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Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Lamprotulini

genus Lamprotula Simpson, 1900: cladomics

17 Recent valid species (26 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 17 (26 total)
analyzed in 137 cladograms in 86 publications
maximum number of terminals in an analysis = 3 (mean = 1.64, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 67
maximum number of included terminals = 3, (mean = 2.3 median = 2)
proportion of cladograms that support basic monophyly = 0.93

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lamprotula cladogram distances
PCoA Ordination

NJ Tree of Lamprotula cladogram distances
NJ Tree

Cladograms

Huang et al., 2019 [Publ 3843], fig. S1

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
3 3 2 6 44 0 0 0 
  +-------- Lamprotula_leaii **          
  |                                   
--+  +----- Lamprotula_cornuumlunae__1 **
  |  |                                
  |  +----- Lamprotula_cornuumlunae__3 **
  +--+                                
     +----- Lamprotula_cornuumlunae__4 **
     |                                
     |  +-- Lamprotula_cornuumlunae__2 **
     +--+                             
        +-- Lamprotula_caveata **

Froufe et al., 2016 [Publ 3727], fig. 3_4

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
1 1 0 1 6 4 2 -- 
-- Lamprotula_leaii **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
3 3 1 3 89 2 2 3 
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
3 3 1 4 42 0 0 4 
  +----- Lamprotula_leaii **       
  |                             
--+  +-- Lamprotula_caveata__1 **  
  |  |                          
  +--+-- Lamprotula_cornuumlunae **
     |                          
     +-- Lamprotula_caveata__2 **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 51 2 2 3 
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
3 3 1 3 11 1 1 3 
  +-------- Potomida               
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
3 3 1 3 93 2 2 3 
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
3 3 2 4 27 1 1 4 
  +----------- Pronodularia           
--+                                   
  |  +-------- Lamprotula_leaii **       
  +--+                                
     |  +----- Lamprotula_caveata__1 **  
     +--+                             
        |  +-- Lamprotula_cornuumlunae **
        +--+                          
           +-- Lamprotula_caveata__2 **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
3 3 1 3 93 2 2 3 
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 198 2 2 3 
        +-- Potomida               
  +-----+                          
  |     +-- Pronodularia           
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
3 3 1 3 198 2 2 3 
        +-- Potomida               
  +-----+                          
  |     +-- Pronodularia           
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iTtTbiriSoTroTroSiD
3 3 2 4 5 1 1 5 
  +----------- Pronodularia           
--+                                   
  |  +-------- Lamprotula_caveata **     
  +--+                                
     |  +----- Lamprotula_leaii__2 **    
     +--+                             
        |  +-- Lamprotula_seomjinensis **
        +--+                          
           +-- Lamprotula_leaii__1 **

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 15 0 0 3 
  +----- Lamprotula_leaii **       
--+                             
  |  +-- Lamprotula_caveata **     
  +--+                          
     +-- Lamprotula_cornuumlunae **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
3 3 1 3 25 2 2 3 
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
3 3 2 5 119 2 2 4 
  +-------------- Pronodularia           
  |                                      
--+  +----------- Potomida               
  |  |                                   
  +--+  +-------- Lamprotula_caveata **     
     |  |                                
     +--+  +----- Lamprotula_leaii__2 **    
        |  |                             
        +--+----- Lamprotula_leaii__3 **    
           |                             
           |  +-- Lamprotula_seomjinensis **
           +--+                          
              +-- Lamprotula_leaii__1 **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
3 3 1 3 196 2 2 3 
        +-- Potomida               
  +-----+                          
  |     +-- Pronodularia           
--+                                
  |  +----- Lamprotula_leaii **       
  +--+                             
     |  +-- Lamprotula_caveata **     
     +--+                          
        +-- Lamprotula_cornuumlunae **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
3 5 0 3 109 2 2 5 
  +-- Lamprotula_leaii **  
  |                     
  +-- Lamprotula **        
  |                     
--+-- Lamprotula_caveata **
  |                     
  +-- Potomida          
  |                     
  +-- Pronodularia

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
3 16 0 3 115 2 2 16 
  +-- Sinosolenaia_iridinea    
  |                            
  +-- Sinosolenaia_recognita__1
  |                            
  +-- Sinosolenaia_carinata    
  |                            
  +-- Leguminaia               
  |                            
  +-- Microcondylaea           
  |                            
  +-- Gonidea                  
  |                            
  +-- Ptychorhynchus           
  |                            
  +-- Parvasolenaia            
  |                            
--+-- Lamprotula_leaii **         
  |                            
  +-- Lamprotula **               
  |                            
  +-- Lamprotula_caveata **       
  |                            
  +-- Potomida                 
  |                            
  +-- Pronodularia             
  |                            
  +-- Sinohyriopsis            
  |                            
  +-- Chamberlainia            
  |                            
  +-- Sinosolenaia_recognita__2
  |                            
  +-- Sinosolenaia_oleivora

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
3 4 2 3 18 0 0 4 
  +-------- Lamprotula_leaii **       
--+                                
  |  +----- Lamprotula_caveata **     
  +--+                             
     |  +-- Gibbosula              
     +--+                          
        +-- Lamprotula_cornuumlunae **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
3 3 2 5 28 2 2 3 
  +-------------- Pronodularia           
  |                                      
--+  +----------- Potomida               
  |  |                                   
  +--+  +-------- Lamprotula_leaii **       
     |  |                                
     +--+  +----- Lamprotula_caveata__3 **  
        |  |                             
        +--+  +-- Lamprotula_cornuumlunae **
           |  |                          
           +--+-- Lamprotula_caveata__1 **  
              |                          
              +-- Lamprotula_caveata__2 **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
3 3 1 3 89 2 2 3 
  +----------- Potomida               
--+                                   
  |  +-------- Pronodularia           
  +--+                                
     |  +----- Lamprotula_leaii **       
     +--+                             
        |  +-- Lamprotula_caveata **     
        +--+                          
           +-- Lamprotula_cornuumlunae **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
2 2 0 2 35 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
2 2 0 2 35 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 72 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 18 3 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 46 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
2 2 0 2 35 1 1 -- 
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Lopes-Lima et al., 2020 [Publ 4080], fig. 14

iTtTbiriSoTroTroSiD
2 2 0 2 4 0 0 -- 
  +-- Lamprotula_leaii **       
--+                          
  +-- Lamprotula_seomjinensis **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 54 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 36 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_leaii **  
  +--+                     
     +-- Lamprotula_caveata **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 51 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Zhang et al., 2024 [Publ 4397], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 6 0 0 -- 
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
2 2 0 2 89 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 137 2 2 -- 
     +-- Potomida          
  +--+                     
  |  +-- Pronodularia      
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
2 2 0 2 20 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 43 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 32 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 72 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 52 2 2 -- 
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Pronodularia      
  +--+                     
     +-- Potomida

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 36 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
2 2 0 2 46 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 18 3 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 72 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 10 0 0 -- 
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 42 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
2 2 0 2 137 2 2 -- 
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Potomida          
  +--+                     
     +-- Pronodularia

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 35 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 86 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
2 34 0 2 32 0 0 33 
                       +----- Prolasmidonta           
                    +--+                              
                    |  |  +-- Alasmidonta             
                    |  +--+                           
                    |     +-- Anemina                 
  +-----------------+                                 
  |                 |     +-- Lamprotula_caveata **      
  |                 |  +--+                           
  |                 |  |  +-- Chamberlainia           
  |                 +--+                              
  |                    |  +-- Pseudodon               
  |                    +--+                           
  |                       +-- Pilsbryoconcha          
  |                                                   
  |                    +----- Schistodesmus           
  |              +-----+                              
  |              |     |  +-- Lepidodesma             
--+              |     +--+                           
  |              |        +-- Aculamprotula           
  |  +-----------+                                    
  |  |           |     +----- Platynaias              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon               
  |  |              |                                 
  |  |              |  +----- Sinanodonta             
  |  |              +--+                              
  |  |                 |  +-- Cristaria               
  +--+                 +--+                           
     |                    +-- Anodonta                
     |                                                
     |                    +-- Utterbackia_peninsularis
     |           +--------+                           
     |           |        +-- Potamilus_alatus        
     |  +--------+                                    
     |  |        |  +-------- Lamprotula_leaii **        
     |  |        +--+                                 
     |  |           |  +----- Sinohyriopsis           
     |  |           +--+                              
     +--+              |  +-- Parvasolenaia           
        |              +--+                           
        |                 +-- Sinosolenaia            
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrula                
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
2 2 0 2 32 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
2 13 0 2 123 2 2 12 
  +-- Chamberlainia         
  |                         
  +-- Sinohyriopsis         
  |                         
  +-- Lamprotula_caveata **    
  |                         
  +-- Lamprotula_leaii **      
  |                         
  +-- Potomida              
  |                         
  +-- Pronodularia          
  |                         
--+-- Gonidea               
  |                         
  +-- Leguminaia_wheatleyi  
  |                         
  +-- Leguminaia            
  |                         
  +-- Microcondylaea        
  |                         
  +-- Sinosolenaia_carinata 
  |                         
  +-- Sinosolenaia_oleivora 
  |                         
  +-- Sinosolenaia_recognita

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
2 2 0 2 78 2 2 -- 
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Potomida          
  +--+                     
     +-- Pronodularia

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 89 2 2 -- 
  +-------- Pronodularia      
--+                           
  |  +----- Potomida          
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
2 2 0 2 35 1 1 -- 
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 84 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 13 0 0 -- 
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 43 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 40 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
2 2 0 2 54 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 42 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_leaii **  
     +--+                     
        +-- Lamprotula_caveata **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 67 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 51 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 45 2 2 -- 
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Pronodularia      
  +--+                     
     +-- Potomida

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 22 2 2 -- 
     +-- Lamprotula_caveata **
  +--+                     
  |  +-- Lamprotula_leaii **  
--+                        
  |  +-- Potomida          
  +--+                     
     +-- Pronodularia

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 29 1 1 -- 
  +----- Potomida          
--+                        
  |  +-- Lamprotula_caveata **
  +--+                     
     +-- Lamprotula_leaii **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 14 0 0 -- 
  +-- Lamprotula_caveata **
--+                     
  +-- Lamprotula_leaii **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
2 2 0 2 39 2 2 -- 
  +-------- Potomida          
--+                           
  |  +----- Pronodularia      
  +--+                        
     |  +-- Lamprotula_caveata **
     +--+                     
        +-- Lamprotula_leaii **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 28 0 0 -- 
-- Lamprotula_cornuumlunae **

Kongim et al., 2023 [Publ 4290], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 2 2 -- 
  +----- Lamprotula_leaii **
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Potomida

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
1 1 0 1 38 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Konopleva et al., 2021 [Publ 4213], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 26 1 1 -- 
-- Lamprotula_leaii **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 20 1 1 -- 
  +-- Lamprotula_leaii **
--+                   
  +-- Pronodularia

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 0 0 -- 
-- Lamprotula_leaii **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
1 1 0 1 81 1 1 -- 
  +-- Lamprotula_leaii **
--+                   
  +-- Potomida

Goncalves et al., 2022 [Publ 4253], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Lamprotula_caveata **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 -- 
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 13 0 0 -- 
-- Lamprotula_caveata **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Lamprotula_cornuumlunae **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Lamprotula_leaii **

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 7 0 0 -- 
-- Lamprotula_leaii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +----- Lamprotula_leaii **
--+                      
  |  +-- Potomida        
  +--+                   
     +-- Pronodularia

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Lamprotula_leaii **
--+                   
  +-- Pronodularia

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 0 0 -- 
-- Lamprotula_cornuumlunae **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 36 1 1 -- 
  +-- Lamprotula_leaii **
--+                   
  +-- Potomida

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Lamprotula_leaii **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
-- Lamprotula_leaii **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
1 1 0 1 22 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 14 0 0 -- 
-- Lamprotula_leaii **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
1 1 0 1 16 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 67 2 2 -- 
  +----- Lamprotula_leaii **
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Potomida

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
1 1 0 1 48 1 1 -- 
-- Lamprotula_leaii **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 -- 
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 3 0 0 -- 
-- Lamprotula_leaii **

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 10 0 0 -- 
-- Lamprotula_leaii **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 39 1 1 -- 
  +-- Lamprotula_caveata **
--+                     
  +-- Potomida

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 23 0 0 -- 
-- Lamprotula_leaii **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 40 2 2 -- 
  +----- Pronodularia    
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Potomida

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 23 0 0 -- 
-- Lamprotula_leaii **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 82 2 2 -- 
-- Lamprotula_leaii **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 1 1 -- 
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 15 1 1 -- 
-- Lamprotula_leaii **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 15 0 0 -- 
-- Lamprotula_leaii **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Lamprotula_leaii **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Lamprotula_leaii **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 28 1 1 -- 
  +-- Potomida        
--+                   
  +-- Lamprotula_leaii **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 13 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
-- Lamprotula_leaii **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 12 0 0 -- 
-- Lamprotula_caveata **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 57 2 2 -- 
  +----- Pronodularia    
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Potomida

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 31 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Lamprotula_leaii **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 19 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 2 2 -- 
  +----- Potomida          
--+                        
  |  +-- Pronodularia      
  +--+                     
     +-- Lamprotula_caveata **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
  +-- Lamprotula_caveata **
--+                     
  +-- Pronodularia

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
1 1 0 1 21 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Lamprotula_leaii **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 67 2 2 -- 
  +----- Lamprotula_leaii **
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Potomida

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
1 1 0 1 12 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 11 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 48 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Pronodularia    
  +--+                   
     +-- Lamprotula_leaii **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 21 1 1 -- 
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 22 0 0 -- 
-- Lamprotula_leaii **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 40 2 2 -- 
  +----- Pronodularia    
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Potomida

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
  +-- Lamprotula_leaii **
--+                   
  +-- Potomida

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 0 0 -- 
-- Lamprotula_leaii **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 49 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +----- Potomida        
--+                      
  |  +-- Lamprotula_leaii **
  +--+                   
     +-- Pronodularia

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
1 1 0 1 12 2 2 -- 
  +----- Lamprotula_leaii **
--+                      
  |  +-- Potomida        
  +--+                   
     +-- Pronodularia

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 67 2 2 -- 
  +----- Lamprotula_leaii **
--+                      
  |  +-- Potomida        
  +--+                   
     +-- Pronodularia

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 32 1 1 -- 
  +-- Pronodularia    
--+                   
  +-- Lamprotula_leaii **

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 6 0 0 -- 
-- Lamprotula_leaii **
 
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