Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Lanceolariini

genus Lanceolaria Conrad, 1853: cladomics

15 Recent valid species (17 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 15 (17 total)
analyzed in 99 cladograms in 61 publications
maximum number of terminals in an analysis = 8 (mean = 2.25, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 73
maximum number of included terminals = 8, (mean = 2.7 median = 2)
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lanceolaria cladogram distances
PCoA Ordination

NJ Tree of Lanceolaria cladogram distances
NJ Tree

Cladograms

Huang et al., 2019 [Publ 3843], fig. S1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
5 5 4 12 42 0 0 0 
  +----------- Lanceolaria_gladiola **       
  |                                       
  |  +-------- Lanceolaria_grayii__5 **      
--+  |                                    
  |  |     +-- Lanceolaria_lanceolata **     
  |  +-----+                              
  |  |     +-- Lanceolaria_triformis **      
  +--+                                    
     |  +----- Lanceolaria_grayii__4 **      
     |  |                                 
     |  |  +-- Lanceolaria_eucylindrica__1 **
     |  |  |                              
     +--+  +-- Lanceolaria_eucylindrica__2 **
        |  |                              
        |  +-- Lanceolaria_eucylindrica__3 **
        |  |                              
        +--+-- Lanceolaria_eucylindrica__4 **
           |                              
           +-- Lanceolaria_grayii__1 **      
           |                              
           +-- Lanceolaria_grayii__2 **      
           |                              
           +-- Lanceolaria_grayii__3 **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
8 8 6 8 114 0 0 9 
  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  +--+           +-- Lanceolaria_bilirata **    
     |  +--------+                           
     |  |        +-- Lanceolaria_grayii **      
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_lanceolata **  
              +--+                           
                 +-- Lanceolaria_triformis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iTtTbiriSoTroTroSiD
8 8 6 8 4 0 0 9 
  +----------------- Lanceolaria_oxyrhyncha **  
  |                                          
--+  +-------------- Lanceolaria_gladiola **    
  |  |                                       
  +--+           +-- Lanceolaria_grayii **      
     |  +--------+                           
     |  |        +-- Lanceolaria_bilirata **    
     +--+                                    
        |  +-------- Lanceolaria_acrorrhyncha **
        +--+                                 
           |  +----- Lanceolaria_kihirai **     
           +--+                              
              |  +-- Lanceolaria_lanceolata **  
              +--+                           
                 +-- Lanceolaria_triformis **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 49 0 0 8 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 5 197 0 0 8 
  +-------- Lanceolaria_gladiola **  
  |                               
--+  +----- Lanceolaria_grayii__1 ** 
  |  |                            
  +--+----- Lanceolaria_grayii__2 ** 
     |                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
4 4 2 4 195 0 0 8 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
4 4 2 4 197 0 0 8 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
4 4 2 4 41 0 0 8 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii **    
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_triformis **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
3 3 1 3 71 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
3 22 5 4 31 0 0 30 
  +----- Lanceolaria_grayii__1 **  
  |                             
  +----- Lanceolaria_grayii__2 **  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata ** 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola **   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 3 51 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
3 3 2 4 77 0 0 9 
  +-------- Lanceolaria_gladiola **  
--+                               
  |  +----- Lanceolaria_grayii__2 ** 
  +--+                            
     |  +-- Lanceolaria_lanceolata **
     +--+                         
        +-- Lanceolaria_grayii__1 **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 71 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 20 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
3 3 1 3 53 0 0 9 
  +----- Lanceolaria_gladiola ** 
--+                           
  |  +-- Lanceolaria_grayii **   
  +--+                        
     +-- Lanceolaria_triformis **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 66 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
3 3 1 3 89 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
3 3 1 3 36 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
3 3 1 3 93 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
3 3 1 3 93 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 53 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
3 3 1 3 89 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 3 88 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
3 21 14 3 21 0 0 22 
  +-------------- Lanceolaria_grayii **    
  |                                     
  +-------------- Lanceolaria_gladiola **  
  |                                     
  +-------------- Lanceolaria_lanceolata **
  |                                     
--+     +-------- Lasmigona             
  |  +--+                               
  |  |  |  +----- Pseudanodonta         
  |  |  +--+                            
  |  |     |  +-- Pyganodon             
  |  |     +--+                         
  +--+        +-- Strophitus            
     |                                  
     |  +-------- Unio                  
     |  |                               
     |  +-------- Aculamprotula         
     |  |                               
     +--+  +----- Cuneopsis             
        |  |                            
        |  |  +-- Nodularia             
        |  +--+                         
        +--+  +-- Schistodesmus         
           |                            
           |  +-- Tchangsinaia          
           |  |                         
           +--+-- Inversiunio           
              |                         
              +-- Pseudocuneopsis

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 51 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
3 3 1 3 10 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
3 21 19 3 21 0 0 21 
                          +-- Lanceolaria_lanceolata **
  +-----------------------+                         
  |                       +-- Lanceolaria_grayii **    
  |                                                 
--+  +----------------------- Lanceolaria_gladiola **  
  |  |                                              
  |  |              +-------- Pseudanodonta         
  +--+  +-----------+                               
     |  |           |  +----- Lasmigona             
     |  |           +--+                            
     |  |              |  +-- Strophitus            
     +--+              +--+                         
        |                 +-- Pyganodon             
        |                                           
        |  +----------------- Unio                  
        |  |                                        
        +--+  +-------------- Aculamprotula         
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              +--+        +-- Nodularia             
                 |  +-----+                         
                 |  |     +-- Schistodesmus         
                 +--+                               
                    |  +----- Tchangsinaia          
                    +--+                            
                       |  +-- Inversiunio           
                       +--+                         
                          +-- Pseudocuneopsis

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 15 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 85 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
3 3 1 3 71 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 14 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
3 3 1 3 83 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 73 0 0 9 
  +----- Lanceolaria_gladiola **  
--+                            
  |  +-- Lanceolaria_grayii **    
  +--+                         
     +-- Lanceolaria_lanceolata **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 19 0 0 -- 
  +-- Lanceolaria_triformis ** 
--+                         
  +-- Lanceolaria_lanceolata **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 47 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 2 0 2 74 0 0 -- 
  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 42 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
2 27 0 2 43 0 0 30 
  +-------------------------- Lanceolaria_gladiola **  
  |                                                 
  +-------------------------- Lanceolaria_oxyrhyncha **
  |                                                 
--+-------------------------- Cristaria             
  |                                                 
  |  +----------------------- Acuticosta            
  |  |                                              
  |  +----------------------- Lepidodesma           
  +--+                                              
     |  +-------------------- Globunio              
     |  |                                           
     +--+  +----------------- Aculamprotula         
        |  |                                        
        |  |              +-- Middendorffinaia      
        +--+  +-----------+                         
           |  |           +-- Nodularia             
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              |  |        +-- Diaurora              
              +--+--------+                         
                 |        +-- Schistodesmus         
                 |                                  
                 |  +-------- Pseudocuneopsis       
                 +--+                               
                    |  +----- Inversiunio           
                    +--+                            
                       |  +-- Tchangsinaia          
                       +--+                         
                          +-- Unio

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 42 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 3 66 0 0 -- 
  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 17 0 3 123 0 0 19 
  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina     
  |                        
  +-- Anodonta_cygnea      
  |                        
  +-- Anodonta_nuttalliana 
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta    
  |                        
  +-- Cristaria_plicata    
  |                        
  +-- Cristaria_radiata    
  |                        
  +-- Pletholophus         
  |                        
  +-- Sinanodonta_lucida   
  |                        
  +-- Sinanodonta_jourdyi  
  |                        
  +-- Lanceolaria_gladiola ** 
  |                        
  +-- Lanceolaria_grayii__1 **
  |                        
  +-- Lanceolaria_grayii__2 **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 17 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 44 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **   
--+                        
  +-- Lanceolaria_triformis **

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 12 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 13 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 3 66 0 0 -- 
  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 21 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 35 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_gladiola **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
2 2 0 2 46 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 81 0 0 -- 
  +-- Lanceolaria_gladiola **
--+                       
  +-- Lanceolaria_grayii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 2 74 0 0 -- 
  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 34 0 0 -- 
  +-- Lanceolaria_grayii **
--+                     
  +-- Lanceolaria **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
2 2 0 2 37 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_gladiola **

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **   
--+                        
  +-- Lanceolaria_triformis **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
2 4 0 2 11 0 0 11 
  +----- Lanceolaria_grayii **  
  |                          
--+----- Acuticosta          
  |                          
  |  +-- Lanceolaria_gladiola **
  +--+                       
     +-- Aculamprotula

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 43 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 30 0 0 -- 
  +-- Lanceolaria_lanceolata **
--+                         
  +-- Lanceolaria_grayii **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 38 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 137 0 0 -- 
  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 2 137 0 0 -- 
  +-- Lanceolaria_grayii **  
--+                       
  +-- Lanceolaria_gladiola **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
2 27 0 2 39 0 0 30 
  +-------------------- Lanceolaria_gladiola **  
  |                                           
  +-------------------- Acuticosta            
  |                                           
  +-------------------- Lepidodesma           
  |                                           
  +-------------------- Lanceolaria_oxyrhyncha **
  |                                           
  +-------------------- Cristaria             
  |                                           
--+                 +-- Alasmidonta           
  |              +--+                         
  |              |  +-- Lasmigona             
  +--------------+                            
  |              |  +-- Anodonta              
  |              +--+                         
  |                 +-- Pseudanodonta         
  |                                           
  |  +----------------- Aculamprotula         
  |  |                                        
  |  |              +-- Middendorffinaia      
  +--+  +-----------+                         
     |  |           +-- Nodularia             
     |  |                                     
     +--+  +----------- Cuneopsis             
        |  |                                  
        |  |        +-- Diaurora              
        +--+--------+                         
           |        +-- Schistodesmus         
           |                                  
           |  +-------- Pseudocuneopsis       
           +--+                               
              |  +----- Inversiunio           
              +--+                            
                 |  +-- Tchangsinaia          
                 +--+                         
                    +-- Unio

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 32 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 3 31 0 0 -- 
  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 0 0 -- 
  +-- Lanceolaria_grayii **    
--+                         
  +-- Lanceolaria_lanceolata **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 39 0 0 -- 
  +-- Lanceolaria_gladiola **  
--+                         
  +-- Lanceolaria_oxyrhyncha **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 3 66 0 0 -- 
  +----- Lanceolaria_grayii__1 ** 
--+                            
  |  +-- Lanceolaria_lanceolata **
  +--+                         
     +-- Lanceolaria_grayii__2 **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Lanceolaria_grayii **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Lanceolaria_lanceolata **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Lanceolaria_lanceolata **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Lanceolaria_lanceolata **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Lanceolaria_gladiola **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Lanceolaria_grayii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Lanceolaria_lanceolata **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Lanceolaria_grayii **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 0 0 -- 
-- Lanceolaria_grayii **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Lanceolaria_grayii **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Lanceolaria_grayii **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Lanceolaria_lanceolata **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Lanceolaria_grayii **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Lanceolaria_lanceolata **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Lanceolaria_lanceolata **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Lanceolaria_lanceolata **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_grayii **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_grayii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Lanceolaria_lanceolata **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Lanceolaria_lanceolata **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Lanceolaria_lanceolata **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Lanceolaria_grayii **