Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina

genus Quadrula Rafinesque, 1820: cladomics

5 Recent valid species (9 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 5 (9 total)
analyzed in 191 cladograms in 115 publications
maximum number of terminals in an analysis = 4 (mean = 1.26, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 35
maximum number of included terminals = 4, (mean = 2.4 median = 2)
proportion of cladograms that support basic monophyly = 0.6

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Quadrula cladogram distances
PCoA Ordination

NJ Tree of Quadrula cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
4 5 3 4 628 12 3 0 
                 +-- Theliderma_johnsoni  
        +--------+                        
        |        +-- Theliderma_metanevra 
        |                                 
  +-----+        +-- Quadrula_nobilis **     
  |     |  +-----+                        
  |     |  |     +-- Quadrula_verrucosa **   
  |     +--+                              
  |        |  +----- Theliderma_cylindrica
  |        +--+                           
  |           |  +-- Quadrula_fragosa **     
--+           +--+                        
  |              +-- Quadrula_quadrula **    
  |                                       
  |           +----- Pustulosa_kieneriana 
  |  +--------+                           
  |  |        |  +-- Pustulosa_infucata   
  |  |        +--+                        
  +--+           +-- Pustulosa_kleiniana  
     |                                    
     |  +----------- Cyclonaias           
     |  |                                 
     +--+        +-- Pustulosa_pustulosa  
        |  +-----+                        
        |  |     +-- Pustulosa_succissa   
        +--+                              
           |  +----- Pustulosa_nodulata   
           +--+                           
              |  +-- Pustulosa_necki      
              +--+                        
                 +-- Pustulosa_petrina

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
3 3 1 3 26 14 3 2 
  +----------------------- Theliderma          
  |                                            
  |                 +----- Quadrula_quadrula **   
--+  +--------------+                          
  |  |              |  +-- Quadrula_nobilis **    
  |  |              +--+                       
  |  |                 +-- Quadrula_verrucosa **  
  +--+                                         
     |  +----------------- Pustulosa_kieneriana
     |  |                                      
     |  |              +-- Pustulosa_infucata  
     +--+  +-----------+                       
        |  |           +-- Pustulosa_kleiniana 
        |  |                                   
        +--+           +-- Pustulosa_pustulosa 
           |  +--------+                       
           |  |        +-- Pustulosa_succissa  
           +--+                                
              |  +-------- Cyclonaias          
              +--+                             
                 |  +----- Pustulosa_necki     
                 +--+                          
                    |  +-- Pustulosa_petrina   
                    +--+                       
                       +-- Pustulosa_nodulata

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iTtTbiriSoTroTroSiD
4 4 2 4 23 13 3 3 
  +----------------- Theliderma        
--+                                    
  |  +-------------- Cyclonaias        
  +--+                                 
     |  +----------- Pustulosa         
     +--+                              
        |  +-------- Quadrula_nobilis **  
        +--+                           
           |  +----- Quadrula_verrucosa **
           +--+                        
              |  +-- Quadrula_fragosa **  
              +--+                     
                 +-- Quadrula_quadrula **

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iTtTbiriSoTroTroSiD
3 8 6 3 70 6 3 9 
  +----------------- Theliderma         
  |                                     
--+  +-------------- Quadrula_verrucosa ** 
  |  |                                  
  |  |           +-- Quadrula_quadrula **  
  +--+  +--------+                      
     |  |        +-- Quadrula_nobilis **   
     |  |                               
     +--+        +-- Pustulosa_infucata 
        |  +-----+                      
        |  |     +-- Pustulosa_kleiniana
        +--+                            
           |  +----- Cyclonaias         
           +--+                         
              |  +-- Pustulosa_pustulosa
              +--+                      
                 +-- Pustulosa_succissa

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
3 8 5 3 99 6 3 7 
  +-------------- Theliderma          
  |                                   
--+  +----------- Quadrula_quadrula **   
  |  |                                
  |  |        +-- Quadrula_nobilis **    
  +--+  +-----+                       
     |  |     +-- Quadrula_verrucosa **  
     |  |                             
     +--+  +----- Pustulosa_kieneriana
        |  |                          
        |  |  +-- Cyclonaias          
        +--+--+                       
           |  +-- Pustulosa_succissa  
           |                          
           |  +-- Pustulosa_infucata  
           +--+                       
              +-- Pustulosa_kleiniana

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iTtTbiriSoTroTroSiD
3 9 6 3 70 6 3 7 
              +-- Quadrula_verrucosa ** 
  +-----------+                      
  |           +-- Quadrula_nobilis **   
  |                                  
--+  +----------- Quadrula_quadrula **  
  |  |                               
  |  +----------- Theliderma         
  +--+                               
     |        +-- Pustulosa_infucata 
     |  +-----+                      
     |  |     +-- Pustulosa_kleiniana
     +--+                            
        |  +----- Cyclonaias         
        +--+                         
           |  +-- Pustulosa_pustulosa
           +--+                      
              +-- Pustulosa_succissa

Szumowski et al., 2012 [Publ 3569], fig. 5

iTtTbiriSoTroTroSiD
3 3 2 4 11 10 2 5 
  +----------- Pustulosa           
--+                                
  |  +-------- Quadrula_quadrula__1 **
  +--+                             
     |  +----- Quadrula_verrucosa **  
     +--+                          
        |  +-- Quadrula_nobilis **    
        +--+                       
           +-- Quadrula_quadrula__2 **

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
3 14 12 3 25 11 3 14 
  +----------------------- Quadrula_quadrula **   
  |                                            
  |                    +-- Quadrula_verrucosa **  
--+  +-----------------+                       
  |  |                 +-- Quadrula_nobilis **    
  |  |                                         
  +--+  +----------------- Theliderma          
     |  |                                      
     |  |           +----- Pustulosa_kieneriana
     +--+  +--------+                          
        |  |        |  +-- Pustulosa_infucata  
        |  |        +--+                       
        +--+           +-- Pustulosa_kleiniana 
           |                                   
           |  +----------- Cyclonaias          
           |  |                                
           +--+        +-- Pustulosa_pustulosa 
              |  +-----+                       
              |  |     +-- Pustulosa_succissa  
              +--+                             
                 |  +----- Pustulosa_nodulata  
                 +--+                          
                    |  +-- Pustulosa_necki     
                    +--+                       
                       +-- Pustulosa_petrina

Serb et al., 2003 [Publ 2659], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 26 10 2 4 
  +----------- Theliderma        
--+                              
  |  +-------- Pustulosa         
  +--+                           
     |  +----- Quadrula_verrucosa **
     +--+                        
        |  +-- Quadrula_quadrula ** 
        +--+                     
           +-- Quadrula_nobilis **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
3 12 10 3 23 11 3 11 
  +----------------------- Theliderma          
  |                                            
--+  +-------------------- Quadrula_quadrula **   
  |  |                                         
  |  |                 +-- Quadrula_verrucosa **  
  +--+  +--------------+                       
     |  |              +-- Quadrula_nobilis **    
     |  |                                      
     |  |           +----- Pustulosa_kieneriana
     +--+  +--------+                          
        |  |        |  +-- Pustulosa_infucata  
        |  |        +--+                       
        |  |           +-- Pustulosa_kleiniana 
        +--+                                   
           |           +-- Pustulosa_pustulosa 
           |  +--------+                       
           |  |        +-- Pustulosa_succissa  
           +--+                                
              |  +-------- Cyclonaias          
              +--+                             
                 |  +----- Pustulosa_necki     
                 +--+                          
                    |  +-- Pustulosa_petrina   
                    +--+                       
                       +-- Pustulosa_nodulata

Serb et al., 2003 [Publ 2659], fig. 3

iTtTbiriSoTroTroSiD
3 13 11 3 26 10 2 10 
        +-- Quadrula_verrucosa **
  +-----+                     
  |     +-- Quadrula_nobilis **  
--+                           
  |  +----- Quadrula_quadrula ** 
  +--+                        
     |  +-- Theliderma        
     +--+                     
        +-- Pustulosa

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 3 99 6 3 4 
  +----------- Theliderma          
  |                                
  |     +----- Quadrula_verrucosa **  
--+  +--+                          
  |  |  |  +-- Quadrula_quadrula **   
  |  |  +--+                       
  |  |     +-- Quadrula_nobilis **    
  +--+                             
     |  +----- Pustulosa_kieneriana
     |  |                          
     |  |  +-- Cyclonaias          
     +--+--+                       
        |  +-- Pustulosa_succissa  
        |                          
        |  +-- Pustulosa_infucata  
        +--+                       
           +-- Pustulosa_kleiniana

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iTtTbiriSoTroTroSiD
2 3 0 2 29 1 1 4 
  +----- Quadrula_verrucosa **
--+                        
  |  +-- Cyclonaias        
  +--+                     
     +-- Quadrula_quadrula **

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
2 5 0 2 16 5 2 6 
  +-------------- Theliderma          
--+                                   
  |  +----------- Quadrula_quadrula **   
  +--+                                
     |  +-------- Pustulosa_infucata  
     +--+                             
        |  +----- Quadrula_verrucosa **  
        +--+                          
           |  +-- Pustulosa_kieneriana
           +--+                       
              +-- Pustulosa_succissa

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
2 5 0 2 103 3 3 7 
  +-- Cyclonaias            
  |                         
  +-- Pustulosa_pustulosa__1
  |                         
  +-- Theliderma            
--+                         
  +-- Pustulosa_pustulosa__2
  |                         
  +-- Quadrula_quadrula **     
  |                         
  +-- Quadrula_verrucosa **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 136 0 0 -- 
  +-- Quadrula_quadrula ** 
--+                     
  +-- Quadrula_verrucosa **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
2 2 0 2 23 0 0 -- 
  +-- Quadrula_quadrula ** 
--+                     
  +-- Quadrula_verrucosa **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
2 2 0 2 196 0 0 -- 
  +-- Quadrula_quadrula ** 
--+                     
  +-- Quadrula_verrucosa **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 44 0 0 -- 
  +-- Quadrula_verrucosa **
--+                     
  +-- Quadrula_quadrula **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 44 0 0 -- 
  +-- Quadrula_verrucosa **
--+                     
  +-- Quadrula_quadrula **

Boyer et al., 2011 [Publ 2718], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 36 4 3 -- 
  +----------- Theliderma         
  |                               
--+        +-- Quadrula_verrucosa ** 
  |  +-----+                      
  |  |     +-- Quadrula_quadrula **  
  +--+                            
     |  +----- Pustulosa_pustulosa
     +--+                         
        |  +-- Cyclonaias         
        +--+                      
           +-- Pustulosa_nodulata

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 2 136 0 0 -- 
  +-- Quadrula_quadrula ** 
--+                     
  +-- Quadrula_verrucosa **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
2 2 0 2 198 0 0 -- 
  +-- Quadrula_quadrula ** 
--+                     
  +-- Quadrula_verrucosa **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
2 2 0 2 78 0 0 -- 
  +-- Quadrula_quadrula ** 
--+                     
  +-- Quadrula_verrucosa **

Johnson et al., 2018 [Publ 3844], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 14 10 3 -- 
  +-------------------- Theliderma          
  |                                         
  |                 +-- Quadrula_quadrula **   
--+  +--------------+                       
  |  |              +-- Quadrula_verrucosa **  
  |  |                                      
  |  |           +----- Pustulosa_kieneriana
  +--+  +--------+                          
     |  |        |  +-- Pustulosa_infucata  
     |  |        +--+                       
     |  |           +-- Pustulosa_kleiniana 
     +--+                                   
        |           +-- Pustulosa_pustulosa 
        |  +--------+                       
        |  |        +-- Pustulosa_succissa  
        +--+                                
           |  +-------- Cyclonaias          
           +--+                             
              |  +----- Pustulosa_necki     
              +--+                          
                 |  +-- Pustulosa_nodulata  
                 +--+                       
                    +-- Pustulosa_petrina

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 45 0 0 -- 
  +-- Quadrula_verrucosa **
--+                     
  +-- Quadrula_quadrula **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 45 0 0 -- 
  +-- Quadrula_verrucosa **
--+                     
  +-- Quadrula_quadrula **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 9 0 2 122 0 0 10 
  +-- Amblema           
  |                     
  +-- Ortmanniana       
  |                     
  +-- Lampsilis         
  |                     
  +-- Cambarunio        
  |                     
--+-- Elliptio          
  |                     
  +-- Eurynia           
  |                     
  +-- Pleurobema        
  |                     
  +-- Quadrula_quadrula ** 
  |                     
  +-- Quadrula_verrucosa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
2 2 0 2 74 0 0 -- 
  +-- Quadrula_quadrula ** 
--+                     
  +-- Quadrula_verrucosa **

Amaro et al., 2017 [Publ 3791], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 3 14 0 0 -- 
  +----- Quadrula_quadrula__2 **
--+                          
  |  +-- Quadrula_quadrula__1 **
  +--+                       
     +-- Quadrula_nobilis **

Boyer et al., 2011 [Publ 2718], fig. 3

iTtTbiriSoTroTroSiD
2 6 0 2 33 4 3 8 
  +-------------- Quadrula_quadrula **  
--+                                  
  |  +----------- Quadrula_verrucosa ** 
  +--+                               
     |  +-------- Theliderma         
     +--+                            
        |  +----- Pustulosa_pustulosa
        +--+                         
           |  +-- Cyclonaias         
           +--+                      
              +-- Pustulosa_nodulata

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iTtTbiriSoTroTroSiD
2 15 0 2 13 0 0 17 
  +-------------------- Quadrula_quadrula **    
  |                                          
  +-------------------- Quadrula_verrucosa **   
  |                                          
--+                 +-- Eurynia              
  |  +--------------+                        
  |  |              +-- Pleurobema           
  |  |                                       
  +--+  +-------------- Amblema              
     |  |                                    
     |  |           +-- Obovaria             
     |  |  +--------+                        
     +--+  |        +-- Ortmanniana          
        |  |                                 
        |  |        +-- Obliquaria           
        |  +--------+                        
        +--+        +-- Lampsilis_ornata     
           |                                 
           |     +----- Lampsilis_radiata__2 
           |  +--+                           
           |  |  |  +-- Lampsilis_siliquoidea
           |  |  +--+                        
           +--+     +-- Lampsilis_radiata__1 
              |                              
              |  +----- Lampsilis_virescens  
              +--+                           
                 |  +-- Lampsilis_hydiana    
                 +--+                        
                    +-- Lampsilis_straminea

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 198 0 0 -- 
  +-- Quadrula_quadrula ** 
--+                     
  +-- Quadrula_verrucosa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
2 2 0 2 74 0 0 -- 
  +-- Quadrula_quadrula ** 
--+                     
  +-- Quadrula_verrucosa **

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iTtTbiriSoTroTroSiD
2 5 0 2 36 3 2 6 
  +----------- Quadrula_quadrula **  
--+                               
  |  +-------- Quadrula_verrucosa ** 
  +--+                            
     |  +----- Pustulosa_pustulosa
     +--+                         
        |  +-- Pustulosa_petrina  
        +--+                      
           +-- Cyclonaias

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Quadrula_quadrula **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
-- Quadrula_quadrula **

Turner et al., 2000 [Publ 2722], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Quadrula_quadrula **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 68 0 0 -- 
-- Quadrula_quadrula **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 43 0 0 -- 
-- Quadrula_quadrula **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Quadrula_quadrula **

Pfeiffer et al., 2018 [Publ 3835], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Quadrula_quadrula **

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Quadrula_quadrula **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Quadrula_quadrula **

Amaro et al., 2017 [Publ 3791], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Quadrula_quadrula **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Quadrula_quadrula **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 43 0 0 -- 
-- Quadrula_quadrula **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Quadrula_quadrula **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Quadrula_quadrula **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Quadrula_quadrula **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Quadrula_quadrula **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Quadrula_quadrula **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Quadrula_quadrula **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 0 0 -- 
-- Quadrula_quadrula **

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Quadrula_quadrula **

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Quadrula_quadrula **

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 42 5 2 -- 
           +-- Pustulosa_infucata 
  +--------+                      
  |        +-- Pustulosa_kleiniana
--+                               
  |  +-------- Quadrula_quadrula **  
  +--+                            
     |  +----- Cyclonaias         
     +--+                         
        |  +-- Pustulosa_succissa 
        +--+                      
           +-- Pustulosa_pustulosa

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Quadrula_quadrula **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Quadrula_quadrula **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Quadrula_quadrula **

Chapman et al., 2008 [Publ 2667], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Quadrula_quadrula **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Quadrula_quadrula **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 75 0 0 -- 
-- Quadrula_quadrula **

Curole & Kocher, 2005 [Publ 2682], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Inoue et al., 2020 [Publ 4128], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Quadrula_quadrula **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 31 0 0 -- 
-- Quadrula_quadrula **

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Quadrula_quadrula **

Freeman et al., 2025 [Publ 4443], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 10 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 0 0 -- 
-- Quadrula_quadrula **

Kandl et al., 2001 [Publ 3384], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 0 0 -- 
-- Quadrula_quadrula **

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Quadrula_quadrula **

Rosenberg et al., 1994 [Publ 2719], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 18 1 1 -- 
-- Quadrula_quadrula **

Freeman et al., 2025 [Publ 4443], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 0 0 -- 
-- Quadrula_quadrula **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Quadrula_quadrula **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Quadrula_quadrula **

Inoue et al., 2020 [Publ 3941], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Pustulosa        
--+                    
  +-- Quadrula_quadrula **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Quadrula_quadrula **

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Quadrula_quadrula **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 0 0 -- 
-- Quadrula_quadrula **

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Quadrula_quadrula **

Inoue et al., 2013 [Publ 3528], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Quadrula_quadrula **

Inoue et al., 2014 [Publ 3635], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Quadrula_quadrula **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Quadrula_quadrula **

Lydeard et al., 2000 [Publ 2654], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 13 2 1 -- 
  +-- Quadrula_nobilis **  
  |                     
--+-- Pustulosa_infucata
  |                     
  +-- Pustulosa_succissa

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Quadrula_quadrula **

Smith et al., 2020 [Publ 4129], fig. 2-3

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Quadrula_quadrula **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
-- Quadrula_quadrula **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 0 0 -- 
-- Quadrula_quadrula **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Quadrula_quadrula **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Quadrula_quadrula **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Quadrula_quadrula **

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Quadrula_quadrula **

Lydeard et al., 2000 [Publ 2654], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 13 2 1 -- 
  +-- Quadrula_nobilis **
--+                   
  +-- Pustulosa

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Quadrula_quadrula **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Quadrula_quadrula **

Curole & Kocher, 2002 [Publ 2712], fig. 1-F

iTtTbiriSoTroTroSiD
1 1 0 1 3 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Freeman et al., 2025 [Publ 4443], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Wen et al., 2017 [Publ 3795], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Quadrula_quadrula **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Quadrula_quadrula **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Quadrula_quadrula **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 0 0 -- 
-- Quadrula_quadrula **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 53 0 0 -- 
-- Quadrula_quadrula **

Inoue et al., 2020 [Publ 3941], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 41 1 1 -- 
  +-- Pustulosa        
--+                    
  +-- Quadrula_quadrula **

Burlakova et al., 2019 [Publ 3899], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 32 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 0 0 -- 
-- Quadrula_quadrula **

Chapman et al., 2008 [Publ 2744], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 20 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 9 0 0 -- 
-- Quadrula_quadrula **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Quadrula_quadrula **

Lydeard et al., 2000 [Publ 2654], fig. 2c

iTtTbiriSoTroTroSiD
1 1 0 1 13 2 1 -- 
  +-- Quadrula_nobilis **
--+                   
  +-- Pustulosa

Hauck et al., 2023 [Publ 4333], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Quadrula_quadrula **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 44 0 0 -- 
-- Quadrula_quadrula **

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 27 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Theliderma

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Quadrula_nobilis **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Quadrula_quadrula **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Quadrula_quadrula **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Quadrula_quadrula **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 37 0 0 -- 
-- Quadrula_quadrula **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Quadrula_quadrula **

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 10 1 1 -- 
-- Quadrula_quadrula **

Huang et al., 2013 [Publ 3629], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Quadrula_quadrula **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Quadrula_quadrula **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Quadrula_quadrula **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 39 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Quadrula_quadrula **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Quadrula_quadrula **

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Quadrula_quadrula **

Smith et al., 2020 [Publ 4129], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 27 0 0 -- 
-- Quadrula_quadrula **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Quadrula_quadrula **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 29 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 0 0 -- 
-- Quadrula_quadrula **

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

iTtTbiriSoTroTroSiD
1 1 0 1 3 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 41 5 2 -- 
  +----------- Quadrula_quadrula **  
  |                               
--+        +-- Pustulosa_infucata 
  |  +-----+                      
  |  |     +-- Pustulosa_kleiniana
  +--+                            
     |  +----- Cyclonaias         
     +--+                         
        |  +-- Pustulosa_succissa 
        +--+                      
           +-- Pustulosa_pustulosa

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Quadrula_quadrula **

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Quadrula_quadrula **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Quadrula_quadrula **

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
1 2 0 2 32 1 1 -- 
  +----- Quadrula_quadrula__2 **
--+                          
  |  +-- Quadrula_quadrula__1 **
  +--+                       
     +-- Theliderma

Lydeard et al., 1996 [Publ 2656], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 26 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Theliderma

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Quadrula_quadrula **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Quadrula_quadrula **

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 11 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Quadrula_quadrula **

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Quadrula_quadrula **

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 41 5 2 -- 
  +----------- Quadrula_quadrula **  
  |                               
--+        +-- Pustulosa_infucata 
  |  +-----+                      
  |  |     +-- Pustulosa_kleiniana
  +--+                            
     |  +----- Cyclonaias         
     +--+                         
        |  +-- Pustulosa_succissa 
        +--+                      
           +-- Pustulosa_pustulosa

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 0 0 -- 
-- Quadrula_quadrula **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 41 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Quadrula_quadrula **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Quadrula_quadrula **

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 41 5 2 -- 
  +----------- Quadrula_quadrula **  
  |                               
--+        +-- Pustulosa_infucata 
  |  +-----+                      
  |  |     +-- Pustulosa_kleiniana
  +--+                            
     |  +----- Cyclonaias         
     +--+                         
        |  +-- Pustulosa_succissa 
        +--+                      
           +-- Pustulosa_pustulosa

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 0 0 -- 
-- Quadrula_quadrula **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Quadrula_quadrula **

Xue et al., 2016 [Publ 3659], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 1 0 0 -- 
-- Quadrula_quadrula **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 0 0 -- 
-- Quadrula_quadrula **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Quadrula_quadrula **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Quadrula_quadrula **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 0 0 -- 
-- Quadrula_quadrula **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 0 0 -- 
-- Quadrula_quadrula **

Wei et al., 2016 [Publ 3665], fig. 0

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Quadrula_quadrula **

Inoue et al., 2020 [Publ 4128], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Quadrula_quadrula **

Smith et al., 2020 [Publ 4136], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Quadrula_quadrula **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 81 0 0 -- 
-- Quadrula_quadrula **

Lydeard et al., 2000 [Publ 2654], fig. 2d

iTtTbiriSoTroTroSiD
1 1 0 1 13 2 1 -- 
  +-- Quadrula_nobilis **  
  |                     
--+-- Pustulosa_infucata
  |                     
  +-- Pustulosa_succissa

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 0 0 -- 
-- Quadrula_quadrula **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Quadrula_quadrula **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Quadrula_quadrula **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 37 2 2 -- 
  +----- Quadrula_quadrula **
--+                       
  |  +-- Cyclonaias       
  +--+                    
     +-- Pustulosa

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Quadrula_quadrula **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 0 0 -- 
-- Quadrula_quadrula **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 0 0 -- 
-- Quadrula_quadrula **

Pieri et al., 2018 [Publ 3867], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

iTtTbiriSoTroTroSiD
1 1 0 1 3 1 1 -- 
  +-- Quadrula_quadrula **
--+                    
  +-- Pustulosa

Smith et al., 2019 [Publ 3951], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Quadrula_quadrula **

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Quadrula_quadrula **

Amaro et al., 2017 [Publ 3791], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Quadrula_quadrula **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 140 0 0 -- 
-- Quadrula_quadrula **

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Quadrula_quadrula **

Krebs, 2004 [Publ 2692], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 4 0 0 -- 
-- Quadrula_quadrula **

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 0 0 -- 
-- Quadrula_quadrula **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 0 0 -- 
-- Quadrula_quadrula **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Quadrula_quadrula **