species Unio crassus Philipsson in Retzius, 1788: synonymy

Mya ovalis Lightfoot, 1786

Lister, 1685. Conch.: pl. 146, fig. 1. [pre-linnaean]
‘S.’ Lightfoot, 1786. Catalogue of the Portland Museum: 134, no. 2983.

Unio crassus Philipsson in Retzius, 1788

Schröter, 1779. Gesch. Flußconch.: 182, pl. 2, fig. 2 [Mya testa crassa]. [non-binomial]
Philipsson in Retzius, 1788. Diss. Nat.-Hist. Test. Gen.: 17.

Margaritana crassa — Simpson (1900)
Unio crassus — Kobelt & Haas (1910), Ortmann (1912), Ortmann (1914), Germain (1931), R.I. Johnson (1953), ... Wu et al. (2025), Gorbunova et al. (2025), Nagel (2025), Campbell & Lydeard (2026), Prie et al. (2026)
Margaritana (Group of M. margaritifera) crassa — Simpson (1914)
Unio (s.s.) crassus — Zhadin (1938)
Unio crassus crassus — Haas (1940), Haas (1969), Fauna Europaea (2012), Prié & Puillandre (2014), Bogatov (2022)
Crassiana crassa crassa — Akromovski (1976), Kantor & Sysoev (2005), Kantor et al. (2010)
Batavusiana (Crassunio) crassa — Zatravkin (1983), Stadnichenko (1984), Zatravkin (1987)
Unio (Crassunio) crassus — Falkner (1994)
Batavusiana (Crassiana) crassus — Korniushin (1998)
Unio (Crassiana) crassus crassus — Falkner et al. (2001), Falkner et al. (2002)
Crassiana (s.s.) crassa — Starobogatov et al. (2004)
“Unio crassus” ** — Soroka (2005), Soroka (2010), Reis et al. (2013), Douda et al. (2014), Froufe et al. (2016), ... Klymus et al. (2025), Wu et al. (2026), Campbell & Lydeard (2026), Wu et al. (2026), Li et al. (2026)
Crassiana crassa — Gozhik (2006)
“Unio crassus crassus” ** — Prié & Puillandre (2014), Lopes-Lima et al. (2017), Klishko et al. (2017)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
“Unio crassus” ** — Araujo et al. (2018), Lyubas et al. (2022)
“Unio crassus” ** — Araujo et al. (2018)
Unio c. crassus — Príe et al. (2021)

Mya pictorum Gmelin, 1791

(misidentified reference to Mya pictorum Linnaeus, 1758)
Lister, 1678. Hist. Anim. Anglic.: pl. 2, fig. 30; [app.] pl. 1, fig. 4. [pre-linnaean]
Lister, 1685. Conch.: pl. 146, fig. 1; pl. 147, figs. 2-3. [pre-linnaean]
Schröter, 1779. Gesch. Flußconch.: pl. 2, fig. 2; pl. 3, figs. 2-5; pl. 4, fig. 6.
Martini & Chemnitz, 1782. Conchyl. Cab. 6: 19, pl. 1, fig. 6.
Gmelin, 1791. Syst. Nat. 1(6): 3218, sp. 3.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)

Unio musivus Spengler, 1793

Spengler, 1793. Skr. Naturhist. Selsk. 3: 67, no. 32.

Unio crassus musivus — Ortmann (1912), Bogatov (2022)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)
Unio crassus crassus — Haas (1940), Haas (1969), Fauna Europaea (2012), Klishko et al. (2017)
Batavusiana (s.s.) musiva — Zatravkin (1983), Zatravkin (1987)
Batavusiana (s.s.) musivus [??] — Korniushin (1998)
Crassiana (Bataviana) musiva — Starobogatov et al. (2004)
Crassiana musiva musiva — Kantor & Sysoev (2005), Kantor et al. (2010)
Crassiana musiva — Gozhik (2006)
Unio crassus — Graf (2007), Bogatov et al. (2018), Anisimova et al. (2018), Lopes-Lima et al. (2024)
Unio (Crassiana) musivus — Vinarski & Kantor (2016)
Unio crassus cf. courtillieri — Klishko et al. (2017)

Mya batava Maton & Rackett, 1807

Schröter, 1779. Gesch. Flußconch.: pl. 3, figs. 2, 4-5.
Martini & Chemnitz, 1782. Conchyl. Cab. 6: 19, pl. 1, fig. 6.
Bruguière, 1797. Encycl. Méth. 2: pl. 248, fig. 3.
Maton & Rackett, 1807. Trans. Linn. Soc. London 8: 37.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (Batavusiana) batavus — Germain (1904)
Unio (Groupe des Batavusiana) batavus — Caziot (1907)
Unio batavus — Kobelt & Haas (1910), Haas (1924), Germain (1931)
Unio crassus musivus — Ortmann (1912)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)
Unio crassus batavus — Haas (1940), R.I. Johnson (1953), Haas (1969), Fauna Europaea (2012), Lopes-Lima et al. (2017)
Batavusiana (s.s.) musiva — Zatravkin (1983), Zatravkin (1987)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Araujo et al. (2018)

Unio lemovicincae Férussac, 1807

‘Linn.’ Draparnaud, 1806. Hist. Moll. France: pl. 11, fig. 5.
Férussac, 1807. Ess. méth.: 114 [as ‘lemovicincæ’].

Unio litoralis Pfeiffer, 1821

(misidentified reference to Unio littoralis Cuvier, 1798; error for Unio littoralis Cuvier, 1798)
Pfeiffer, 1821. Natur. Deutsch. Land Sussw. Moll. 1: 117, pl. 5, fig. 12.
Rossmässler, 1853. Zeitschr. f. Malakozool. 10: 92.
Mørch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): ??.

Margaritana crassa — Simpson (1900)
Margaritana (Group of M. margaritifera) crassa — Simpson (1914)
Batavusiana (Crassunio) crassa — Stadnichenko (1984)
Crassiana crassa crassa — Kantor & Sysoev (2005), Kantor et al. (2010)

Unio ater Nilsson, 1822

‘Linn.’ Draparnaud, 1806. Hist. Moll. France: pl. 11, fig. 5 [as ‘Unio margaritifera’].
Nilsson, 1822. Hist. Moll. Svec.: 107.
LECTOTYPE SMF 10953 (by Zilch, 1967: 53), Höjeå River near Lund, Sweden.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)
Unio (s.s.) ater — Zhadin (1938)
Unio crassus crassus — Haas (1940), Haas (1969)
Batavusiana (s.s.) musiva — Zatravkin (1987)
Crassiana ater — Gozhik (2006)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio rubens Menke, 1828

Menke, 1828. Syn. Method. Moll.: 64, 90.
Menke, 1830. Syn. Method. Moll.: 107, 149.
LECTOTYPE SMF 10950 (by Haas, 1912: 527, pl. 5, figs. 10-14), mill race on Wupper River, Barmen, Germany.

Quadrula (sec. Fusconaia) coccinea — Simpson (1900)
Quadrula (sec. Fusconaia) (Group of Q. undata) coccinea — Simpson (1914)
Quadrula (Obliquata) coccinea — Frierson (1927)
Unio crassus crassus — Haas (1940), Haas (1969)
Pleurobema sintoxia — Parmalee & Bogan (1998), Williams et al. (2008)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio rugatus Menke, 1828

Menke, 1828. Syn. Method. Moll.: 64, 90.
Menke, 1830. Syn. Method. Moll.: 107, 149.
LECTOTYPE SMF 10951 (by Zilch, 1967: 54), Emmer River, Pyrmont, Germany.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio consentaneus var. — Kobelt & Haas (1910)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio badius Kokeil, 1831

Kokeil, 1831. Mich. Comp.: pl. 16, fig. 36.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)

Unio arcuata Bouchard-Chantereaux, 1838

(renamed Unio bouchardi Locard, 1889)
Bouchard-Chantereaux, 1838. Cat. Moll. Terr. Fluv. Départ. Pas-de-Calais: 91, pl. 1, fig. 1-2.

Margaritana auricularia — Germain (1931)
Unio consentaneus var. moquini — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus moquinianus — Prie (2012)

Unio corrugata Mauduyt, 1839

Mauduyt, 1839. Tableau Indic. Moll. Vienne: 8, pl. 1, fig. 1.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)

Unio littoralis var. minor Rossmässler, 1842

(intro'd as intra-sp. of Unio littoralis Cuvier, 1798; homonym of Unio margaritifer var. minor Rossmässler, 1835)
Rossmässler, 1842. Icon. Land- und Süssw.-Moll. 2 (5): 14, pl. 55, figs. 744. [in part]
SYNTYPE SMF 10949, Schulensee, near Kiel, Germany.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)
Unio (Tumidusiana) tumidus var. minor — Stadnichenko (1984)
Unio crassus — Nagel (2025)

Unio crassissimus Hanley, 1843

(homonym of Unio crassissimus J. de C. Sowerby, 1817)
Schlüter, 1838. Kurzgef. Syst. Verz. Conch. Samml.: 33 [as ‘Mysca tumida var. crassissima’]. [nomen nudum]
‘Fer.’ Hanley, 1843. Illustr. Descr. Cat. Rec. Bivalve Shells: 209.

Margaritana crassa — Simpson (1900)
Margaritana (Group of M. margaritifera) crassa — Simpson (1914)

Unio manca ‘Lamarck’ of Authors

(misidentified reference to Unio manca Lamarck, 1819)
‘Lamarck’ of Authors.
Millet, 1843. Mag. Zool.: 4, pl. 64, fig. 2.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (Mancusiana) mancus — Germain (1904)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)
Unio mancus — Germain (1931)

Unio batavus var. pusillus Rossmässler, 1844

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio pusillus Lea, 1840)
Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): ??.

Unio drouetii Dupuy, 1849

Dupuy, 1849. Cat. Extramar. Galliae Test.: no. 326.
Dupuy, 1852. Hist. Natur. Moll. France (6): 639, pl. 24, fig. 12.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969), Fauna Europaea (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio ventricosus Scholtz, 1853

(homonym of Unio ventricosus Barnes, 1823)
Scholtz, 1853. unknown publication: ??.

Unio capigliolo ‘Payraudeau’ of Authors

(misidentified reference to Unio capigliolo Payraudeau, 1826)
‘Payraudeau’ of Authors.
Küster, 1855. Conchyl. Cab. 9(2): 125, pl. 34, figs. 5-6. [in part]
Sowerby, 1868. Conch. Icon. 16: pl. 75, sp. 392.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)

Unio rivularis Moquin-Tandon, 1855

(homonym of Unio rivularis Conrad, 1853)
Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): fig. 414.
Moquin-Tandon, 1855. Hist. Natur. Moll. France 2: 571.

Unio (Crassiana) crassus — Vinarski & Kantor (2016)

Unio batavus var. sarratinus ‘St. Germain, in litt.’ Moquin-Tandon, 1855

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)
‘St. Germain, in litt.’ Moquin-Tandon, 1855. Hist. Natur. Moll. France 2: 571.

Unio gangraenosus ‘Ziegler’ of Authors

(error for Unio gangrenosus J.F. Schmidt, 1840)
‘Ziegler’ of Authors.
Küster, 1855. Conchyl. Cab. 9(2): 124.
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 81.
Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 130.
Haas, 1969. Das Tierreich 88: 46.

Unio (Crassiana) crassus — Vinarski & Kantor (2016)

Unio courtillieri Hattemann, 1859

Hattemann, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 232.

Unio (Fusculusiana) fusculus — Germain (1904)
Unio crassus var. piscinalis — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969), Lopes-Lima et al. (2017)
Unio (Crassiana) crassus courtillieri — Falkner et al. (2001), Falkner et al. (2002)
Unio crassus courtillieri — Prié & Puillandre (2014), Lopes-Lima et al. (2017)
Unio crassus cf. courtillieri — Klishko et al. (2017)
Unio crassus courtillierii [sic] — Araujo et al. (2018)
Unio crassus — Anisimova et al. (2018), Bogatov et al. (2018), Lopes-Lima et al. (2024)
Unio c. courtillieri — Príe et al. (2021)
Unio cf. courtillieri — Lopes-Lima et al. (2021)

Unio batavus var. baraceus de Joannis, 1859

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 285, pl. 9, fig. 7.

Unio batavus var. baraceus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio batavus var. minimus de Joannis, 1859

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio requienii var. minima Drouët, 1857)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 289, pl. 9, fig. 6.

Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio batavus var. longus de Joannis, 1859

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio longus Philippi, 1869)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 288, pl. 10, fig. 3.

Unio batavus — Germain (1931)

Unio batavus var. acutus de Joannis, 1859

(homonym of Unio acutus J. de C. Sowerby, 1813)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 287, pl. 10, fig. 1.

Unio batavus var. senotius de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 288, pl. 10, fig. 4.

Unio sanderi Bourguignat, 1862

Bourguignat, 1862. Malacol. Lac Quatre-Cantons: 55.
Bourguignat, 1864. Malac. Aix-les-Baines: 76.

Unio mancus var. macrorhynchus — Germain (1931)

Unio merdiger Reeve, 1865

Jay, 1839. Cat. Rec. Shells: 25, sp. 1065.
‘Walsh.’ Lea, 1852. Synopsis Fam. Naiades: ??. [nomen nudum]
‘Watts’ Reeve, 1865. Conch. Icon. 16: pl. 28, sp. 145.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)

Unio lambottei Malzine, 1867

Malzine, 1867. Essai faun. malac. Belg.: 32, pl. 2, figs, 3-4.

Unio (Ateriana) lambottei — Germain (1904)
Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio ryckholtii Malzine, 1867

Malzine, 1867. Essai faun. malac. Belg.: 32, pl. 2, figs. 1-2.

Unio (Batavusiana) ryckholtii — Germain (1904)
Unio batavus — Germain (1931)

Unio batavus var. squamosus Brot, 1867

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; misidentified reference to Unio batavus var. squamosus Charpentier, 1837)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): pl. 9, fig. 1.

Unio consentaneus — Germain (1931)

Unio (tumidus) elegans Westerlund, 1867

(homonym of Unio elegans Lea, 1831)
Westerlund, 1867. Malak. Blatt. 14: 202, 206.

Unio (Crassiana) crassus — Vinarski & Kantor (2016)

Unio ryckholti var. cuneata Malzine, 1867

(intro'd as intra-sp. of Unio ryckholtii Malzine, 1867; homonym of Unio cuneatus Jacquemin, 1835)
Malzine, 1867. Essai faun. malac. Belg.: 32.

Unio batavus — Germain (1931)

Unio batavus var. belgicus Colbeau, 1868

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)
Colbeau, 1868. Ann. Soc. Malac. Belgique 3: 106, pl. 4, fig. 3.

Unio crassus var. maximus Kobelt, 1872

(intro'd as intra-sp. of Unio crassus Philipsson in Retzius, 1788; homonym of Unio pictorum maxima Mørch, 1863)
Rossmässler, 1853. Zeitschr. f. Malakozool. 10: 92 [as ‘Unio litoralis’].
Kobelt, 1872. Malak. Blatt. 19: 142, pl. 5.
Kobelt, 1884. Icon. Land- und Süssw.-Moll. 1: 67, pl. 29, figs. 218-219.
LECTOTYPE SMF 10967 (by Zilch, 1967: 53), in a stream near Aller, Denmark.

Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio pseudolitoralis Clessin, 1877

Clessin, 1875. Verh. Ver. Naturgesch. Hamburg 2: 258 [as ‘pseudo-litoralis’]. [nomen nudum]
Clessin, 1877. Deutsche Excurs. Moll. Fauna: 469, text fig. 305 [as ‘pseudo-litoralis’].

Unio (s.s.) crassus var. pseudolitoralis — Zhadin (1938)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio crassus var. pseudolitoralis — Zhadin (1965)
Batavusiana (Crassunio) crassa — Stadnichenko (1984), Zatravkin (1987)
Crassiana crassa crassa — Kantor & Sysoev (2005), Kantor et al. (2010)
Crassiana pseudolitoralis — Gozhik (2006)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio subtilis Drouët, 1879

Drouët, 1879. J. de Conch. 27: 142.
SYNTYPES SMF 3712, Aube, France.

Unio (Groupe des Mancusiana) subtilis — Caziot (1907)
Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)
Unio mancus — Nagel (2025)

Unio vescoi var. conemenosi Blanc in Westerlund & Blanc, 1879

(intro'd as intra-sp. of Unio vescoi Bourguignat, 1856)
Blanc in Westerlund & Blanc, 1879. Aperçu Faune Malac. Grèce: 145.

Unio requieni Brevière, 1880

(misidentified reference to Unio requienii Michaud, 1831)
‘Mich.’ Brevière, 1880. Cat. moll. Nièvre: 26.

Unio consentaneus — Germain (1931)

Unio moquinianus Brevière, 1880

(misidentified reference to Unio moquinianus Dupuy, 1843)
Brevière, 1880. Cat. moll. Nièvre: 26.

Unio consentaneus — Germain (1931)

Unio ligericus Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 289, 358.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 142 [36].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 169.
IDIOTYPE MNHN_22310, La Loire à Ingrandes.

Unio (Batavusiana) ligericus — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio sequanicus Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 289, 358.
‘Coutagne’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 142 [36].
‘Coutagne’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 169, fig. 183.
LECTOTYPE MNHN_22529 (by Graf, 2011: 504), La Marne à Lagny (Seine-et-Marne).

Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio pilloti Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 291, 360.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 128 [22].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 157.
IDIOTYPE MNHN_22460, Passavant (Haute-Saône).

Unio pilloti — Kobelt & Haas (1910)
Unio mancus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus mancus — Prie (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio rayi Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 291, 360.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 127 [21].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 155.
LECTOTYPE MNHN_22483 (by Graf, 2011: 500), Gigny (Jura).

Unio (Groupe des Ninusiana) [sic] rayi — Caziot (1907)
Unio rayi — Kobelt & Haas (1910)
Unio mancus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus mancus — Prie (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassatellus Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 286, 356.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 136 [30].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 162.
LECTOTYPE MNHN_22182 (by Graf, 2011: 469), La Canne, à Jailly (Nièvre).

Unio crassatellus — Rochebrune (1903)
Unio crassus — Germain (1931), Lopes-Lima et al. (2024)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio locardianus Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 287, 356.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 138 [32].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 164, fig. 178.
IDIOTYPE MNHN_22316, l’Albane (Côte-d’Or).

Unio crassus — Germain (1931), Lopes-Lima et al. (2024)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio lagnisicus Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 291, 359.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 127 [21] [as ‘lagnysicus’].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 154 [as ‘lagnysicus’].
SYNTYPE MNHN_22303, Condal (Saône-et-Loire).

Unio lagnysicus [sic] — Rochebrune (1903), Kobelt & Haas (1910)
Unio mancus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus mancus — Prie (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio carantoni Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 295, 364.
‘Coutagne’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 129 [23].
‘Coutagne’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 157.
IDIOTYPE MNHN_22162, [illegible] (Charente).

Unio carantoni — Rochebrune (1903)
Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio berthelini Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 290, 359.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 137 [31].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 163.
IDIOTYPE MNHN_22132, La Marne, à Charenton (Seine).

Unio mancus var. amnicus form riparius — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio andegavensis Locard, 1882

‘Servain’ Locard, 1882. Prodr. Malac. Franç. 1: 289, 359.
‘Servain’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 142 [36].
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 167.
POSSIBLE SYNTYPE MNHN_22100, La Maine à Angers.

Unio (Batavusiana) andegavensis — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio cyprinorum Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 289, 358.
‘Berthier’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 142 [36].
‘Berthier’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 168.
IDIOTYPE MNHN_22185, l’Ornain (Meuse).

Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Crassiana (Bataviana) cyprinorum — Starobogatov et al. (2004)
Crassiana cyprinorum — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus — Graf (2007), Lopes-Lima et al. (2024)
Unio (Crassiana) cyprinorum — Vinarski & Kantor (2016)
Unio crassus musivus — Bogatov (2022)

Unio matronicus Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 289, 358.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 143 [37].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 170.
LECTOTYPE MNHN_22355 (by Graf, 2011: 489), La Marne à Lagny (Seine-et-Marne).

Unio (Batavusiana) matronicus — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio septentrionalis Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 284, 354.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 149 [43].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 181, fig. 194.
IDIOTYPE MNHN_22528, le Mandrezey, à Saulcy-sur-Meurthe (Vosges).

Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio oxyrhynchus Bourguignat in Locard, 1882

(homonym of Unio oxyrhynchus Martens, 1861)
Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 285, 355.
‘Brevière’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 149 [43].
‘Brevière’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 180.
LECTOTYPE MNHN_22438 (by Graf, 2011: 495), La Canne, dans la Nièvre.

Unio consentaneus var. oxyrhynchus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio marcellinus Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 285, 355.
‘Berthier’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 149 [43].
‘Berthier’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 180.

Unio (Ateriana) marcellinus — Germain (1904)
Unio consentaneus var. moquini — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus moquinianus — Prie (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio feliciani Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 285, 355.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 134 [28].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 161, fig. 175.
Bourguignat in Locard, 1899. Conchyliol. Portug.: 217.
POSSIBLE SYNTYPE MNHN_22215, La Moselle.

Unio (Socardiana) feliciani — Germain (1904)
Unio batavus var. baraceus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio arenarum Bourguignat in Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 289, 358.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 144 [38].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 171.
IDIOTYPE MNHN_22110, Basse Indre (Loire-Inf.).

Unio (Batavusiana) arenarum — Germain (1904)
Unio batavus var. baraceus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio brevierei Locard, 1882

(correct name for Unio brevieri ‘Bourguignat in Locard’ of Authors)
‘Bourguignat’ Locard, 1882. Prodr. Malac. Franç. 1: 286 [as ‘brevieri’], 356.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 148 [42].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 178.
LECTOTYPE MNHN_22145 (by Graf, 2011: 465), La Canne, à Goulnot.

Unio (Ateriana) brevieri [sic] — Germain (1904)
Unio (Groupe des Ateriana) brevierei — Caziot (1907)
Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio brevieri ‘Bourguignat in Locard’ of Authors

(incorrect original spelling of Unio brevierei Locard, 1882)
‘Bourguignat in Locard’ of Authors.
Locard, 1882. Prodr. Malac. Franç. 1: 286.
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 265.
Germain, 1931. Faune de France 22: 732.
Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 130.
Haas, 1969. Das Tierreich 88: 49.

Unio crassus — Lopes-Lima et al. (2024)

Unio batavellus Locard, 1885

‘Letourneux’ Locard, 1885. Bull. Soc. Sci. Nat. Rouen 21: 11.
‘Letourneux’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 144 [38].
‘Letourneux’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 170, fig. 184.
LECTOTYPE MNHN_22127 (by Graf, 2011: 462), La Bouille, près de Rouen (Seine-Inf.).

Unio batavellus — Rochebrune (1903), Kobelt & Haas (1910)
Unio (Batavusiana) batavellus — Germain (1904)
Unio batavus var. baraceus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio carcasinus Sourbieu, 1887

Sourbieu, 1887. Bull. Soc. Malac. France 4: 235.

Unio crassus var. piscinalis — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio baroni Servain, 1887

Servain, 1887. Bull. Soc. Malac. France 4: 254.

Unio hamburgiensis Servain, 1888

Servain, 1888. Bull. Soc. Malac. France 5: 315.

Unio crassus — Germain (1931), Lopes-Lima et al. (2024)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)

Unio baudoni Folin, 1888

(homonym of Unio baudoni Folin & Bérillon, 1874)
Folin, 1888. Le Naturaliste 10: 273, 4 text figs.

Unio baudoni — Kobelt (1902)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio crassulus Drouët, 1888

(homonym of Unio requienii crassulus Drouët, 1867)
Drouët, 1888. J. de Conch. 36: 106.
Drouët, 1889. Mém. Acad. Sci. Dijon 1: 66, pl. 2, fig. 5.
SYNTYPE MNHN-IM-2000-1666, Saone.

Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio vegesackensis Bourguignat in Servain, 1888

Bourguignat in Servain, 1888. Bull. Soc. Malac. France 5: 317.

Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio visurgisinus Servain, 1888

Servain, 1888. Bull. Soc. Malac. France 5: 316.

Unio (Batavusiana) visurgicus [sic] — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio minutulus Servain, 1888

‘Ray’ Locard, 1882. Prodr. Malac. Franç. 1: 290 [as ‘Unio minutus’]. [nomen nudum]
‘Ray’ Servain, 1888. Bull. Soc. Malac. France 5: 316.

Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1940)

Unio besnardianus Bourguignat in Locard, 1889

(renamed Unio besnardi Germain, 1904)
Servain, 1888. Bull. Soc. Malac. France 5: 317. [nomen nudum]
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 141 [35], 206 [100].
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 172, fig. 186 [as ‘besnardinus’].
LECTOTYPE MNHN_22133 (by Graf, 2011: 463), La Loire, à Saint-Gemmes (Maine-et-Loire).

Unio (Batavusiana) besnardi — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio catalaunicus Bourguignat in Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 133 [27], 192 [86].
‘Coutagne’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 160.
LECTOTYPE MNHN_22171 (by Graf, 2011: 467), La Marne, à Charenton (Seine).

Unio crassus var. piscinalis — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio subamnicus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 137 [31], 194 [88].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 163.
SYNTYPE MNHN_22559, La Saône, à Auxonne (Côte-d’Or).

Unio subamnicus — Kobelt & Haas (1910)
Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio aturicus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 129 [23], 188 [82].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 157.
HOLOTYPE (MONOTYPY) MNHN_22118, Bahus (Landes).

Unio mancus var. amnicus form riparius — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio lemotheuxi Bourguignat in Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 145 [39], 208 [102].
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 175, fig. 189.
LECTOTYPE MNHN_22306 (by Graf, 2011: 484), le Reyran (Var).

Unio (Lemotheuxiana) lemotheuxi — Germain (1904), Caziot (1910)
Unio mancus var. elongatulus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio andeliacus Bourguignat in Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 138 [32], 196 [90].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 165.
LECTOTYPE MNHN_22101 (by Graf, 2011: 458), le Gouet, à Saint-Brieue (Côtes-du-Nord).

Unio mancus var. elongatulus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio diptychus Bourguignat in Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 143 [37], 201 [95].
‘Surault’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 169.
POSSIBLE SYNTYPE MNHN_22192, La Loire à Ingrandes.

Unio (Batavusiana) diptychus — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio ingrandiensis Bourguignat in Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 143 [37], 202 [96].
‘Sarault’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 173.
POSSIBLE SYNTYPE MNHN_22282, Ingrandes (Maine-et-Loire).

Unio (Batavusiana) ingrandiensis — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio materniacus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 143 [37], 203 [97].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 168, fig. 182.
LECTOTYPE MNHN_22347 (by Graf, 2011: 488), la Somme à Abbeville.

Unio materniacus — Rochebrune (1903)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio melantatus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 147 [41], 216 [110].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 177.
LECTOTYPE MNHN_22360 (by Graf, 2011: 489), La Loire à Balbigny (Loire).

Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio balbignyanus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 147 [41], 217 [111].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 178.
LECTOTYPE MNHN_22122 (by Graf, 2011: 462), La Loire, à Balbigny (Loire).

Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio occidentalis Locard, 1889

(homonym of Unio occidentalis Conrad, 1836)
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 149 [43], 220 [114].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 180.
‘Bourguignat’ Locard, 1899. Conchyliol. Portug.: 220.
LECTOTYPE MNHN_22424 (by Graf, 2011: 494), Balbigny (Loire).

Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio bouchardi Locard, 1889

(new name for Unio arcuata Bouchard-Chantereaux, 1838)
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 149 [43].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 180.
LECTOTYPE FIGURED SPECIMEN (by Graf, 2011: 464), ditches of Saint-Omer, fed by the Aa.

Unio consentaneus var. moquini — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus moquinianus — Prie (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio surraulti Bourguignat in Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 144 [38], 204 [98].
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174 [as ‘suraulti’].
SYNTYPE MNHN_22569, La Loire à Ingrandes.

Unio (Batavusiana) surraulti — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio adonus Locard, 1889

‘Servain’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 146 [40], 211 [105].
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 175, fig. 188.
LECTOTYPE MNHN_22085 (by Graf, 2011: 456), La Loire a Basse Indre (Loire-Inf.).

Unio (Lemotheuxiana) adonus — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio caumonti Bourguignat in Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 145 [39], 207 [101].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 171.

Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio hattemani Bourguignat in Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 146 [40], 212 [106].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 176.
SYNTYPE MNHN_22254, La Loire, à Basse-Indre.

Unio (Lemotheuxiana) hattemanni [sic] — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio seneauxi Bourguignat in Locard, 1889

‘Linn.’ Draparnaud, 1806. Hist. Moll. France: 131, pl. 11, figs. 3 [as ‘Unio pictorum var. B’].
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 145 [39], 208 [102].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174, fig. 187 [as ‘senauxi’].
LECTOTYPE MNHN_22523 (by Graf, 2011: 503), La Loire à Orléans.

Unio (Batavusiana) seneauxi — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio visurgicus ‘Servain’ of Authors

(error for Unio visurgisinus Servain, 1888)
‘Servain’ of Authors.
Locard, 1889. Contr. Faune Malacol. Franc. 13: 141 [35].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 173.
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 262.

Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio caroliensis Bourguignat in Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 159 [53], 237 [131].
‘F. Pacome’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 172.
LECTOTYPE MNHN_22167 (by Graf, 2011: 467), Charolles (Saône-et-Loire).

Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio mariae Bourguignat in Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 159 [53], 236 [130] [as ‘mariæ’].
‘F. Pacome’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 172 [as ‘mariæ’].
IDIOTYPE MNHN_22342, La Loire à Nantes (Loire-Infereur).

Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio nicolloni Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 141 [35], 200 [94].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 173.
HOLOTYPE (MONOTYPY) MNHN_22404, Nantes.

Unio (Batavusiana) nicolloni — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio ignariformis Bourguignat in Locard, 1889

‘Dup.’ Drouët, 1857. Étud. Naïades. France, Unio: 100, pl. 9, fig. 1 [as ‘U. platyrhynchoideus’].
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 147 [41], 214 [108].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 177.

Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio financei Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 143 [37], 205 [99].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 171.
IDIOTYPE MNHN_22217, La Marne à Châlons (Marne).

Unio batavus — Germain (1931)

Unio lagnysicus ‘Bourguignat’ of Authors

(error for Unio lagnisicus Bourguignat in Locard, 1882)
‘Bourguignat’ of Authors.
Locard, 1889. Contr. Faune Malacol. Franc. 13: 127 [21].
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 84.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 154.
Dumas, 1895. Rev. Sci. Bourbonnais, Suppl.: 47.
Rochebrune, 1903. Bull. Mus. Nation. Hist. Natur. 9: 389.
Kobelt & Haas, 1910. Icon. Land- und Süssw.-Moll. 17: 37.

Unio subrobustus var. curta Locard, 1889

(homonym of Unio curtus Lea, 1859; intro'd as intra-sp. of Unio subrobustus Servain, 1885)
Locard, 1889. Contr. Faune Malacol. Franc. 13: 135 [29].

Unio gliscerus Locard, 1893

Locard, 1893. Coqu. Eau Douce Saumâtre France: 168.
LECTOTYPE MNHN_22241 (by Graf, 2011: 477), La Loire à Nantes (Loire-Infereur).

Unio (Batavusiana) gliscerus — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio ampullaceus Locard, 1893

(homonym of Unio ampullaceus Lea, 1866)
Locard, 1893. Coqu. Eau Douce Saumâtre France: 174.
LECTOTYPE MNHN_22098 (by Graf, 2011: 458), Le Mans (Sarthe).

Unio (Batavusiana) ampullaceus — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio chardoni Locard, 1893

‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 173.

Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio lancelevei Locard, 1893

Locard, 1893. Bull. Soc. Sci. Nat. Elbeuf 12: 1.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 170.
SYNTYPE MNHN_22305, le Seine au nord d. Rouen.

Unio batavus var. baraceus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio lamboltei ‘Malzine’ of Authors

(error for Unio lambottei Malzine, 1867)
‘Malzine’ of Authors.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 177.
Germain, 1931. Faune de France 22: 732.
Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 130.

Unio lepidulus Drouët, 1898

Drouët, 1898. Mém. Acad. Sci. Dijon ??: 60, pl. 1, fig. 7.

Unio mancus var. elongatulus — Germain (1931)

Unio batavus var. major Drouët, 1898

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio (Anodontes) sirionos var. major d’Orbigny, 1835)
Drouët, 1898. Mém. Acad. Sci. Dijon ??: 61.

Unio batavus var. baraceus — Germain (1931)

Unio ellipsopsis Bourguignat in Locard, 1899

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 140 [34]. [nomen nudum]
Bourguignat in Locard, 1899. Conchyliol. Portug.: 218.

Unio ater var. irgizlaicus Lindholm, 1904

(intro'd as intra-sp. of Unio ater Nilsson, 1822)
Lindholm, 1904. Ann. Mus. St. Pétersbourg 8(3-4): 343.

Unio crassus crassus — Haas (1969), Klishko et al. (2017)
Batavusiana (s.s.) irgislaica [sic] — Zatravkin (1987)
Crassiana (Bataviana) irgizlaica — Starobogatov et al. (2004)
Crassiana irgizlaica — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus — Graf (2007), Lopes-Lima et al. (2024)
Unio (Crassiana) irgizlaicus — Vinarski & Kantor (2016)
Unio crassus musivus — Bogatov (2022)

Unio batavus var. radiata Germain, 1904

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio (Elliptio) viridis var. radiata Rafinesque, 1820)
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 257 [19].

Unio (Batavusiana) batavus radiata — Germain (1904)

Unio ligericus var. subsinuatus Germain, 1904

(intro'd as intra-sp. of Unio ligericus Bourguignat in Locard, 1882)
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 259 [21].

Unio (Batavusiana) ligericus subsinuatus — Germain (1904)

Unio batavellus var. perconvexus Germain, 1904

(intro'd as intra-sp. of Unio batavellus Locard, 1885)
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 260 [22].

Unio (Batavusiana) batavellus perconvexus — Germain (1904)

Unio besnardi Germain, 1904

(new name for Unio besnardianus Bourguignat in Locard, 1889)
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 261 [23].

Unio (Batavusiana) besnardi — Germain (1904)

Unio pseudolittoralis [sic] var. curonicus Riemenschneider, 1907

(intro'd as intra-sp. of Unio pseudolitoralis Clessin, 1877)
Riemenschneider, 1907. Nachr. Bl. Dtsch. Malak. Ges. 39: 150.

Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassus var. okae Kobelt & Haas, 1910

(intro'd as intra-sp. of Unio crassus Philipsson in Retzius, 1788)
Kobelt & Haas, 1910. Icon. Land- und Süssw.-Moll. 17: 58, index, pl. 480, fig. 2567.
HOLOTYPE SMF 10971, Oka River, Serpukhov, Russia.

Unio (s.s.) crassus var. okae — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio crassus heimburgi Haas, 1912

(intro'd as intra-sp. of Unio crassus Philipsson in Retzius, 1788)
Haas, 1912. Verh. Naturhist. Ver. Preuß. Rheinl. 68: 527, pl. 6, fig. 20.
HOLOTYPE SMF 4389, Hatfeld, Germany.

Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio kungurensis Kobelt, 1912

Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 24, pl. 490, figs. 2622-2623.
HOLOTYPE SMF 10972, Reka Iren’, Kungur, Perm Krai, Russia.

Unio (s.s.) crassus kungurensis — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio kungurensis irenjensis Kobelt, 1912

(intro'd as intra-sp. of Unio kungurensis Kobelt, 1912)
Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 26, pl. 491, fig. 2626.
HOLOTYPE SMF 10975, Flüschen Irenj, Kungur, Russia.

Unio (s.s.) crassus var. chlebnikovi [sic] — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969), Graf (2007)
Batavusiana (s.s.) irenjensis — Zatravkin (1983), Stadnichenko (1984), Zatravkin (1987)
Crassiana irenjensis — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio (Crassiana) irenjensis — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio kungurensis chlebnikowi Kobelt, 1912

(intro'd as intra-sp. of Unio kungurensis Kobelt, 1912; renamed Unio crassus var. chlebnikovi Zhadin, 1938)
Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 26, pl. 491, fig. 2624.
HOLOTYPE SMF 10973, Iren River, Kungur, Russia.

Unio (s.s.) crassus var. chlebnikovi [sic] — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio kungurensis sylwensis Kobelt, 1912

(intro'd as intra-sp. of Unio kungurensis Kobelt, 1912)
Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 26, pl. 491, fig. 2625.
LECTOTYPE SMF 10974 (by Zilch, 1967: 67), Kungur, Russia.

Unio (s.s.) crassus var. chlebnikovi [sic] — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio roseni Kobelt, 1912

Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 27, pl. 492, fig. 2627.
LECTOTYPE SMF 10976 (by Zilch, 1967: 67), near former Kuriazh monastery, W Kjharvin, Ukraine.

Unio (s.s.) crassus var. roseni — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024), Nagel (2025)

Unio crassus thuringiacus Israel, 1913

(intro'd as intra-sp. of Unio crassus Philipsson in Retzius, 1788)
Israel, 1913. [in] Thüring. Lehrerverein Naturk.: 28, pl. 6, figs. 4-6.

Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassus polonicus Polinksi, 1917

(intro'd as intra-sp. of Unio crassus Philipsson in Retzius, 1788)
Polinksi, 1917. C. R. Soc. Sci. Varsovie 10: 902, pl. ??, figs. 1, 3.

Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassus ornatus Polinksi, 1917

(homonym of Unio ovatus var. ornatus Conrad, 1835; intro'd as intra-sp. of Unio crassus Philipsson in Retzius, 1788)
Polinksi, 1917. C. R. Soc. Sci. Varsovie 10: 904, pl. ??, fig. 2.

Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio cantianus Kennard & Woodward, 1924

Kennard & Woodward, 1924. Proc. Malac. Soc. London 16: 136, text figs. 8-9.

Unio crassus batavus — Haas (1940), Haas (1969)

Unio lagnysiacus ‘Bourguignat in Locard’ of Authors

(error for Unio lagnisicus Bourguignat in Locard, 1882)
‘Bourguignat in Locard’ of Authors.
Germain, 1931. Faune de France 22: 724.
Prie, 2012. MalaCo 8: 430.

Unio crassus var. ishmensis Zhadin, 1938

(intro'd as intra-sp. of Unio crassus Philipsson in Retzius, 1788)
Zhadin, 1938. Faune USSR Mollusques 4(1): 160.

Unio (s.s.) crassus var. ishmensis — Zhadin (1938)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassus var. chlebnikovi Zhadin, 1938

(new name for Unio kungurensis chlebnikowi Kobelt, 1912)
‘Kobelt’ Zhadin, 1938. Faune USSR Mollusques 4(1): 89.

Unio intermedius ‘Rm.’ of Authors

‘Rm.’ of Authors.

Unio inflatus Locard 1881 of Authors

Locard 1881 of Authors.

Unio courtilieri ‘Hattemann’ of Authors

(error for Unio courtillieri Hattemann, 1859)
‘Hattemann’ of Authors.
Locard, 1889. Contr. Faune Malacol. Franc. 13: 132 [26].
‘Hattemann’ Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 256 [18].
‘Hattmann’ Germain, 1931. Faune de France 22: 724, 727.
Lopes-Lima et al., 2024. MPE 195: suppl. append. 2, p. 3.

Unio platyrhynchoideus de Cessac of Authors

de Cessac of Authors.

Unio coactatus Ziégl. in sched. of Authors

Ziégl. in sched. of Authors.

Unio pseudolittoralis ‘Clessin’ of Authors

(error for Unio pseudolitoralis Clessin, 1877)
‘Clessin’ of Authors.
Riemenschneider, 1907. Nachr. Bl. Dtsch. Malak. Ges. 39: 150.
Vinarski & Kantor, 2016. Analyt. Cat. Fr. Brack. Moll. Russia: 25.

Unio pictorum Draparnaud, 1806

(misidentified reference to Mya pictorum Linnaeus, 1758)
‘Linn.’ Draparnaud, 1806. Hist. Moll. France: 131, pl. 11, figs. 3 [var. B]. [in part]

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (Batavusiana) seneauxi — Germain (1904)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)
Unio batavus — Germain (1931)

Unio margaritifera Draparnaud, 1806

(misidentified reference to Mya margaritifera Linnaeus, 1758)
‘Linn.’ Draparnaud, 1806. Hist. Moll. France: pl. 11, fig. 5.

Unio consentaneus — Germain (1931)

Unio crassus Nilsson, 1822

(reference to Unio crassus Philipsson in Retzius, 1788)
‘Retzius’ Nilsson, 1822. Hist. Moll. Svec.: 108.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)

Unio batavus Nilsson, 1822

(reference to Mya batava Maton & Rackett, 1807)
‘Lamarck’ Nilsson, 1822. Hist. Moll. Svec.: 112.

Unio rubens Menke, 1830

(reference to Unio rubens Menke, 1828)
Menke, 1830. Syn. Method. Moll.: 149.

Unio (sec. Lymnium) littoralis — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) littoralis — Simpson (1914)

Unio lemovicensis Kickx, 1830

(error for Unio lemovicincae Férussac, 1807)
‘Daud.’ Kickx, 1830. Synopsin Molluscorum Brabantiae: 82.

Unio gibba Lea, 1836

(intro'd as syn of Mya batava Maton & Rackett, 1807)
‘Mühl. Pfeif.’ Lea, 1836. Synopsis Fam. Naiades: 27.

Unio fuscus Lea, 1836

(intro'd as syn of Mya batava Maton & Rackett, 1807)
‘Zieg.’ Lea, 1836. Synopsis Fam. Naiades: 27.

Unio acutus Rossmässler, 1837

Rossmässler, 1837. Icon. Land- und Süssw.-Moll. 1 (5-6): 56, pl 29, fig. 413 [as ‘Unio an n. sp.?’]. [not available]

Unio gargottae Rossmässler, 1838

(misidentified reference to Unio gargottae Philippi, 1836)
Rossmässler, 1838. Icon. Land- und Süssw.-Moll. 2 (1-2): 26, pl. 35, fig. 493.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)

Mysca reniformis subvar. kokeilli Schlüter, 1838

(intro'd as intra-sp. of Unio reniformis Rossmässler, 1836)
Schlüter, 1838. Kurzgef. Syst. Verz. Conch. Samml.: 32. [infra-subsp.]

Mysca batava var. fuscula subvar. pallipes Schlüter, 1838

(intro'd as intra-sp. of Unio fusculus Rossmässler, 1836)
‘Ziegl.’ Schlüter, 1838. Kurzgef. Syst. Verz. Conch. Samml.: 33. [infra-subsp.]

Unio requieni Rossmässler, 1844

(misidentified reference to Unio requienii Michaud, 1831)
‘Mich.’ Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): 29, pl. 57, fig. 757-761.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) batavus var. requieni — Simpson (1914)

Unio glaucinus Stabile, 1845

(misidentified reference to Unio glaucinus Porro, 1838)
Stabile, 1845. Faun. Lug.: f. 75.

Unio (sec. Lymnium) batavus — Simpson (1900)

Unio retzii C.B. Adams, 1847

‘Küst.’ C.B. Adams, 1847. Cat. Gen. Sp. Adams: 14. [nomen nudum]

Unio piscinalis J.F. Schmidt, 1847

(misidentified reference to Unio piscinalis Rossmässler, 1836)
J.F. Schmidt, 1847. Conch. Krain: 27.

Unio moquinianus Sandberger & Koch, 1851

(misidentified reference to Unio moquinianus Dupuy, 1843)
Sandberger & Koch, 1851. Jb. Nassau. Ver. Naturk. 7: 245.

Unio nanus Dupuy, 1852

(misidentified reference to Unio nana Lamarck, 1819)
‘Lamarck’ Dupuy, 1852. Hist. Natur. Moll. France (6): pl. 25, fig. 16.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (Rhomboïdiana) nana — Germain (1904)
Unio nana — Kobelt & Haas (1910)
Unio (sec. Lymnium) (Group U. littoralis) batavus var. requieni — Simpson (1914)
Unio mancus var. nanus — Germain (1931)

Unio ovalis Dupuy, 1852

(misidentified reference to Mya ovalis Montagu, 1803)
Dupuy, 1852. Hist. Natur. Moll. France (6): pl. 25, fig. 13.

Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (Batavusiana) besnardi — Germain (1904)
Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)
Unio batavus — Germain (1931)

Unio argens Lea, 1852

(intro'd as syn of Mya batava Maton & Rackett, 1807)
‘Küst.’ Lea, 1852. Synopsis Fam. Naiades: 30.

Unio kurrii Lea, 1852

(intro'd as syn of Mya batava Maton & Rackett, 1807)
‘Küst.’ Lea, 1852. Synopsis Fam. Naiades: 30.

Unio drouatii Lea, 1852

(error for Unio drouetii Dupuy, 1849)
‘Dup.’ Lea, 1852. Synopsis Fam. Naiades: 39.

Unio moquinianus Drouët, 1852

(misidentified reference to Unio moquinianus Dupuy, 1843)
Drouët, 1852. Etudes Nayades de la France: pl. 6, fig. 3.

Unio consentaneus — Germain (1931)

Unio crassus Küster, 1854

(reference to Unio crassus Philipsson in Retzius, 1788)
‘Retz.’ Küster, 1854. Conchyl. Cab. 9(2): 113, pl. 31, fig. 5. [in part]

Lampsilis (sec. Eurynia) radiatus — Simpson (1900)
Lampsilis (Eurynia) (Group of L. luteola) radiata — Simpson (1914)

Unio ater Moquin-Tandon, 1855

(reference to Unio ater Nilsson, 1822)
‘Nilss.’ Moquin-Tandon, 1855. Hist. Natur. Moll. France 2: 570.

Unio ater Drouët, 1857

(reference to Unio ater Nilsson, 1822)
‘Nilss.’ Drouët, 1857. Étud. Naïades. France, Unio: 72, pl. 4, fig. 1.

Unio consentaneus — Germain (1931)

Unio platyrhynchoideus Drouët, 1857

(misidentified reference to Unio platyrinchoideus Dupuy, 1849)
‘Dup.’ Drouët, 1857. Étud. Naïades. France, Unio: 100, pl. 9, fig. 1.

Unio nanus Drouët, 1857

(misidentified reference to Unio nana Lamarck, 1819)
‘Lam.’ Drouët, 1857. Étud. Naïades. France, Unio: pl. 5, fig. 2.

Unio (Rhomboïdiana) nana — Germain (1904)
Unio nana — Kobelt & Haas (1910)
Unio mancus var. nanus — Germain (1931)

Unio batavus var. ovalis de Joannis, 1859

(misidentified reference to Mya ovalis Montagu, 1803)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 10, fig. 2.

Unio (Batavusiana) ligericus — Germain (1904)
Unio batavus — Germain (1931)

Unio batavus var. nanus de Joannis, 1859

(misidentified reference to Unio nana Lamarck, 1819)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 8. [in part]

Unio (Melasiana) melas — Germain (1904)
Unio mancus var. amnicus — Germain (1931)

Unio batavus var. courtilieri de Joannis, 1859

(reference to Unio courtillieri Hattemann, 1859; error for Unio courtillieri Hattemann, 1859)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 3.

Unio (Batavusiana) batavus courtilieri — Germain (1904)
Unio batavus — Germain (1931)

Unio batavus var. nanus de Joannis, 1859

(misidentified reference to Unio nana Lamarck, 1819)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 10. [in part]

Unio (Rhomboïdiana) nana — Germain (1904)

Unio batavus de Joannis, 1859

(reference to Mya batava Maton & Rackett, 1807)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 2. [in part]

Unio (Batavusiana) batavus radiata — Germain (1904)

Unio orientalis ‘Parreis in sched.’ Küster, 1862

(intro'd as syn of Unio kotschyi Küster, 1862)
‘Parreis in sched.’ Küster, 1862. Conchyl. Cab. 9(2): 267.

Unio ater Bourguignat, 1864

(reference to Unio ater Nilsson, 1822)
Bourguignat, 1864. Malac. Aix-les-Baines: 73.

Unio mancus var. macrorhynchus — Germain (1931)

Unio batavus var. fontainei Colbeau, 1868

(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)
‘Lecomte’ Colbeau, 1868. Ann. Soc. Malac. Belgique 3: 106. [nomen nudum]

Unio maceerthyanus Lea, 1870

(error for Unio maccarthyanus Bourguignat, 1866; misidentified reference to Unio maccarthyanus Bourguignat, 1866)
‘Bourg.’ Lea, 1870. Synopsis Fam. Naiades: 48.

Unio batavus Miller, 1873

(reference to Mya batava Maton & Rackett, 1807)
Miller, 1873. Schalth. Bodensees: pl. 2, fig. 8.

Unio crassus — Germain (1931)

Unio diagonalis Drouët, 1881

(intro'd as syn of Unio batavus var. squamosus Charpentier, 1837)
‘Held’ Drouët, 1881. J. de Conch. 29: 27.

Unio minutus Locard, 1882

‘Ray’ Locard, 1882. Prodr. Malac. Franç. 1: 290. [nomen nudum]

Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio sandrii Locard, 1882

(misidentified reference to Unio sandrii Rossmässler, 1844)
Locard, 1882. Prodr. Malac. Franç. 1: 292.

Unio hueti Kobelt, 1886

(misidentified reference to Unio hueti Bourguignat, 1855)
‘Bourg.’ Kobelt, 1886. Icon. Land- und Süssw.-Moll. 2: 22, pl. 41, fig. 255.

Unio (sec. Lymnium) pseudonymus — Simpson (1900)
Unio (sec. Lymnium) (Group U. littoralis) pseudonymus — Simpson (1914)

Unio sieversi Kobelt, 1886

(misidentified reference to Unio sieversi Drouët, 1881)
Kobelt, 1886. Icon. Land- und Süssw.-Moll. 2: pl. 53, fig. 286.

Unio dilophius Servain, 1888

‘Bourguignat’ Servain, 1888. Bull. Soc. Malac. France 5: 316, footnote. [nomen nudum]
Locard, 1889. Contr. Faune Malacol. Franc. 13: 136 [30], footnote. [nomen nudum]

Unio suborbicularis var. minor Locard, 1889

(intro'd as intra-sp. of Unio suborbicularis Drouët, 1888)
Locard, 1889. Contr. Faune Malacol. Franc. 13: 127 [21]. [nomen nudum]
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 84. [nomen nudum]

Unio sperchinus Locard, 1889

‘Thièse’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 141 [35], footnote. [nomen nudum]

Unio pilloti var. minor Locard, 1889

(intro'd as intra-sp. of Unio pilloti Bourguignat in Locard, 1882)
Locard, 1889. Contr. Faune Malacol. Franc. 13: 129 [23]. [nomen nudum]

Unio besnardinus Locard, 1893

(error for Unio besnardianus Bourguignat in Locard, 1889)
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 172.

Unio (Batavusiana) besnardi — Germain (1904)

Unio senauxi Locard, 1893

(error for Unio seneauxi Bourguignat in Locard, 1889)
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174.

Unio suraulti ‘Servain in Locard’ Locard, 1893

(error for Unio surraulti Bourguignat in Locard, 1889)
‘Servain in Locard’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174.

Unio musicus Simpson, 1900

(error for Unio musivus Spengler, 1793)
‘Spengler’ Simpson, 1900. Proc. U.S. Nat. Mus. 22: 744.

Unio (sec. Lymnium) (Group U. littoralis) batavus — Simpson (1914)

Unio hattemanni Germain, 1904

(error for Unio hattemani Bourguignat in Locard, 1889)
‘Bourguignat’ Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 265 [27].

Unio visursisinus Germain, 1931

(error for Unio visurgisinus Servain, 1888)
‘Servain’ Germain, 1931. Faune de France 22: 728.

Unio carioliensis Germain, 1931

(error for Unio caroliensis Bourguignat in Locard, 1889)
‘Pacôme’ Germain, 1931. Faune de France 22: 728.

Unio kungurensis sylvensis Haas, 1940

(error for Unio kungurensis sylwensis Kobelt, 1912)
‘Kobelt’ Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 132.
‘Kobelt’ Haas, 1969. Das Tierreich 88: 53.
Vinarski & Kantor, 2016. Analyt. Cat. Fr. Brack. Moll. Russia: 25.

Unio hattmanni Haas, 1940

(error for Unio hattemani Bourguignat in Locard, 1889)
‘Bourguignat’ Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 131.

Unio lancelevi Haas, 1969

(error for Unio lancelevei Locard, 1893)
‘Locard’ Haas, 1969. Das Tierreich 88: 51.

Unio crassus var. irgislaicus Zatravkin, 1987

(error for Unio ater var. irgizlaicus Lindholm, 1904)
‘Lindholm’ Zatravkin, 1987. Byull. Mosk. Obshch.. Ispyt. Prir. 92(4): 57.