** 
  |                        
  +-- Lanceolaria_grayii__1 **
  |                        
  +-- Lanceolaria_grayii__2 **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 35 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_gladiola **

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **   
--+                        
  +-- Lanceolaria_triformis **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
2 4 0 2 11 0 0 11 
  +----- Lanceolaria_grayii **  
  |                          
--+----- Acuticosta          
  |                          
  |  +-- Lanceolaria_gladiola **
  +--+                       
     +-- Scriptolamprotula

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 42 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **   
--+                        
  +-- Lanceolaria_triformis **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 47 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 42 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 3 66 0 0 -- 
  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 2 74 0 0 -- 
  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
2 2 0 2 46 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 19 0 0 -- 
  +-- Lanceolaria_triformis ** 
--+                         
  +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 3 66 0 0 -- 
  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 2 0 2 74 0 0 -- 
  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_grayii **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Lanceolaria_grayii **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Lanceolaria_grayii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Lanceolaria_lanceolata **

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Lanceolaria_lanceolata **

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Lanceolaria_lanceolata **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Lanceolaria_grayii **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Lanceolaria_grayii **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Lanceolaria_gladiola **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Lanceolaria_grayii **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Lanceolaria_lanceolata **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 0 0 -- 
-- Lanceolaria_grayii **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Lanceolaria_lanceolata **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Lanceolaria_lanceolata **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_grayii **

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Lanceolaria_grayii **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Lanceolaria_lanceolata **

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Lanceolaria_grayii **