MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Tue 05 May 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini
genus Potamilus Rafinesque, 1818: cladomics
10 Recent valid species (11 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 10 (11 total)
analyzed in 109 cladograms in 63 publications
maximum number of terminals in an analysis = 10 (mean = 2.14, median = 2)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 56
maximum number of included terminals = 10, (mean = 3.21 median = 2)
proportion of cladograms that support basic monophyly = 0.77
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 582 | 651 | 15 | 622 | 83 | 26 | 0 |
+----- Amuranodonta
+-----------------------------------+
| | +-- Anemina
| +--+
| +-- Buldowskia
|
| +-- Beringiana
| +-----+
| | +-- Pletholophus
| +-----------------------------+
+-----------------------+ | | +----- Mimo
| | | +--+
| | | | +-- Acudonta
| | | +--+
| | | +-- Sinanodonta
| | |
| | | +----------------------------------- Cristaria
| | | |
| | | | +----------------------------- Lepidodesma
| +--+ | |
| | | | +----------- Lanceolaria
| | | | +--------------+
| | | +--+ | | +-------- Pseudoacuticosta_retiaria__2
| | | | | | +--+
| | | | | | | +----- Acuticosta_chinensis__2
| | | | | | +--+
| | | | +--+ | +-- Acuticosta_chinensis__1
| | | | | +--+
| | | | | +-- Pseudoacuticosta_retiaria__1
| +--+ | |
| | | | +----------------------- Globunio
| | | | |
| | | +--+ +-------------------- Scriptolamprotula
| | | | |
| | | +--+ +----------------- Middendorffinaia
| | | | |
| | | +--+ +-------------- Unio
| | | | |
| | | | | +-- Schistodesmus
| | | +--+ +--------+
| | | | | +-- Nodularia
| | | | |
| | | +--+ +-- Tchangsinaia
| +--+ | +-----+
| | | | +-- Inversiunio
| | +--+
| | | +----- Pseudocuneopsis
| | +--+
| | | +-- Diaurora
| | +--+
| | +-- Cuneopsis
| |
| | +-------- Pseudodontoideus
| | +--------------------+
| | | | +----- Lasmigona
| | | +--+
| | | | +-- Pterosyna
| | | +--+
| | | +-- Arcidens
| | |
| | | +----------------------- Anodonta_anatina
| | | |
+-----------------------+ | | +--+ +-------------------- Pseudanodonta
| | | | | | |
| | +--+ | +--+ +----------------- Brachyanodon_oregonensis__2
| | | | | |
| | | | | | +----- Anodonta_cygnea
| | | | +--+ +--------+
| | | | | | | +-- Anodonta_seddonae
| | | | | | +--+
| | | | +--+ +-- Anodonta_exulcerata
| | | | |
| | | | | +----------- Brachyanodon_impura
| | | | | |
| | | | +--+ +-------- Brachyanodon
| | | | | |
| | +--+ +--+ +----- Brachyanodon_nuttalliana__1
| | | | |
| | | +--+----- Brachyanodon_nuttalliana__2
| | | |
| | | | +-- Brachyanodon_californiensis
| | | +--+
| | | +-- Brachyanodon_oregonensis__1
| | |
| | | +-------------- Utterbackia
| | | +--+
| | | | | +----------- Pyganodon_grandis
| | | | +--+
| | | | | +-------- Pyganodon_lacustris
| | | | +--+
| | | | | +----- Pyganodon_cataracta__2
| | +--------+ +--+
| | | | +-- Pyganodon_cataracta__1
| | | +--+
| | | +-- Potamilus_fragilis__3 **
| | |
| | | +-------------- Utterbackiana
| | | |
| | +--+ +----------- Simpsonaias
| | | |
| | | | +-- Alasminota
| | +--+ +-----+
| | | | +-- Alasmidonta
| | | |
| | +--+ +-- Prolasmidonta
| | | +--+
| | | | +-- Platynaias
| | +--+
| | | +-- Anodontoides
| | +--+
| | +-- Strophitus
| |
| | +-------------------------------------------------------------- Lamellidens
| | |
| | | +-- Balwantia
| +--+ +--------------------------------------------------------+
| | | +-- Trapezidens
| | |
| +--+ +-------------------------------------------------------- Leoparreysia
| | |
| | | +----------------------------------------------------- Pseudomulleria
| | | |
| +--+ | +-- Pseudoscabies
| | | +--------------+
| | | | +-- Indonaia
| | | +--------------------------------+
| | | | | +-------------- Indochinella
| +--+ | | |
| | | +--+ +----------- Radiatula
| | | | |
| | | +--+ +-- Songkhlanaia_songkhramensis__2
| | | | +-----+
| | | | | +-- Unionetta_fabagina__2
| | | +--+
| | | | +----- Harmandia
| +--+ +--+
| | | +-- Unionetta_fabagina__1
| | +--+
| | +-- Scabies
| |
| | +-- Assamnaia
| | +--------------------------------------------+
| | | +-- Parreysia
| | |
| | | +----- Nitia_monceti__2
| +--+ +--------------------------------------+
| | | | +-- Nitia_teretiuscula
| | | +--+
| | | +-- Nitia_monceti__1
| +--+
| | +----------------------------------------- Grandidieria
| | |
| | | +----- Pseudospatha_tanganyicensis__2
| | | +--------------------------------+
| +--+ | | +-- Moncetia
| | | +--+
| | | +-- Pseudospatha_tanganyicensis__1
| | |
| | | +----- Coelatura_leopoldvillensis__2
| +--+ +-----------------------------+
| | | | +-- Coelatura_leopoldvillensis__1
| | | +--+
| | | +-- Coelatura_mesafricana
| | |
| | | +-------------------------------- Coelatura_aegyptiaca__3
| | | |
| +--+ | +-------- Coelatura_hauttecoeuri__2
| | | |
| | | +-------- Nitia_acuminata
| | | |
| | | | +----- Coelatura_aegyptiaca__2
| | | +--------------------+ |
| | | | | +----- Coelatura_alluaudi__2
| +--+ | | |
| | | | +----- Coelatura_hauttecoeuri__1
| | | | |
| | | +--+----- Coelatura_stuhlmanni__2
--+ | | |
| | | | +-- Coelatura_aegyptiaca__1
| | | | |
| | | | +-- Coelatura_stuhlmanni__1
| | | | |
| | | +--+-- Coelatura_alluaudi__1
| +--+ |
| | +-- Coelatura_bakeri__1
| | |
| | +-- Coelatura_bakeri__2
| |
| | +----------- Coelatura_horei__3
| | |
| | +--------------+ +-------- Coelatura_ratidota
| | | | |
| | | +--+ +----- Coelatura__2
| | | | |
| | | +--+ +-- Coelatura_horei__1
| | | | |
| +--+ +--+-- Coelatura__1
| | |
| | +-- Coelatura_horei__2
| |
| | +----------------------- Coelatura_gabonensis
| | |
| +--+ +-------------------- Coelatura_stanleyvillensis
| | |
| | | +----------------- Coelatura_kunenensis
| +--+ |
| | | +-------------- Nyassunio
| | | |
| +--+ | +----------- Coelatura_hypsiprymna__2
| | | |
| | | +----------- Coelatura_mossambicensis__1
| | | |
| +--+ | +-- Coelatura_hypsiprymna__3
| | +--------+
| | | +-- Coelatura_mossambicensis__2
| | |
| | | +----- Coelatura_luapulaensis__2
| | | |
| | | +----- Coelatura_choziensis__2
| +--+-----+
| | +----- Coelatura_hypsiprymna__1
| | |
| | | +-- Coelatura_choziensis__1
| | +--+
| | +-- Coelatura_luapulaensis__1
| |
| | +----- Coelatura_briarti
| | +--+
| | | | +-- Prisodontopsis_aviculaeformis__1
| | | +--+
| +--+ +-- Prisodontopsis_mweruensis__1
| |
| | +-- Prisodontopsis_aviculaeformis__2
| | |
| +-----+-- Prisodontopsis_aviculaeformis__3
| |
| +-- Prisodontopsis_mweruensis__2
|
| +-------------------------- Khairuloconcha
| |
| | +----------------------- Ctenodesma
| | |
| +-----------------------------------------------------------+ | +-- Physunio_modelli
| | | | +-----------+
| | | | | +-- Physunio_pellucidus
| | | | |
| | +--+ +-----+ +-- Physunio_massini
| | | | | +--------+
| | | | | | +-- Physunio_superbus
| | | | +--+
| | | | | +-------- Physunio
| | | | +--+
| | | | | +----- Trapezoideus
| | +--+ +--+
| | | | +-- Yaukthwa
| | | +--+
| | | +-- Lens
| | |
| | | +----------------- Ensidens
| | | |
| | | | +----- Rectidens
| | +--+ +--------+
| | | | | +-- Hyriopsis_delaportei
| | | | +--+
| | | | +-- Hyriopsis_desowitzi
| | +--+
| | | +----------- Hyriopsis_panhai
| | | |
| | | | +-- Hyriopsis_bialata
| | +--+ +-----+
| | | | +-- Hyriopsis_khoratensis
| | | |
| | +--+ +-- Hyriopsis_kratiensis
| | | +--+
| | | | +-- Hyriopsis_phuphaniensis
| | +--+
| | | +-- Hyriopsis_myersiana
| | +--+
| | +-- Hyriopsis_sakhonensis
| |
| | +-------- Pseudunio
| | +--------------------------------------------------------------------------+
| | | | +----- Gibbosula
| | | +--+
| | | | +-- Cumberlandia
| | | +--+
| | | +-- Margaritifera
| | |
| | | +----------------------------- Indopseudodon
| | | |
| | | | +-- Lannanaia
| | | +--+ +-----------------------+
+--+ | | | | +-- Pilsbryoconcha
| | | | |
| | | | | +----- Songkhlanaia_tamodienica
| | | +--+ +-----------------+
| | | | | | +-- Songkhlanaia_moreleti
| | | | | +--+
| | | | | +-- Songkhlanaia_songkhramensis__1
| | | | |
| | | +--+ +-- Nyeinchanconcha
| | | | +-----------------+
| | | | | +-- Sundadontina
| | | | |
| | | | | +-- Isannaia
| | | +--+ +--+
| | | | | +-- Namkongnaia
| | | | +-----------+
| | | | | | +-- Thaiconcha
| | | | | +--+
| | | +--+ +-- Bineurus
| | | |
| | | | +-------------- Pseudodon_walpolei
| | | | |
| | | +--+ +----------- Pseudodon_vagulus
| | | | |
| | +-----------------------------------------------+ | | +----- Pseudodon_vondembuschianus
| | | | +--+ +--+
| | | | | | | +-- Pseudodon_mekongi
| | | | | | +--+
| | | | +--+ +-- Schepmania
| | | | |
| | | | | +----- Pseudodon_lenyanensis__2
| | | | +--+
| | | | | +-- Pseudodon_inoscularis
| | | | +--+
| | | | +-- Pseudodon_lenyanensis__1
| | | |
| | | | +-- Chamberlainia
| | | | +-----------------+
| | | | | +-- Sinohyriopsis
| | | | +--+
| | | | | | +----------------- Parvasolenaia
| | | | | | |
| | | | | +--+ +-------------- Pseudopostolata
| | | | | | |
| | | | | +--+ +----------- Koreosolenaia
| | | | | | |
| | | | | +--+ +-- Cosmopseudodon
| | | | | | +-----+
| | | | | | | +-- Guiunio
| | | | | +--+
+--+ | +--------+ | +----- Ptychorhynchus
| | | +--+
| | | | +-- Obovalis
| | | +--+
| | | +-- Postolata
| | |
| | | +-------- Caudiculatus
| | | +--------+
| | | | | +----- Lamprotula
| | | | +--+
| | | | | +-- Pronodularia
| | | | +--+
| | +-----+ +-- Potomida
| | |
| | | +-------------- Sinosolenaia_carinata
| | | |
| | | | +----- Gonidea
| | +--+ +-----+
| | | | | +-- Microcondylaea
| | | | +--+
| | +--+ +-- Leguminaia
| | |
| | | +-------- Inversidens
| | +--+
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
| |
| | +----- Amblema
| | +--------------------------------------------------------------------+
| | | | +-- Nephronaias
| | | +--+
| | | +-- Psoronaias
| | |
| | +--+ +----------------------------------------------------------------------- Reginaia
| | | | |
| | | | | +----- Popenaias
| | | | | +--------------------------------------------------------------+
| | | | | | | +-- Cyrtonaias
| | | +--+ | +--+
| | | | | +-- Glebula
| | | | |
| | | | | +----------------------------------------------------------------- Paetulunio
| | | | | |
| | | +--+ | +----------------- Toxolasma_parvum
| | | | | |
| | | | | | +----- Toxolasma_lividum__2
| | | | | | +--------+
| | | | | +--------------------------------------------+ | | +-- Toxolasma_cylindrellus
| | | | | | | | +--+
| | | | | | | | +-- Toxolasma_lividum__1
| | | | | | | |
| | | +--+ | +--+ +-------- Toxolasma_texasiense__2
| | | | | | +--+
| | | | | | | | +----- Toxolasma_glans
| | | | | | | +--+
| | | | | | | | +-- Toxolasma_paulum
| | | | | +--+ +--+
| | | | | | +-- Toxolasma_texasiense__1
| | | | | |
+--+ | | | | +-------- Toxolasma__4
| | | | +--+
| | +--+ | +----- Toxolasma_granulatum
| | | +--+
| | | | +-- Toxolasma__2
| | | +--+
| | | +-- Toxolasma__3
| | |
| | | +----- Toxolasma_pullus
| | | +-----------------------------------------------------+
| | | | | +-- Toxolasma_germanum
| | | | +--+
| | | | +-- Toxolasma__1
| | | |
| | | | +-- Obliquaria
| | | | +-----------------------------------------------------+
| | | | | +-- Plectomerus
| | +--+ |
| | | | +-------- Potamilus_inflatus **
| | | | +--------+
| | | | | | +----- Potamilus_ohiensis **
| | | | | +--+
| | | | | | +-- Potamilus_amphichaenus **
| | | | | +--+
| | | | +-----------------------------------+ +-- Potamilus_streckersoni **
| | | | | |
| | +--+ | | +----- Potamilus_fragilis__2 **
| | | | | +--------+
| | | | | | | +-- Potamilus_leptodon **
| | | | | | +--+
| | | | +--+ +-- Potamilus_fragilis__1 **
| | | | |
| | | | | +----------- Potamilus_metnecktayi **
| | | | | |
| | | | +--+ +-------- Potamilus_alatus__3 **
| | | | | |
| | | | +--+ +----- Potamilus_alatus__2 **
| | +--+ | |
| | | +--+----- Potamilus_purpuratus__2 **
| | | |
| | | | +-- Potamilus_alatus__1 **
| | | +--+
| | | +-- Potamilus_purpuratus__1 **
| | |
| | | +-- Villosa_villosa
| | | +-----------------------------------------------+
| | | | +-- Atlanticoncha
| | | |
| | | | +-- Cambarunio_nebulosus
| | | | +--------------------------------------------+
| | +--+ | +-- Ligumia
| | | |
| | | | +-------- Leaunio_umbrans
| | | | +-----------------------------------+
| | | | | | +----- Cambarunio_taeniatus
| | +--+ | +--+
| | | | | +-- Cambarunio_iris
| | | | +--+
| | | | +-- Cambarunio_simus
| | | |
| | | | +----------------------------------------- Hamiota
| | | | |
| | +--+ | +----------------------- Pachynaias
| | | | |
| | | | +--------------+ +-------------------- Sagittunio
| | | | | | |
| | | | | | | +----- Ellipsaria
| | | | | +--+ +-----------+
| | | | | | | | +-- Potamilus_capax **
| | | | | | | +--+
| | | | | | | +-- Truncilla
| | +--+ | +--+
| | | | | +----- Medionidus_acutissimus
| | | | | +--------+
| | | | | | | +-- Lemiox
| | | | | | +--+
| | | | +--+ +-- Medionidus_penicillatus
| | | | |
| | | | | +----------- Ptychobranchus
| | | | | |
| | | | +--+ +-- Cyprogenia_aberti__2
| | | | | +-----+
| | +--+ | | +-- Dromus
| | | +--+
| | | | +----- Cyprogenia_stegaria__2
| | | +--+
| | | | +-- Cyprogenia_aberti__1
| | | +--+
| | | +-- Cyprogenia_stegaria__1
+--+ |
| | +-- Villosa_delumbis
| | +--------------------------------+
| | | +-- Villosa_vibex
| | |
| | | +-------------- Obovaria_olivaria
| | | |
| | | +-----------------+ +-- Venustaconcha
| | | | | +--------+
| | | | | | +-- Epioblasma
| | | | +--+
| | | | | +-------- Obovaria_retusa
| +--+ | +--+
| | | | +----- Obovaria_choctawensis
| | | +--+
| | | | +-- Obovaria_unicolor
| | | +--+
| | | +-- Obovaria_subrotunda
| | |
| | | +-- Mesonaias
| | | +-----------------------+
| | | | +-- Obovaria_arkansasensis
| | | |
| +--+ +--+ +-- Lampsilis_splendida
| | | | +--------------------+
| | | | | +-- Lampsilis_bracteata
| | | | |
| | | +--+ +-- Lampsilis_fasciola
| | | | +-----------------+
| | | | | +-- Ortmanniana_pectorosa
| | | +--+
| | | | +----------------- Lampsilis_ornata
| | | | |
| | | +--+ +-------------- Lampsilis_ovata__2
| | | | |
| | | +--+ +----------- Lampsilis_dolabraeformis
| | | | |
| | | | | +----- Lampsilis_cardium__2
| | | +--+ +--+
| | | | | | +-- Lampsilis_cardium__1
| +--+ | | +--+
| | +--+ +-- Lampsilis_satura
| | |
| | | +----- Lampsilis_cariosa__2
| | +--+
| | | +-- Lampsilis_ovata__1
| | +--+
| | +-- Lampsilis_cariosa__1
| |
| | +----------- Ortmanniana_ligamentina__2
| | +--------------+
| | | | +-------- Lampsilis_abrupta
| | | +--+
| | | | +----- Lampsilis_higginsii__2
| | | +--+
| | | | +-- Ortmanniana_ligamentina__1
| | | +--+
| +--+ +-- Lampsilis_higginsii__1
| |
| | +----- Medionidus_conradicus
| | +-----------------+
| | | | +-- Leaunio_lienosus
| | | +--+
| | | +-- Leaunio_vanuxemensis
| | |
| +--+ +-- Lampsilis_floridensis
| | +-----------------+
| | | +-- Lampsilis_teres
| | |
| | | +-- Lampsilis_siliquoidea__1
| +--+ |
| | +-- Lampsilis_siliquoidea__2
| | +--------------+
| | | +-- Lampsilis_powellii__1
| | | |
| +--+ +-- Lampsilis_powellii__2
| |
| | +-------------- Lampsilis_sietmani
| | |
| +--+ +-- Lampsilis_straminea__2
| | +--------+
| | | +-- Lampsilis_virescens
| +--+
| | +-------- Lampsilis_hydiana__2
| +--+
| | +----- Lampsilis_hydiana__1
| +--+
| | +-- Lampsilis_bergmanni
| +--+
| +-- Lampsilis_straminea__1
|
| +-- Megalonaias
| +-----------------+
| | +-- Uniomerus
| |
| | +-- Theliderma_johnsoni
| | +--------+
| | | +-- Theliderma_metanevra
| | |
| +--------------------------------------------+ +-----+ +-- Quadrula_nobilis
| | | | | +-----+
| | | | | | +-- Quadrula_verrucosa
| | | | +--+
| | | | | +----- Theliderma_cylindrica
| | | | +--+
| | | | | +-- Quadrula_fragosa
| | +--+ +--+
| | | +-- Quadrula_quadrula
| | |
| | | +----- Pustulosa_kieneriana
| | | +--------+
| | | | | +-- Pustulosa_infucata
| | | | +--+
| | +--+ +-- Pustulosa_kleiniana
| | |
+-----------+ | +----------- Cyclonaias
| | |
| +--+ +-- Pustulosa_pustulosa
| | +-----+
| | | +-- Pustulosa_succissa
| +--+
| | +----- Pustulosa_nodulata
| +--+
| | +-- Pustulosa_necki
| +--+
| +-- Pustulosa_petrina
|
| +-------------------------------------------------------------- Elliptoideus
| |
| | +----------------------------------------------------------- Parvaspina
| | |
| | | +-- Elliptio_fisheriana
+--+ | +-----------------------------------------------------+
| | | +-- Elliptio_nasutidus
| | |
| | | +----------------------------------------------------- Elliptio_spinosa
| | | |
+--+ | | +----------------------------- Elliptio_dariensis__5
| | | |
| | | | +----- Elliptio_arctata__3
| | | | |
| | | | +----- Elliptio_crassidens__4
| | | | +--------------------+
| | | | | +----- Elliptio_mcmichaeli__2
| | | | | |
| | | | | | +-- Elliptio_crassidens__5
+--+ | | | +--+
| | +--------------------+ | +-- Elliptio_roanokensis
| | | | |
| | | | | +----- Elliptio_complanata__2
| | | | | |
| | | | | +----- Elliptio_pullata__2
| | | | | |
| | | | | +----- Elliptio_icterina__3
| | | | | |
| | | | +--------------------+ +-- Elliptio_dariensis__4
| | | | | +--+
| | | | | | +-- Elliptio_jayensis__2
| | | +--+ |
| | | | | +-- Elliptio_congaraea__2
+--+ | | | |
| | | | +-- Elliptio_crassidens__3
| | | +--+
| | | +-- Elliptio_fumata__2
| | | |
| | | +-- Elliptio_mcmichaeli__1
| | |
| | | +----------------------- Elliptio_crassidens__2
| | | |
| | | | +-- Elliptio_arca__2
| | | +--------------------+
| | | | +-- Elliptio_crassidens__1
| | | |
| | | | +-------- Elliptio_icterina__2
| | | | +-----------+
| | +--+ | | +----- Elliptio_dariensis__3
| | | | +--+
| | | | | +-- Elliptio_dariensis__2
| | | | +--+
| | | | +-- Elliptio_mediocris
| | | |
| | | | +-- Elliptio_folliculata__2
| | | | +-----------+
| | | | | +-- Elliptio_purpurella__2
| | +--+ |
| | | | +-- Elliptio_jayensis__1
+--+ | +--+-----------+
| | | | +-- Elliptio_pullata__1
| | | |
| | | | +----------- Elliptio_arca__1
| | | +--+
| | | | +-------- Elliptio_arctata__2
| | | +--+
| | | | +----- Elliptio_fumata__1
| | | +--+
| +--+ | +-- Elliptio_arctata__1
| | +--+
| | +-- Elliptio_purpurella__1
| |
| | +-- Elliptio_congaraea__1
| | |
| | +--------+-- Elliptio__1
| | | |
| | | +-- Elliptio__2
| | |
| +-----+ +-- Elliptio_complanata__1
| | +-----+
| | | +-- Elliptio_hopetonensis__1
| | |
| +--+ +----- Elliptio_dariensis__1
| | |
| | +----- Elliptio_producta
| +--+
| | +-- Elliptio_hopetonensis__2
| | |
| +--+-- Elliptio_hopetonensis__3
| |
| +-- Elliptio_icterina__1
|
| +----- Elliptio_shepardiana
| +-----------------------------------------+
| | | +-- Elliptio_angustata
| | +--+
| | +-- Elliptio_folliculata__1
| |
+--+ +-------------------------------------------- Eurynia
| |
| | +-- Hemistena
| | +--------------------------------------+
| | | +-- Plethobasus_cyphyus
+--+ |
| | +-------------------------------------- Elliptio_fraterna
| | |
| | | +-------------------------------- Pleurobema_plenum
| | | |
+--+ | +--+ +----------------------------- Pleurobema_stabile
| | | | |
| | | | | +-------------------------- Pleuronaia
| | | +--+ |
| | | | | +-- Pleurobema_decisum
| | | | | +-----+
| | | | | | +-- Pleurobema_hanleyianum
| | | +--+ +--------------+
+--+ | | | | +----- Pleurobema_pyriforme
| | | | +--+
| | | | | +-- Pleurobema_athearni
| | | | +--+
| | | | +-- Pleurobema_georgianum
| | +--+
| | | +-- Pleurobema_clava__1
| | | +--+
| | | | +-- Pleurobema_oviforme__1
| | | +--------------+
| | | | | +-- Pleurobema_clava__2
| | | | +--+
| | +--+ +-- Pleurobema_oviforme__2
+--+ |
| | +----------------- Pleurobema_beadleianum__2
| +--+
| | +-------------- Pleurobema_rubellum
| +--+
| | +----------- Pleurobema_strodeanum__2
| +--+
| | +-------- Pleurobema_beadleianum__1
| +--+
| | +----- Pleurobema_taitianum
| +--+
| | +-- Pleurobema_strodeanum__1
| +--+
| +-- Pleurobema_perovatum
|
| +----- Pleurobema_riddellii
| +-----------------+
| | | +-- Plethobasus_cooperianus
| | +--+
| | +-- Pleurobema_sintoxia
| |
+-----------+ +-------- Pleurobema_cordatum
| +-----------+
| | | +----- Fusconaia_subrotunda
| | +--+
| | | +-- Fusconaia_cor
| | +--+
+--+ +-- Fusconaia_cuneolus
|
| +----------------- Fusconaia_ozarkensis
| |
| | +----- Fusconaia_escambia__2
| | +--------+
+--+ | | +-- Fusconaia_burkei
| | +--+
| | +-- Fusconaia_escambia__1
| |
+--+ +----- Fusconaia_masoni
| |
| +-----+ +-- Fusconaia_cerina__1
| | | |
| | +--+-- Fusconaia_cerina__2
| | |
+--+ +-- Fusconaia_flava__2
|
| +-- Fusconaia_askewi
| +-----+
| | +-- Fusconaia_chunii
+--+
| +----- Fusconaia_flava__1
+--+
| +-- Fusconaia_iheringi
+--+
+-- Fusconaia_mitchelli
Smith et al., 2020 [Publ 4129], fig. 2-3
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 10 | 18 | 10 | 7 | 683 |
+----------------------------- Obliquaria | --+ +-------------------------- Cyrtonaias | | | | +-- Pachynaias +--+ +--------------------+ | | +-- Lampsilis | | +--+ +-- Ellipsaria | +-----------------+ | | +-- Truncilla | | +--+ +----------------- Atlanticoncha | | | | +-- Potamilus_capax ** +--+ +-----------+ | | +-- Potamilus_inflatus ** | | | | +----- Potamilus_ohiensis ** +--+ +-----+ | | | +-- Potamilus_streckersoni ** | | +--+ | | +-- Potamilus_amphichaenus ** +--+ | +-- Potamilus_fragilis ** | +-----+ | | +-- Potamilus_leptodon ** +--+ | +----- Potamilus_metnecktayi ** +--+ | +-- Potamilus_alatus ** +--+ +-- Potamilus_purpuratus **
Smith et al., 2020 [Publ 4129], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 8 | 10 | 18 | 10 | 7 | 683 |
+-------------------------- Obliquaria | --+ +----------------------- Cyrtonaias | | | | +-- Pachynaias +--+ +-----------------+ | | +-- Lampsilis | | | | +-- Ellipsaria +--+ +--------------+ | | +-- Truncilla | | | | +-------------- Atlanticoncha +--+ | | | +-- Potamilus_fragilis ** | | +-----+ | | | +-- Potamilus_leptodon ** +--+ +--+ | | | +----- Potamilus_metnecktayi ** | | +--+ | | | +-- Potamilus_alatus ** | | +--+ +--+ +-- Potamilus_purpuratus ** | | +-- Potamilus_capax ** | +-----+ | | +-- Potamilus_inflatus ** +--+ | +----- Potamilus_ohiensis ** +--+ | +-- Potamilus_streckersoni ** +--+ +-- Potamilus_amphichaenus **
Smith et al., 2019 [Publ 3951], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 10 | 10 | 9 | 11 | 31 | 26 | 19 | 683 |
+----------------------------- Obliquaria | | +-- Medionidus | +--+ | | +-- Lemiox | +--------------------+ | | | +-- Cyprogenia --+ | +--+ | | +-- Ptychobranchus | | | | +-- Hamiota | | +-----------+ | | | +-- Villosa | | | +--+ +--------+ +----------- Ligumia | | | | | | | | +----- Venustaconcha | | +--+ +--+ | | | | | +-- Obovaria | | | | +--+ | | +--+ +-- Epioblasma | | | | | | +----- Ortmanniana_pectorosa +--+ +--+ | | +-- Ortmanniana_ligamentina | +--+ | +-- Lampsilis | | +----- Atlanticoncha | +--------------+ | | | +-- Ellipsaria | | +--+ | | +-- Truncilla +--+ | +----------------- Potamilus_capax ** | | | | +----- Potamilus_inflatus ** +--+ +--------+ | | | +-- Potamilus_fragilis ** | | +--+ | | +-- Potamilus_leptodon ** +--+ | +----- Potamilus_ohiensis ** | +-----+ | | | +-- Potamilus_amphichaenus ** | | +--+ +--+ +-- Potamilus_streckersoni ** | | +-------- Potamilus_metnecktayi ** +--+ | +----- Potamilus_purpuratus__1 ** +--+ | +-- Potamilus_purpuratus__2 ** +--+ +-- Potamilus_alatus **
Roe & Lydeard, 1998 [Publ 2693], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 8 | 4 | 8 | 3 | 2 | 2 | 685 |
+----------- Potamilus_capax ** | +----------- Lampsilis --+ | +-------- Potamilus_fragilis ** | | | | +----- Potamilus_inflatus ** +--+ | | | +-- Potamilus_ohiensis ** | +--+ +--+ +-- Potamilus_amphichaenus ** | | +-- Potamilus_purpuratus__1 ** | | +--+-- Potamilus_purpuratus__2 ** | +-- Potamilus_alatus **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 7 | 49 | 41 | 22 | 686 |
+-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
| +-------------------------------- Paetulunio
--+ |
| | +-- Obliquaria
| | +--------------------------+
| | | +-- Toxolasma
+--+ |
| | +-------------------------- Sagittunio
| | |
| | | +-- Medionidus_conradicus
| | | +-----+
+--+ | | +-- Ptychobranchus
| | +--------+
| | | | +----- Medionidus_acutissimus
| | | | |
| | | +--+----- Lemiox
| | | |
| | +-----+ | +-- Dromus
+--+ | | +--+
| | | +-- Cyprogenia
| | |
| | | +-------------- Potamilus_leptodon **
| | | |
| | +--+ +----------- Potamilus_fragilis **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +-----+
| | | | +-- Potamilus_purpuratus **
| | +--+
+--+ | +----- Potamilus_inflatus **
| +--+
| | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_amphichaenus **
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 7 | 7 | 5 | 7 | 50 | 41 | 22 | 686 |
+-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
| | | +-- Glebula
--+ | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus
| | +--+
| | | | +-- Dromus
| | | +--+
+--+ +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
| | +-- Lemiox
| |
| | +----------------------------- Sagittunio
+--+ |
| | +-------------- Potamilus_leptodon **
| | |
| +--------------+ +----------- Potamilus_fragilis **
| | | |
| | +--+ +-- Potamilus_alatus **
| | | +-----+
| | | | +-- Potamilus_purpuratus **
| | +--+
| | | +----- Potamilus_inflatus **
+--+ +--+
| | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_amphichaenus **
|
| +-- Truncilla
| +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Inoue et al., 2020 [Publ 3941], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 5 | 36 | 21 | 6 | 686 |
+----------------------------------- Toxolasma | --+ +-------------------------------- Ellipsaria | | | | +----------------------------- Truncilla +--+ | | | +-------- Potamilus_fragilis ** | | | | | +-----------------+ +-- Potamilus_ohiensis ** +--+ | | +--+ | | | | +-- Potamilus_amphichaenus ** | | +--+ | | | +-- Potamilus_metnecktayi ** | | +--+ | | +-- Potamilus_purpuratus ** +--+ | +----------------------- Leaunio | | | | +----------------- Lampsilis_bracteata | | | | | +--+ +-- Lampsilis_ornata | | | | +-----------+ +--+ | | | +-- Lampsilis_fasciola | | +--+ | | | +----------- Lampsilis_ovata | | +--+ | | | +-------- Lampsilis_cardium__3 | | +--+ +--+ | +----- Lampsilis_cardium__2 | +--+ | | +-- Lampsilis_satura | +--+ | +-- Lampsilis_cardium__1 | | +-------------- Lampsilis_teres | | | | +-- Lampsilis_higginsii +-----+ +--------+ | | +-- Ortmanniana | | +--+ +-- Lampsilis_radiata | +-----+ | | +-- Lampsilis_siliquoidea | | +--+ +-- Lampsilis_hydiana | +--+ | | +-- Lampsilis_bergmanni +--+ | +-- Lampsilis_virescens +--+ +-- Lampsilis_straminea
Inoue et al., 2020 [Publ 3941], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 5 | 5 | 3 | 5 | 37 | 22 | 6 | 686 |
+----------------------------------- Toxolasma | | +-------- Potamilus_fragilis ** | | | +-----------------------+ +-- Potamilus_ohiensis ** --+ | | +--+ | | | | +-- Potamilus_amphichaenus ** | | +--+ | | | +-- Potamilus_metnecktayi ** | | +--+ +--+ +-- Potamilus_purpuratus ** | | +-- Ellipsaria | +--------------------------+ | | +-- Truncilla | | | | +-------------------------- Leaunio +--+ | | | +-------------------- Lampsilis_bracteata | | | | | +--+ +----------------- Lampsilis_fasciola | | | | | +--+ | +--+ +-------------- Lampsilis_ornata | | | | | | +--+ +-- Lampsilis_ovata | | | +--------+ | | | | +-- Lampsilis_cariosa | | +--+ | | | +-------- Lampsilis_cardium__3 +--+ +--+ | | +----- Lampsilis_cardium__2 | +--+ | | +-- Lampsilis_satura | +--+ | +-- Lampsilis_cardium__1 | | +-- Lampsilis_higginsii | +--------------+ | | +-- Ortmanniana +-----+ | +-------------- Lampsilis_teres | | +--+ +-- Lampsilis_radiata | +--------+ | | +-- Lampsilis_siliquoidea +--+ | +-------- Lampsilis_bergmanni +--+ | +----- Lampsilis_hydiana +--+ | +-- Lampsilis_virescens +--+ +-- Lampsilis_straminea
Campbell et al., 2005 [Publ 2660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 4 | 98 | 39 | 22 | 685 |
+----- Obliquaria
+--------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
| +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
--+ +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
| | +-------- Ortmanniana_pectorosa
| +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_cardium
+--+ +--+
| +-- Lampsilis_ovata
|
| +-- Potamilus_fragilis **
| +--+
| | +-- Potamilus_leptodon **
+-----+
| | +-- Potamilus_alatus **
| +--+
| +-- Potamilus_purpuratus **
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Campbell et al., 2005 [Publ 2660], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 4 | 4 | 2 | 4 | 98 | 39 | 22 | 685 |
+-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-- Potamilus_fragilis **
| +--+
| | +-- Potamilus_leptodon **
| +--------+
| | | +-- Potamilus_alatus **
--+ | +--+
| | +-- Potamilus_purpuratus **
| |
| +-----------+ +-------- Medionidus_acutissimus
| | | +--+
| | | | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
+--+ | | +-- Medionidus_conradicus
| +-----+
| | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 93 | 8 | 4 | 688 |
+----------------- Toxolasma | | +----- Potamilus_leptodon ** --+ +--------+ | | | +-- Potamilus_alatus ** | | +--+ | | +-- Potamilus_streckersoni ** +--+ | +----------- Leaunio | | | | +----- Lampsilis_ornata +--+ +--+ | | | +-- Lampsilis_cardium | | +--+ +--+ +-- Lampsilis_cariosa | | +----- Venustaconcha +--+ | +-- Lampsilis_powellii +--+ +-- Lampsilis_siliquoidea
Boyer et al., 2011 [Publ 2718], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 35 | 13 | 9 | 688 |
+----- Potamilus_fragilis **
+-----------+
| | +-- Potamilus_ohiensis **
| +--+
| +-- Potamilus_alatus **
--+
| +-- Ellipsaria
| +-----------+
| | +-- Truncilla
| |
+--+ +----------- Lampsilis_cardium
| |
| | +----- Ligumia
| +-----+
+--+ | +-- Obovaria
| +--+
| +-- Epioblasma
|
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis_higginsii
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 93 | 8 | 4 | 688 |
+----------------- Toxolasma | | +----- Potamilus_leptodon ** --+ +--------+ | | | +-- Potamilus_alatus ** | | +--+ | | +-- Potamilus_streckersoni ** +--+ | +----------- Leaunio | | | | +----- Lampsilis_ornata +--+ +--+ | | | +-- Lampsilis_cardium | | +--+ +--+ +-- Lampsilis_cariosa | | +----- Venustaconcha +--+ | +-- Lampsilis_powellii +--+ +-- Lampsilis_siliquoidea
Smith & Johnson, 2020 [Publ 4229], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 0 | 0 | 0 | 686 |
+----- Potamilus_inflatus ** --+ | +-- Potamilus_purpuratus ** +--+ +-- Potamilus_fragilis **
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 89 | 8 | 4 | 688 |
+----------------- Toxolasma | | +----- Potamilus_leptodon ** --+ +--------+ | | | +-- Potamilus_alatus ** | | +--+ | | +-- Potamilus_streckersoni ** +--+ | +----------- Leaunio | | | | +----- Lampsilis_ornata +--+ +--+ | | | +-- Lampsilis_cardium | | +--+ +--+ +-- Lampsilis_cariosa | | +----- Venustaconcha +--+ | +-- Lampsilis_powellii +--+ +-- Lampsilis_siliquoidea
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 12 | 6 | 3 | 688 |
+----- Potamilus_leptodon **
+-----+
| | +-- Potamilus_streckersoni **
| +--+
| +-- Potamilus_alatus **
--+
| +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Boyer et al., 2011 [Publ 2718], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 32 | 13 | 9 | 688 |
+-------------- Toxolasma | | +----------- Obliquaria | | --+ | +-------- Obovaria | | | | | +-------- Lampsilis_cardium | | | +--+ | +-- Epioblasma | +-----+ | | +-- Lampsilis_siliquoidea | | | | +-- Ortmanniana | +-----+ +--+ +-- Lampsilis_higginsii | | +-- Ligumia +-----+ | +-- Lampsilis_teres | | +-- Ellipsaria +-----+ | +-- Truncilla | | +----- Potamilus_fragilis ** +--+ | +-- Potamilus_ohiensis ** +--+ +-- Potamilus_alatus **
Krebs et al., 2003 [Publ 2691], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 9 | 7 | 3 | 30 | 8 | 7 | 693 |
+----- Ptychobranchus
+--------+
| | +-- Potamilus_fragilis **
| +--+
| +-- Medionidus
--+
| +-- Villosa
| +--------+
| | +-- Ellipsaria
+--+
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Potamilus_purpuratus **
+--+
+-- Potamilus_alatus **
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 89 | 8 | 4 | 688 |
+----------------- Toxolasma | | +----- Potamilus_leptodon ** --+ +--------+ | | | +-- Potamilus_alatus ** | | +--+ +--+ +-- Potamilus_streckersoni ** | | +----------- Leaunio | | +--+ +-------- Venustaconcha | | | | +-- Lampsilis_powellii +--+-----+ | +-- Lampsilis_siliquoidea | | +----- Lampsilis_ornata +--+ | +-- Lampsilis_cardium +--+ +-- Lampsilis_cariosa
Dai et al., 2025 [Publ 4440], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 85 | 7 | 4 | 688 |
+----------------- Toxolasma | | +----- Potamilus_leptodon ** --+ +--------+ | | | +-- Potamilus_streckersoni ** | | +--+ +--+ +-- Potamilus_alatus ** | | +----------- Leaunio | | +--+ +-- Lampsilis_cardium | +-----+ | | +-- Lampsilis_ornata +--+ | +----- Venustaconcha +--+ | +-- Lampsilis_powellii +--+ +-- Lampsilis_siliquoidea
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5
| tT | bi | riS | oT | roT | roS | iD | |
| 3 | 3 | 1 | 3 | 138 | 9 | 5 | 688 |
+-------------------- Toxolasma | | +----- Potamilus_leptodon ** --+ +-----------+ | | | +-- Potamilus_alatus ** | | +--+ | | +-- Potamilus_streckersoni ** +--+ | +-------------- Leaunio | | | | +----- Ortmanniana +--+ +-----+ | | | +-- Lampsilis_hydiana | | +--+ +--+ +-- Lampsilis_siliquoidea | | +-------- Venustaconcha +--+ | +----- Lampsilis_ornata +--+ | +-- Lampsilis_cardium +--+ +-- Lampsilis_cariosa
Wu et al., 2021 [Publ 4366], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 67 | 6 | 3 | -- |
+-------------- Toxolasma | | +-- Potamilus_alatus ** --+ +--------+ | | +-- Potamilus_leptodon ** | | +--+ +-- Lampsilis_ornata | +-----+ | | +-- Lampsilis_cardium +--+ | +----- Venustaconcha +--+ | +-- Lampsilis_siliquoidea +--+ +-- Lampsilis_powellii
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 15 | 0 | 2 | 71 | 13 | 10 | 692 |
+-------------- Potamilus_alatus ** | +-------------- Potamilus_capax ** | +-------------- Glebula | | +-- Toxolasma +-----------+ --+ +-- Paetulunio | | +-- Dromus +-----------+ | +-- Lemiox | | +-- Lampsilis_ovata | +--------+ | | +-- Obliquaria +--+ | +-------- Lampsilis_virescens | | +--+ +-- Epioblasma | +--+ | | +-- Obovaria_arkansasensis +--+ | +-- Obovaria_olivaria +--+ +-- Venustaconcha
Froufe et al., 2019 [Publ 3936], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 42 | 3 | 3 | -- |
+-------- Toxolasma | --+ +-- Lampsilis | +--+ | | +-- Venustaconcha +--+ | +-- Potamilus_leptodon ** +--+ +-- Potamilus_alatus **
Wen et al., 2017 [Publ 3795], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 14 | 3 | 3 | -- |
+-------- Toxolasma | --+ +-- Lampsilis | +--+ | | +-- Venustaconcha +--+ | +-- Potamilus_leptodon ** +--+ +-- Potamilus_alatus **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 12 | 0 | 2 | 103 | 9 | 7 | 687 |
+-- Dromus | +-- Epioblasma | +-- Glebula | +-- Lampsilis_ovata | +-- Lampsilis_virescens | +-- Lemiox --+ +-- Obliquaria | +-- Obovaria_olivaria | +-- Obovaria_unicolor | +-- Plectomerus | +-- Potamilus_alatus ** | +-- Potamilus_capax **
Froufe et al., 2019 [Publ 3936], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 42 | 3 | 3 | -- |
+-------- Toxolasma | --+ +-- Potamilus_leptodon ** | +--+ | | +-- Potamilus_alatus ** +--+ | +-- Lampsilis +--+ +-- Venustaconcha
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 17 | 6 | 3 | -- |
+-------------- Toxolasma | | +-- Potamilus_alatus ** --+ +--------+ | | +-- Potamilus_leptodon ** | | +--+ +-- Lampsilis_cardium | +-----+ | | +-- Lampsilis_ornata +--+ | +----- Venustaconcha +--+ | +-- Lampsilis_powellii +--+ +-- Lampsilis_siliquoidea
Wu et al., 2021 [Publ 4366], fig. 3-ML
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 72 | 7 | 4 | -- |
+----------------- Toxolasma | --+ +-- Potamilus_alatus ** | +-----------+ | | +-- Potamilus_leptodon ** +--+ | +----------- Leaunio | | +--+ +-- Lampsilis_ornata | +-----+ | | +-- Lampsilis_cardium +--+ | +----- Venustaconcha +--+ | +-- Lampsilis_siliquoidea +--+ +-- Lampsilis_powellii
Huang et al., 2019 [Publ 3843], fig. S3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 12 | 0 | 2 | 196 | 12 | 11 | 692 |
+-------------------- Potamilus_leptodon ** | --+ +----------------- Potamilus_alatus ** | | | | +-- Medionidus +--+ +-----------+ | | +-- Ptychobranchus | | +--+ +-- Ellipsaria | +--------+ | | +-- Truncilla +--+ | +-------- Leaunio | | +--+ +-- Venustaconcha | +--+ | | +-- Cambarunio +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Wang et al., 2018 [Publ 3865], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 29 | 3 | 3 | -- |
+-------- Toxolasma | --+ +-- Potamilus_leptodon ** | +--+ | | +-- Potamilus_alatus ** +--+ | +-- Venustaconcha +--+ +-- Lampsilis
Wu et al., 2019 [Publ 3864], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 44 | 3 | 3 | -- |
+-------- Toxolasma | --+ +-- Lampsilis | +--+ | | +-- Venustaconcha +--+ | +-- Potamilus_leptodon ** +--+ +-- Potamilus_alatus **
Dai et al., 2023 [Publ 4285], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 51 | 7 | 4 | -- |
+----------------- Toxolasma | --+ +-- Potamilus_leptodon ** | +-----------+ | | +-- Potamilus_alatus ** +--+ | +----------- Leaunio | | +--+ +-- Lampsilis_cardium | +-----+ | | +-- Lampsilis_ornata +--+ | +----- Venustaconcha +--+ | +-- Lampsilis_powellii +--+ +-- Lampsilis_siliquoidea
Wu et al., 2022 [Publ 4238], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 72 | 7 | 4 | -- |
+----------------- Toxolasma | --+ +-- Potamilus_alatus ** | +-----------+ | | +-- Potamilus_leptodon ** +--+ | +----------- Leaunio | | +--+ +-- Lampsilis_cardium | +-----+ | | +-- Lampsilis_ornata +--+ | +----- Venustaconcha +--+ | +-- Lampsilis_siliquoidea +--+ +-- Lampsilis_powellii
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 71 | 13 | 10 | -- |
+----------------- Obliquaria | | +-- Dromus | +--+ --+ | +-- Lemiox | +--------+ | | | +-- Potamilus_alatus ** | | +--+ +--+ +-- Potamilus_capax ** | | +----------- Lampsilis_ovata | | +--+ +-------- Lampsilis_virescens | | +--+ +-- Epioblasma | +--+ | | +-- Obovaria_arkansasensis +--+ | +-- Obovaria_olivaria +--+ +-- Venustaconcha
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 4 | 3 | 2 | -- |
+-- Potamilus_streckersoni **
+--+
| +-- Potamilus_alatus **
--+
| +-- Venustaconcha
+--+
+-- Lampsilis
Huang et al., 2019 [Publ 3843], fig. S4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 12 | 0 | 2 | 198 | 12 | 11 | 688 |
+----- Ellipsaria | +----- Potamilus_leptodon ** | +----- Medionidus | +----- Potamilus_alatus ** | +----- Ptychobranchus --+ +----- Truncilla | +----- Venustaconcha | +----- Cambarunio | +----- Leaunio | | +-- Ortmanniana +--+ +-- Lampsilis
Wu et al., 2020 [Publ 4121], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 47 | 3 | 3 | -- |
+-------- Toxolasma | --+ +-- Venustaconcha | +--+ | | +-- Lampsilis +--+ | +-- Potamilus_alatus ** +--+ +-- Potamilus_leptodon **
Huang et al., 2002 [Publ 2657], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 6 | 0 | 2 | 32 | 6 | 6 | 692 |
+----------------- Medionidus --+ | +-------------- Potamilus_fragilis ** +--+ | +----------- Ptychobranchus +--+ | +-------- Potamilus_purpuratus ** +--+ | +----- Lampsilis +--+ | +-- Villosa +--+ +-- Ellipsaria
Huang et al., 2019 [Publ 3843], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 54 | 5 | 4 | -- |
+----------- Toxolasma | --+ +-- Potamilus_leptodon ** | +-----+ | | +-- Potamilus_alatus ** +--+ | +----- Leaunio +--+ | +-- Venustaconcha +--+ +-- Lampsilis
Huang et al., 2002 [Publ 2657], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 10 | 0 | 2 | 32 | 6 | 6 | 688 |
+----- Potamilus_fragilis ** | +----- Medionidus | +----- Amblema | | +-- Obliquaria --+--+ | +-- Plectomerus | | +-- Villosa | | | +-- Ellipsaria | | +--+-- Lampsilis | +-- Potamilus_purpuratus ** | +-- Ptychobranchus
Dai et al., 2024 [Publ 4358], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 51 | 4 | 4 | -- |
+----------- Toxolasma | --+ +-- Potamilus_leptodon ** | +-----+ | | +-- Potamilus_alatus ** +--+ | +----- Leaunio +--+ | +-- Lampsilis +--+ +-- Venustaconcha
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 52 | 6 | 3 | -- |
+-- Potamilus_leptodon **
+-----------+
| +-- Potamilus_alatus **
--+
| +----------- Leaunio
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Burlakova et al., 2019 [Publ 3899], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 31 | 24 | 17 | -- |
+----------------- Toxolasma | | +-- Potamilus_fragilis ** | +-----------+ | | +-- Potamilus_metnecktayi ** | | --+ | +-- Ellipsaria | +-----------+ | | +-- Truncilla | | | | +----- Medionidus | +--------+ | | | +-- Lemiox +--+ +--+ | +-- Dromus | | +----------- Cambarunio | | | +----------- Leaunio | | | +----------- Villosa | | | | +-- Sagittunio | +--------+ +--+ +-- Lampsilis_ovata | | +----- Venustaconcha +-----+ | | +-- Epioblasma | +--+ | +-- Obovaria | | +-------- Lampsilis_teres +--+ | +----- Lampsilis_siliquoidea +--+ | +-- Lampsilis_hydiana +--+ +-- Lampsilis_straminea
Lydeard et al., 1996 [Publ 2656], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 6 | 0 | 2 | 26 | 7 | 6 | 685 |
+-------------------- Obliquaria --+ | +----------------- Medionidus +--+ | +-------------- Ptychobranchus +--+ | +----------- Potamilus_fragilis ** +--+ | +-------- Potamilus_purpuratus ** +--+ | +----- Ellipsaria +--+ | +-- Villosa +--+ +-- Lampsilis
Cha et al., 2018 [Publ 3916], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 36 | 2 | 2 | -- |
+-------- Toxolasma --+ | +----- Lampsilis +--+ | +-- Potamilus_alatus ** +--+ +-- Potamilus_leptodon **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 46 | 3 | 3 | -- |
+-------- Toxolasma | --+ +-- Potamilus_alatus ** | +--+ | | +-- Potamilus_leptodon ** +--+ | +-- Venustaconcha +--+ +-- Lampsilis
Wang et al., 2018 [Publ 3846], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 29 | 3 | 3 | -- |
+-------- Toxolasma | --+ +-- Venustaconcha | +--+ | | +-- Lampsilis +--+ | +-- Potamilus_leptodon ** +--+ +-- Potamilus_alatus **
Huang et al., 2018 [Publ 3828], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 40 | 3 | 3 | -- |
+-------- Toxolasma | --+ +-- Lampsilis | +--+ | | +-- Venustaconcha +--+ | +-- Potamilus_leptodon ** +--+ +-- Potamilus_alatus **
Wu et al., 2021 [Publ 4366], fig. 3-BI
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 72 | 7 | 4 | -- |
+----------------- Toxolasma | --+ +-- Potamilus_alatus ** | +-----------+ | | +-- Potamilus_leptodon ** +--+ | +----------- Leaunio | | +--+ +-- Lampsilis_ornata | +-----+ | | +-- Lampsilis_cardium +--+ | +----- Venustaconcha +--+ | +-- Lampsilis_siliquoidea +--+ +-- Lampsilis_powellii
Lydeard et al., 1996 [Publ 2656], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 8 | 0 | 2 | 25 | 7 | 6 | 691 |
+-------------- Potamilus_fragilis ** | --+ +----------- Ptychobranchus | | +--+ +-------- Medionidus | | +--+ +----- Potamilus_purpuratus ** | | +--+----- Lampsilis | | +-- Ellipsaria +--+ +-- Villosa
Chen et al., 2020 [Publ 4170], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 16 | 0 | 2 | 32 | 4 | 2 | 691 |
+-- Utterbackia
+--------+
| +-- Potamilus_alatus **
+--------+
| | +-------- Lamprotula
| +--+
| | +----- Sinohyriopsis
| +--+
--+ | +-- Parvasolenaia
| +--+
| +-- Sinosolenaia
|
| +----------------- Gibbosula
+--+
| +-------------- Quadrula
+--+
| +----------- Toxolasma
+--+
| +-------- Lampsilis_ornata
+--+
| +----- Potamilus_leptodon **
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Campbell & Lydeard, 2026 [Publ 4810], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 45 | 4 | 3 | -- |
+-- Potamilus_alatus **
+-----+
| +-- Potamilus_streckersoni **
--+
| +----- Venustaconcha
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Huang et al., 2019 [Publ 3843], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 12 | 0 | 2 | 198 | 12 | 11 | 689 |
+-------------- Potamilus_leptodon ** | | +----------- Potamilus_alatus ** --+ | | | +-------- Venustaconcha | | | +--+ +-------- Leaunio | | | | +-- Ellipsaria | +-----+ +--+ +-- Truncilla | | +-- Medionidus +-----+ | +-- Ptychobranchus | | +----- Cambarunio +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Chen et al., 2025 [Publ 4448], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 2 | 2 | 0 | 2 | 84 | 7 | 4 | -- |
+----------------- Toxolasma | --+ +-- Potamilus_leptodon ** | +-----------+ | | +-- Potamilus_alatus ** +--+ | +----------- Leaunio | | +--+ +-- Lampsilis_cardium | +-----+ | | +-- Lampsilis_ornata +--+ | +----- Venustaconcha +--+ | +-- Lampsilis_powellii +--+ +-- Lampsilis_siliquoidea
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 11 | 11 | -- |
+-- Cyrtonaias
+--------------------+
| +-- Glebula
--+
| +-------------------- Potamilus_alatus **
| |
+--+ +-- Truncilla
| +--------------+
| | +-- Ptychobranchus
+--+
| +-------------- Pachynaias
| |
+--+ +----------- Mesonaias
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Ortmanniana
+--+
| +----- Cambarunio
+--+
| +-- Epioblasma
+--+
+-- Ligumia
Huang et al., 2019 [Publ 3843], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- |
+-- Venustaconcha --+ +-- Potamilus_alatus **
Walker et al., 2006 [Publ 2666], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 14 | 4 | 4 | -- |
+----------- Potamilus_purpuratus ** --+ | +-------- Mesonaias +--+ | +----- Ligumia +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Feng et al., 2016 [Publ 3800], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 13 | 1 | 1 | -- |
+-- Potamilus_leptodon ** --+ +-- Lampsilis
Hoeh et al., 2001 [Publ 2652], fig. 14f2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- |
+----------- Ortmanniana --+ | +-------- Glebula +--+ | +----- Potamilus_alatus ** +--+ | +-- Ligumia +--+ +-- Toxolasma
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 33 | 5 | 5 | -- |
+----- Potamilus_alatus ** --+ | +-- Ortmanniana +--+ +-- Ligumia
Walker et al., 2006 [Publ 2666], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 35 | 4 | 4 | -- |
+-------- Potamilus_purpuratus ** --+ | +----- Lampsilis +--+ | +-- Ligumia +--+ +-- Ortmanniana
Hoeh et al., 2001 [Publ 2652], fig. 14f4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- |
+-- Cyrtonaias
+--------+
| +-- Glebula
--+
| +-------- Toxolasma
+--+
| +----- Potamilus_alatus **
+--+
| +-- Ligumia
+--+
+-- Ortmanniana
Hoeh et al., 2002 [Publ 2673], fig. 1-F
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 8 | 4 | 4 | -- |
+----------- Cyrtonaias --+ | +-------- Ortmanniana +--+ | +----- Lampsilis +--+ | +-- Potamilus_purpuratus ** +--+ +-- Ligumia
Curole & Kocher, 2005 [Publ 2682], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 10 | 4 | 3 | -- |
+-------- Leaunio --+ | +----- Lampsilis_straminea +--+ | +-- Lampsilis_teres +--+ +-- Potamilus_purpuratus **
Inoue et al., 2020 [Publ 4128], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 21 | 10 | 10 | -- |
+----------------------- Cyrtonaias | --+ +-------------------- Ptychobranchus | | +--+ +----------------- Ligumia | | +--+ +-------------- Potamilus_alatus ** | | +--+ +----------- Pachynaias | | | | +-- Epioblasma +--+ +-----+ | | +-- Truncilla | | +--+ +-- Mesonaias | +--+ | | +-- Cambarunio +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 39 | 2 | 2 | -- |
+----- Potamilus_alatus ** --+ | +-- Lampsilis +--+ +-- Truncilla
Hoeh et al., 2002 [Publ 2653], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- |
+----- Potamilus_alatus ** --+ | +-- Ligumia +--+ +-- Ortmanniana
Szumowski et al., 2012 [Publ 3569], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 15 | 14 | 9 | -- |
+----------------------- Ellipsaria | --+ +-------------------- Truncilla | | +--+ +----------------- Obliquaria | | | | +-------------- Potamilus_alatus ** +--+ | | | +----- Lampsilis_ornata | | +-----+ | | | | +-- Lampsilis_cardium +--+ | +--+ | | +-- Lampsilis_ovata | | | | +-------- Lampsilis_teres | +--+ +--+ | +----- Lampsilis_siliquoidea | +--+ | | +-- Ortmanniana | +--+ | +-- Lampsilis_higginsii | | +----- Hamiota | | +-----+ +-- Epioblasma | | +--+-- Cambarunio | +-- Ligumia
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- |
+-- Potamilus_alatus ** --+ +-- Venustaconcha
Guerra et al., 2019 [Publ 4168], fig. 2-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- |
+-- Venustaconcha --+ +-- Potamilus_alatus **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 27 | 1 | 1 | -- |
+-- Lampsilis --+ +-- Potamilus_leptodon **
Klymus et al., 2025 [Publ 4814], fig. 3B
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 35 | 4 | 3 | -- |
+-------- Potamilus_alatus ** --+ | +----- Venustaconcha +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Grobler et al., 2011 [Publ 3359], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 16 | 15 | 9 | -- |
+-------- Lemiox
+-----+
| | +----- Medionidus
| +--+
| | +-- Dromus
| +--+
--+ +-- Cyprogenia
|
| +----------- Potamilus_alatus **
| |
| | +----- Lampsilis_siliquoidea
| | +--+
+--+ | | +-- Lampsilis_higginsii
| | +--+
| | +-- Ortmanniana
| |
| | +----- Lampsilis_fasciola
+--+--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-- Cambarunio
| |
+-----+-- Ligumia
|
+-- Epioblasma
Walker et al., 2006 [Publ 2666], fig. 6-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 13 | 4 | 4 | -- |
+----------- Potamilus_purpuratus ** --+ | +-------- Cyrtonaias +--+ | +----- Ligumia +--+ | +-- Lampsilis +--+ +-- Ortmanniana
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 38 | 5 | 3 | -- |
+----------- Toxolasma | --+ +-------- Potamilus_leptodon ** | | +--+ +-- Lampsilis_cardium | +--+ | | +-- Lampsilis_ornata +--+ | +-- Lampsilis_siliquoidea +--+ +-- Venustaconcha
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- |
+-- Cyrtonaias
+--------+
| +-- Glebula
--+
| +-------- Toxolasma
+--+
| +----- Ortmanniana
+--+
| +-- Ligumia
+--+
+-- Potamilus_alatus **
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- |
+-- Venustaconcha --+ +-- Potamilus_alatus **
Zhou et al., 2017 [Publ 3788], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 15 | 1 | 1 | -- |
+-- Potamilus_leptodon ** --+ +-- Lampsilis
Serb et al., 2003 [Publ 2659], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 28 | 6 | 5 | -- |
-- Potamilus_alatus **
Smith, 2021 [Publ 4331], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 19 | 3 | 2 | -- |
+----- Potamilus_streckersoni ** --+ | +-- Venustaconcha +--+ +-- Lampsilis
Wen et al., 2017 [Publ 3795], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- |
+-- Potamilus_alatus ** --+ +-- Venustaconcha
Guerra et al., 2019 [Publ 4168], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- |
+-- Venustaconcha --+ +-- Potamilus_alatus **
Wu et al., 2021 [Publ 4366], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 90 | 3 | 3 | -- |
+-------- Obliquaria --+ | +----- Potamilus_alatus ** +--+ | +-- Lampsilis +--+ +-- Truncilla
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 30 | 5 | 5 | -- |
+----------- Ortmanniana --+ | +-------- Glebula +--+ | +----- Potamilus_alatus ** +--+ | +-- Ligumia +--+ +-- Toxolasma
Hoeh et al., 2002 [Publ 2673], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 8 | 4 | 4 | -- |
+----------- Potamilus_purpuratus ** --+ | +-------- Cyrtonaias +--+ | +----- Ligumia +--+ | +-- Lampsilis +--+ +-- Ortmanniana
Wu et al., 2023 [Publ 4339], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 18 | 3 | 3 | -- |
+-------- Obliquaria --+ | +----- Truncilla +--+ | +-- Lampsilis +--+ +-- Potamilus_alatus **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 25 | 1 | 1 | -- |
+-- Lampsilis --+ +-- Potamilus_leptodon **
Walker et al., 2006 [Publ 2666], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 14 | 4 | 4 | -- |
+----------- Mesonaias --+ | +-------- Potamilus_purpuratus ** +--+ | +----- Ligumia +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Froufe et al., 2019 [Publ 3936], fig. 3-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- |
+-- Potamilus_alatus ** --+ +-- Venustaconcha
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 11 | 4 | 3 | -- |
+-- Potamilus_purpuratus **
+--+
| +-- Glebula
--+
| +-- Leaunio
+--+
+-- Lampsilis
Wu et al., 2021 [Publ 4366], fig. 2_I
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 90 | 3 | 3 | -- |
+-------- Obliquaria --+ | +----- Potamilus_alatus ** +--+ | +-- Lampsilis +--+ +-- Truncilla
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 36 | 5 | 3 | -- |
+----------- Toxolasma | --+ +-------- Potamilus_leptodon ** | | +--+ +-- Lampsilis_cardium | +--+ | | +-- Lampsilis_ornata +--+ | +-- Lampsilis_siliquoidea +--+ +-- Venustaconcha
Guerra et al., 2019 [Publ 4168], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 40 | 2 | 2 | -- |
+----- Toxolasma --+ | +-- Venustaconcha +--+ +-- Potamilus_alatus **
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 28 | 1 | 1 | -- |
+-- Lampsilis --+ +-- Potamilus_leptodon **
Wu et al., 2024 [Publ 4388], fig. 6
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 52 | 4 | 4 | -- |
+----------- Toxolasma --+ | +-------- Potamilus_alatus ** +--+ | +----- Leaunio +--+ | +-- Lampsilis +--+ +-- Venustaconcha
Keogh & Simons, 2019 [Publ 3935], fig. S1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 9 | 7 | 2 | -- |
+----- Potamilus_fragilis ** --+ | +-- Leaunio +--+ +-- Lampsilis
Inoue et al., 2020 [Publ 4128], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 22 | 11 | 11 | -- |
+-- Glebula
+--------+
| +-- Cyrtonaias
|
--+ +-------- Potamilus_alatus **
| |
| | +-- Ptychobranchus
+--+ +--+
| | +-- Truncilla
| |
| | +-- Ortmanniana
+--+ |
| +-- Epioblasma
| |
| +-- Lampsilis
| |
+--+-- Ligumia
|
+-- Cambarunio
|
+-- Pachynaias
|
+-- Mesonaias
Serb et al., 2003 [Publ 2659], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 28 | 6 | 5 | -- |
-- Potamilus_alatus **
Breton et al., 2011 [Publ 3376], fig. 3
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 37 | 12 | 7 | -- |
+----------------- Toxolasma | --+ +-------------- Ellipsaria | | +--+ +----------- Truncilla | | +--+ +-- Lemiox | +-----+ | | +-- Potamilus_metnecktayi ** +--+ | +----- Lampsilis +--+ | +-- Cambarunio +--+ +-- Venustaconcha
Froufe et al., 2019 [Publ 3936], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- |
+-- Potamilus_alatus ** --+ +-- Venustaconcha
Huang et al., 2018 [Publ 3828], fig. 6-M
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 16 | 1 | 1 | -- |
+-- Venustaconcha --+ +-- Potamilus_alatus **
Walker et al., 2006 [Publ 2666], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 14 | 4 | 4 | -- |
+----------- Mesonaias --+ | +-------- Potamilus_purpuratus ** +--+ | +----- Ligumia +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Chase et al., 2018 [Publ 3847], fig. 5
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 9 | 2 | 1 | -- |
+-- Potamilus_alatus ** --+ +-- Lampsilis
Klymus et al., 2025 [Publ 4814], fig. 3A
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 35 | 4 | 3 | -- |
+-------- Potamilus_alatus ** --+ | +----- Venustaconcha +--+ | +-- Ortmanniana +--+ +-- Lampsilis
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 37 | 5 | 5 | -- |
+-- Cyrtonaias
+-----+
| +-- Glebula
|
--+ +-- Lampsilis
| +--+
| | +-- Leaunio
+--+
| +-- Truncilla
+--+
+-- Potamilus_fragilis **
Guerra et al., 2019 [Publ 4168], fig. 1
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 40 | 2 | 2 | -- |
+----- Toxolasma --+ | +-- Venustaconcha +--+ +-- Potamilus_alatus **
Chase et al., 2018 [Publ 3847], fig. 4
| tT | bi | riS | oT | roT | roS | iD | |
| 1 | 1 | 0 | 1 | 38 | 12 | 7 | -- |
+----------------- Toxolasma | --+ +-------------- Venustaconcha | | +--+ +-- Lemiox | +--------+ | | +-- Lampsilis +--+ | +-------- Potamilus_metnecktayi ** +--+ | +----- Cambarunio +--+ | +-- Ellipsaria +--+ +-- Truncilla
