Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina
8 Recent freshwater species (9 total species)
Frierson, 1927. Check List N. Amer. Naiades: 48.
(homonym of Jousseaume, 1875 [Gastr.]; intro'd as sub-gen. of Quadrula Rafinesque, 1820; renamed Pustulosa Frierson, 1927)
Type species: Obliquaria (Quadrula) bullata Rafinesque, 1820 (original designation) = Pustulosa pustulosa
Frierson, 1927. Check List N. Amer. Naiades: errata et corrigenda.
Thiele, 1934. Handb. Syst. Weichtierk. 3: 827 [1260].
(intro'd as sub-gen. of Quadrula Rafinesque, 1820; new name for Bullata Frierson, 1927)
Type species: Obliquaria (Quadrula) bullata Rafinesque, 1820 = Pustulosa pustulosa
The Taxonomic History lists the ways authors have classified the species currently assigned to Pustulosa, since 1900.
Obovaria — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) — Simpson (1900), Simpson (1914)
Unio (s.s.) — Call (1900)
Quadrula (sec. Fusconaia) — Simpson (1900), Simpson (1914)
Quadrula — Ihering (1901), Ortmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910), ... Inoue et al. (2018), Marshall et al. (2018), Chase et al. (2018), Klymus et al. (2020), Gasparini et al. (2020)
Unio — de Gregorio (1914), Henderson (1935)
Fusconaia — Ortmann (1923), Ortmann (1925), Clench & Turner (1956), Haas (1969), Burch (1975), ... Graf & Cummings (2007), Zhou et al. (2007), Cummings & Graf (2010), ITIS (2012), Szumowski et al. (2012)
Quadrula (Bullata) — Frierson (1927)
Quadrula (Quincuncina) — Frierson (1927)
Quadrula (Fusconia [sic]) — Frierson (1927)
Quadrula (Striata) — Frierson (1927)
Quadrula (s.s.) — F.C. Baker (1928)
Quincuncina — Clench & Turner (1956), Haas (1969), Burch (1975), Davis & Fuller (1981), Davis (1983), ... Graf & Cummings (2007), Zhou et al. (2007), Cummings & Graf (2010), ITIS (2012), Szumowski et al. (2012)
Pustulosa — Modell (1957), Neemuchwala et al. (2023), FMCS (2025)
Quadrula (Pustulosa) — Haas (1969)
Pleurobema — Morrison (1969), Vidrine (1993)
Obliquaria — Burch (1975)
Amphinaias — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010), IUCN (2011), ITIS (2013), Roe & Boyer (2015)
Rotundaria — Campbell & Lydeard (2012), Campbell & Lydeard (2012)
Cyclonaias — Williams et al. (2017), Johnson et al. (2018), Burlakova et al. (2018), Pieri et al. (2018), Lopes-Lima et al. (2019), Inoue et al. (2020), Graf & Cummings (2021), FMCS (2021), Bieler (2021), Hopper et al. (2023), Hauck et al. (2023), Freeman et al. (2025)
Cyclonaias (Quadrula) — Porto-Hannes et al. (2021)