MUSSELp

MUSSELpdb

Introduction

Browse

d'basing

Search the MUSSELpdb

Taxonomy

all genera

Unionidae

Margaritiferidae

Hyriidae

Etheriidae

Mycetopodidae

Iridinidae


Species & Genus Richness by
Taxonomy

Geography

Summary

North America

Central America

South America

Afrotropics

North Eurasia

East Asia

Australasia


Species Richness & Endemism by
Geography

Data last updated
Mon 1 Dec 2025

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini | Unio

species Unio crassus Philipsson in Retzius, 1788: synonymy

98 specimens + 264 Genbank vouchers

Synonymy

Mya ovalis Lightfoot, 1786

Lister, 1685. Conch.: pl. 146, fig. 1. [pre-linnaean]
‘S.’ Lightfoot, 1786. Catalogue of the Portland Museum: 134, no. 2983.

Unio crassus Retzius, 1788

Schröter, 1779. Gesch. Flußconch.: 182, pl. 2, fig. 2 [Mya testa crassa]. [non-binomial]
Philipsson in Retzius, 1788. Diss. Nat.-Hist. Test. Gen.: 17.

Margaritana crassaSimpson (1900), Simpson (1914)
Unio crassusKobelt & Haas (1910), Ortmann (1912), Ortmann (1914), Germain (1931), Zhadin (1965), ... Dai et al. (2025), Wu et al. (2025), Chen et al. (2025), Hou et al. (2025), Wang et al. (2025)
Unio (s.s.) crassusZhadin (1938)
Unio crassus crassusHaas (1940), Haas (1969), Fauna Europaea (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017), Klishko et al. (2017), Bogatov (2022)
Crassiana crassa crassaAkromovski (1976), Kantor & Sysoev (2005), Kantor et al. (2010)
Batavusiana (Crassunio) crassaZatravkin (1983), Stadnichenko (1984), Zatravkin (1987)
Unio (Crassunio) crassusFalkner (1994)
Batavusiana (Crassiana) crassusKorniushin (1998)
Unio (Crassiana) crassus crassusFalkner et al. (2001), Falkner et al. (2002)
Crassiana (s.s.) crassaStarobogatov et al. (2004)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio c. crassusPríe et al. (2021)

Mya pictorum Gmelin, 1791

Lister, 1678. Hist. Anim. Anglic.: pl. 2, fig. 30; [app.] pl. 1, fig. 4. [pre-linnaean]
Lister, 1685. Conch.: pl. 146, fig. 1; pl. 147, figs. 2-3. [pre-linnaean]
Schröter, 1779. Gesch. Flußconch.: pl. 2, fig. 2; pl. 3, figs. 2-5; pl. 4, fig. 6.
specimen image from publicationMartini & Chemnitz, 1782. Conchyl. Cab. 6: 19, pl. 1, fig. 6.
Gmelin, 1791. Syst. Nat. 1(6): 3218, sp. 3.
(misidentified reference to Mya pictorum Linnaeus, 1758)

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)

Unio musivus Spengler, 1793

Spengler, 1793. Skr. Naturhist. Selsk. 3: 67, no. 32.

Unio musivusSimpson (1900)
Unio crassus musivusOrtmann (1912), Bogatov (2022)
Unio (sec. Lymnium) batavusSimpson (1914)
Unio crassus crassusHaas (1940), Haas (1969), Fauna Europaea (2012), Klishko et al. (2017)
Batavusiana (s.s.) musivaZatravkin (1983), Zatravkin (1987)
Batavusiana (s.s.) musivus [??]Korniushin (1998)
Crassiana (Bataviana) musivaStarobogatov et al. (2004)
Crassiana musiva musivaKantor & Sysoev (2005), Kantor et al. (2010)
Unio crassusGraf (2007), Bogatov et al. (2018), Anisimova et al. (2018), Lopes-Lima et al. (2024)
Unio (Crassiana) musivusVinarski & Kantor (2016)
Unio crassus cf. courtillieriKlishko et al. (2017)

Mya batava Maton & Rackett, 1807

Schröter, 1779. Gesch. Flußconch.: pl. 3, figs. 2, 4-5.
Martini & Chemnitz, 1782. Conchyl. Cab. 6: 19, pl. 1, fig. 6.
Bruguière, 1797. Encycl. Méth. 2: pl. 248, fig. 3.
Maton & Rackett, 1807. Trans. Linn. Soc. London 8: 37.

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio (Batavusiana) batavusGermain (1904)
Unio (Groupe des Batavusiana) batavusCaziot (1907)
Unio batavusKobelt & Haas (1910), Haas (1924), Germain (1931)
Unio crassus musivusOrtmann (1912)
Unio crassus batavusHaas (1940), Haas (1969), Fauna Europaea (2012), Lopes-Lima et al. (2017)
Batavusiana (s.s.) musivaZatravkin (1983), Zatravkin (1987)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusAraujo et al. (2018)

Unio lemovicincae Férussac, 1807

‘Linn.’ Draparnaud, 1806. Hist. Moll. France: pl. 11, fig. 5.
Férussac, 1807. Ess. méth.: 114 [as ‘lemovicincæ’].

Unio litoralis Pfeiffer, 1821

Pfeiffer, 1821. Natur. Deutsch. Land Sussw. Moll. 1: 117, pl. 5, fig. 12.
Rossmässler, 1853. Z. Malak. 10: 92.
Mörch, 1863. Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): ??.
(misidentified reference to Unio littoralis Cuvier, 1798; error for Unio littoralis Cuvier, 1798)

Margaritana crassaSimpson (1900), Simpson (1914)
Batavusiana (Crassunio) crassaStadnichenko (1984)
Crassiana crassa crassaKantor & Sysoev (2005), Kantor et al. (2010)

Unio ater Nilsson, 1822

‘Linn.’ Draparnaud, 1806. Hist. Moll. France: pl. 11, fig. 5 [as ‘Unio margaritifera’].
Nilsson, 1822. Hist. Moll. Svec.: 107.

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio (s.s.) aterZhadin (1938)
Unio crassus crassusHaas (1940), Haas (1969)
Batavusiana (s.s.) musivaZatravkin (1987)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio rubens Menke, 1828

Menke, 1828. Syn. Method. Moll.: 64, 90.
Menke, 1830. Syn. Method. Moll.: 107, 149.

Quadrula (sec. Fusconaia) coccineaSimpson (1900), Simpson (1914)
Quadrula (Obliquata) coccineaFrierson (1927)
Unio crassus crassusHaas (1940), Haas (1969)
Pleurobema sintoxiaParmalee & Bogan (1998), Williams et al. (2008)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio rugatus Menke, 1828

Menke, 1828. Syn. Method. Moll.: 64, 90.
Menke, 1830. Syn. Method. Moll.: 107, 149.

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio consentaneus var.Kobelt & Haas (1910)
Unio crassus crassusHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio badius Kokeil, 1831

Kokeil, 1831. Mich. Comp.: pl. 16, fig. 36.

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)

Unio arcuata Bouchard-Chantereaux, 1838

Bouchard-Chantereaux, 1838. Cat. Moll. Terr. Fluv. Départ. Pas-de-Calais: 91, pl. 1, fig. 1-2.
(renamed Unio bouchardi Locard, 1889)

Margaritana auriculariaGermain (1931)
Unio consentaneus var. moquiniGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus moquinianusPrie (2012)

Unio corrugata Mauduyt, 1839

Mauduyt, 1839. Tableau Indic. Moll. Vienne: 8, pl. 1, fig. 1.

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)

Unio gangrenosus J.F. Schmidt, 1840

‘Ziegler’ J.F. Schmidt, 1840. Bull. Soc. imp. Nat. Moscou 13: 436.
J.F. Schmidt, 1847. Conch. Krain: 26.

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio crassus cythereaHaas (1940), Haas (1969)

Unio littoralis var. minor Rossmässler, 1842

Rossmässler, 1842. Icon. Land- und Süssw.-Moll. 2 (5): 14, pl. 55, figs. 744. [in part]
(misidentified reference to Unio littoralis var. minor Rossmässler, 1842)

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio (Tumidusiana) tumidus var. minorStadnichenko (1984)

Unio crassissimus Hanley, 1843

Schlüter, 1838. Kurzgef. Syst. Verz. Conch. Samml.: 33 [as ‘Mysca tumida var. crassissima’]. [nomen nudum]
specimen image from publication‘Fer.’ Hanley, 1843. Illustr. Descr. Cat. Rec. Bivalve Shells: 209.

Margaritana crassaSimpson (1900), Simpson (1914)

Unio manca Millet, 1843

‘Lamarck’ of Authors.
Millet, 1843. Mag. Zool.: 4, pl. 64, fig. 2.
(misidentified reference to Unio manca Lamarck, 1819)

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio (Mancusiana) mancusGermain (1904)
Unio mancusGermain (1931)

Unio batavus var. pusillus Rossmässler, 1844

Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): ??.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio pusillus Lea, 1840)

Unio drouetii Dupuy, 1849

Dupuy, 1849. Cat. Extramar. Galliae Test.: no. 326.
Dupuy, 1852. Hist. Natur. Moll. France (6): 639, pl. 24, fig. 12.

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969), Fauna Europaea (2012)
Unio crassusLopes-Lima et al. (2024)

Unio ventricosus Scholtz, 1853

Scholtz, 1853. unknown publication: ??.
(homonym of Unio ventricosus Barnes, 1823)

Unio capigliolo Küster, 1855

‘Payraudeau’ of Authors.
specimen image from publicationKüster, 1855. Conchyl. Cab. 9(2): 125, pl. 34, figs. 5-6. [in part]
Sowerby, 1868. Conch. Icon. 16: pl. 75, sp. 392.
(misidentified reference to Unio capigliolo Payraudeau, 1826)

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)

Unio gangraenosus Küster, 1855

‘Ziegler’ of Authors.
Küster, 1855. Conchyl. Cab. 9(2): 124.
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 81.
Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 130.
Haas, 1969. Das Tierreich 88: 46.
(error for Unio gangrenosus J.F. Schmidt, 1840)

Unio (Crassiana) crassusVinarski & Kantor (2016)

Unio rivularis Moquin-Tandon, 1855

Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): fig. 414.
Moquin-Tandon, 1855. Hist. Natur. Moll. France 2: 571.
(homonym of Unio rivularis Conrad, 1853)

Unio (Crassiana) crassusVinarski & Kantor (2016)

Unio batavus var. sarratinus Moquin-Tandon, 1855

‘St. Germain, in litt.’ Moquin-Tandon, 1855. Hist. Natur. Moll. France 2: 571.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio courtillieri Hattemann, 1859

Hattemann, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 232.

Unio (Fusculusiana) fusculusGermain (1904)
Unio crassus var. piscinalisGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969), Lopes-Lima et al. (2017)
Unio (Crassiana) crassus courtillieriFalkner et al. (2001), Falkner et al. (2002)
Unio crassus courtillieriPrié & Puillandre (2014), Lopes-Lima et al. (2017)
Unio crassus cf. courtillieriKlishko et al. (2017)
Unio crassus courtillierii [sic]Araujo et al. (2018)
Unio crassusAnisimova et al. (2018), Bogatov et al. (2018), Lopes-Lima et al. (2024)
Unio c. courtillieriPríe et al. (2021)
Unio cf. courtillieriLopes-Lima et al. (2021)

Unio batavus var. acutus de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 287, pl. 10, fig. 1.
(homonym of Unio acutus Rossmässler, 1837)

Unio batavus var. baraceus de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 285, pl. 9, fig. 7.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus var. baraceusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio batavus var. longus de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 288, pl. 10, fig. 3.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio longus Philippi, 1869)

Unio batavusGermain (1931)

Unio batavus var. minimus de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 289, pl. 9, fig. 6.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio requienii var. minima Drouët, 1857)

Unio mancus var. amnicusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio batavus var. senotius de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 288, pl. 10, fig. 4.

Unio sanderi Bourguignat, 1862

Bourguignat, 1862. Malacol. Lac Quatre-Cantons: 55.
Bourguignat, 1864. Malac. Aix-les-Baines: 76.

Unio mancus var. macrorhynchusGermain (1931)

Unio merdiger Reeve, 1865

Jay, 1839. Cat. Rec. Shells: 25, sp. 1065.
‘Walsh.’ Lea, 1852. Synopsis Fam. Naiades: ??. [nomen nudum]
specimen image from publication‘Watts’ Reeve, 1865. Conch. Icon. 16: pl. 28, sp. 145.

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)

Unio lambottei Malzine, 1867

Malzine, 1867. Essai faun. malac. Belg.: 32, pl. 2, figs, 3-4.

Unio (Ateriana) lambotteiGermain (1904)
Unio consentaneusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio ryckholtii Malzine, 1867

Malzine, 1867. Essai faun. malac. Belg.: 32, pl. 2, figs. 1-2.

Unio (Batavusiana) ryckholtiiGermain (1904)
Unio batavusGermain (1931)

Unio batavus var. squamosus Brot, 1867

Brot, 1867. Assoc. Zool. Léman Ann. (1866): pl. 9, fig. 1.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; misidentified reference to Unio batavus var. squamosus Charpentier, 1837)

Unio consentaneusGermain (1931)

Unio ryckholti var. cuneata Malzine, 1867

Malzine, 1867. Essai faun. malac. Belg.: 32.
(homonym of Unio cuneatus Jacquemin, 1835; intro'd as intra-sp. of Unio ryckholtii Malzine, 1867)

Unio batavusGermain (1931)

Unio (tumidus) elegans Westerlund, 1867

Westerlund, 1867. Malak. Blatt. 14: 202, 206.
(homonym of Unio elegans Lea, 1831)

Unio (Crassiana) crassusVinarski & Kantor (2016)

Unio batavus var. belgicus Colbeau, 1868

Colbeau, 1868. Ann. Soc. Malac. Belgique 3: 106, pl. 4, fig. 3.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio crassus var. maximus Kobelt, 1872

Kobelt, 1872. Malak. Blatt. 19: 142, pl. 5.
Kobelt, 1884. Icon. Land- und Süssw.-Moll. 1: 67, pl. 29, figs. 218-219.
(intro'd as intra-sp. of Unio crassus Retzius, 1788; homonym of Unio pictorum maxima Mörch, 1863)

Unio crassus crassusHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio pseudolitoralis Clessin, 1877

Clessin, 1875. Verh. Ver. Naturgesch. Hamburg 2: 258 [as ‘pseudo-litoralis’]. [nomen nudum]
Clessin, 1877. Deutsche Excurs. Moll. Fauna: 469, text fig. 305 [as ‘pseudo-litoralis’].

Unio (s.s.) crassus var. pseudolitoralisZhadin (1938)
Unio crassus crassusHaas (1940), Haas (1969)
Unio crassus var. pseudolitoralisZhadin (1965)
Batavusiana (Crassunio) crassaStadnichenko (1984), Zatravkin (1987)
Crassiana crassa crassaKantor & Sysoev (2005), Kantor et al. (2010)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio subtilis Drouët, 1879

Drouët, 1879. J. de Conch. 27: 142.

Unio (Groupe des Mancusiana) subtilisCaziot (1907)
Unio mancus var. amnicusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio vescoi var. conemenosi Westerlund & Blanc, 1879

Blanc in Westerlund & Blanc, 1879. Aperçu Faune Malac. Grèce: 145.
(intro'd as intra-sp. of Unio vescoi Bourguignat, 1856)

Unio moquinianus Brevière, 1880

Brevière, 1880. Cat. moll. Nièvre: 26.
(misidentified reference to Unio moquinianus Dupuy, 1843)

Unio consentaneusGermain (1931)

Unio requieni Brevière, 1880

‘Mich.’ Brevière, 1880. Cat. moll. Nièvre: 26.
(misidentified reference to Unio requienii Michaud, 1831)

Unio consentaneusGermain (1931)

image of type lot

Unio andegavensis Locard, 1882

‘Servain’ Locard, 1882. Prodr. Malac. Franç. 1: 289, 359.
‘Servain’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 142 [36].
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 167.
Type(s): POSSIBLE SYNTYPE MNHN_22100, La Maine à Angers.

Unio (Batavusiana) andegavensisGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio arenarum Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 289, 358.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 144 [38].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 171.
Type(s): IDIOTYPE MNHN_22110, Basse Indre (Loire-Inf.).

Unio (Batavusiana) arenarumGermain (1904)
Unio batavus var. baraceusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio berthelini Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 290, 359.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 137 [31].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 163.
Type(s): IDIOTYPE MNHN_22132, La Marne, à Charenton (Seine).

Unio mancus var. amnicus form ripariusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio brevierei Locard, 1882

‘Bourguignat’ Locard, 1882. Prodr. Malac. Franç. 1: 286 [as ‘brevieri’], 356.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 148 [42].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 178.
(correct name for Unio brevieri Locard, 1882)
Type(s): LECTOTYPE MNHN_22145 (by Graf, 2011: 465), La Canne, à Goulnot.

Unio (Ateriana) brevieri [sic]Germain (1904)
Unio (Groupe des Ateriana) breviereiCaziot (1907)
Unio consentaneusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio brevieri Locard, 1882

‘Bourguignat in Locard’ of Authors.
Locard, 1882. Prodr. Malac. Franç. 1: 286.
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 265.
Germain, 1931. Faune de France 22: 732.
Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 130.
Haas, 1969. Das Tierreich 88: 49.
(incorrect original spelling of Unio brevierei Locard, 1882)

Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio carantoni Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 295, 364.
‘Coutagne’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 129 [23].
‘Coutagne’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 157.
Type(s): IDIOTYPE MNHN_22162, [illegible] (Charente).

Unio carantoniRochebrune (1903)
Unio mancus var. amnicusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio crassatellus Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 286, 356.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 136 [30].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 162.
Type(s): LECTOTYPE MNHN_22182 (by Graf, 2011: 469), La Canne, à Jailly (Nièvre).

Unio crassatellusRochebrune (1903)
Unio crassusGermain (1931), Lopes-Lima et al. (2024)
Unio crassus batavusHaas (1940), Haas (1969)

image of type lot

Unio cyprinorum Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 289, 358.
‘Berthier’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 142 [36].
‘Berthier’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 168.
Type(s): IDIOTYPE MNHN_22185, l’Ornain (Meuse).

Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Crassiana (Bataviana) cyprinorumStarobogatov et al. (2004)
Crassiana cyprinorumKantor & Sysoev (2005), Kantor et al. (2010)
Unio crassusGraf (2007), Lopes-Lima et al. (2024)
Unio (Crassiana) cyprinorumVinarski & Kantor (2016)
Unio crassus musivusBogatov (2022)

image of type lot

Unio feliciani Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 285, 355.
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 134 [28].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 161, fig. 175.
Bourguignat in Locard, 1899. Conchyliol. Portug.: 217.
Type(s): POSSIBLE SYNTYPE MNHN_22215, La Moselle.

Unio (Socardiana) felicianiGermain (1904)
Unio batavus var. baraceusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio lagnisicus Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 291, 359.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 127 [21] [as ‘lagnysicus’].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 154 [as ‘lagnysicus’].
Type(s): SYNTYPE MNHN_22303, Condal (Saône-et-Loire).

Unio lagnysicus [sic]Rochebrune (1903), Kobelt & Haas (1910)
Unio mancusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus mancusPrie (2012)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio ligericus Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 289, 358.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 142 [36].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 169.
Type(s): IDIOTYPE MNHN_22310, La Loire à Ingrandes.

Unio (Batavusiana) ligericusGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio locardianus Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 287, 356.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 138 [32].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 164, fig. 178.
Type(s): IDIOTYPE MNHN_22316, l’Albane (Côte-d’Or).

Unio crassusGermain (1931), Lopes-Lima et al. (2024)
Unio crassus batavusHaas (1940), Haas (1969)

Unio marcellinus Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 285, 355.
‘Berthier’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 149 [43].
‘Berthier’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 180.

Unio (Ateriana) marcellinusGermain (1904)
Unio consentaneus var. moquiniGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus moquinianusPrie (2012)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio matronicus Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 289, 358.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 143 [37].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 170.
Type(s): LECTOTYPE MNHN_22355 (by Graf, 2011: 489), La Marne à Lagny (Seine-et-Marne).

Unio (Batavusiana) matronicusGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio oxyrhynchus Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 285, 355.
‘Brevière’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 149 [43].
‘Brevière’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 180.
(homonym of Unio oxyrhynchus Martens, 1861)
Type(s): LECTOTYPE MNHN_22438 (by Graf, 2011: 495), La Canne, dans la Nièvre.

Unio consentaneus var. oxyrhynchusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio pilloti Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 291, 360.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 128 [22].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 157.
Type(s): IDIOTYPE MNHN_22460, Passavant (Haute-Saône).

Unio pillotiKobelt & Haas (1910)
Unio mancusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus mancusPrie (2012)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio rayi Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 291, 360.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 127 [21].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 155.
Type(s): LECTOTYPE MNHN_22483 (by Graf, 2011: 500), Gigny (Jura).

Unio (Groupe des Ninusiana) [sic] rayiCaziot (1907)
Unio rayiKobelt & Haas (1910)
Unio mancusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus mancusPrie (2012)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio septentrionalis Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 284, 354.
‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 149 [43].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 181, fig. 194.
Type(s): IDIOTYPE MNHN_22528, le Mandrezey, à Saulcy-sur-Meurthe (Vosges).

Unio consentaneusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)

image of type lot

Unio sequanicus Locard, 1882

Bourguignat in Locard, 1882. Prodr. Malac. Franç. 1: 289, 358.
‘Coutagne’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 142 [36].
‘Coutagne’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 169, fig. 183.
Type(s): LECTOTYPE MNHN_22529 (by Graf, 2011: 504), La Marne à Lagny (Seine-et-Marne).

Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio batavellus Locard, 1885

‘Letourneux’ Locard, 1885. Bull. Soc. Sci. Nat. Rouen 21: 11.
‘Letourneux’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 144 [38].
‘Letourneux’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 170, fig. 184.
Type(s): LECTOTYPE MNHN_22127 (by Graf, 2011: 462), La Bouille, près de Rouen (Seine-Inf.).

Unio batavellusRochebrune (1903), Kobelt & Haas (1910)
Unio (Batavusiana) batavellusGermain (1904)
Unio batavus var. baraceusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio baroni Servain, 1887

Servain, 1887. Bull. Soc. Malac. France 4: 254.

Unio carcasinus Sourbieu, 1887

Sourbieu, 1887. Bull. Soc. Malac. France 4: 235.

Unio crassus var. piscinalisGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)

Unio baudoni Folin, 1888

Folin, 1888. Le Naturaliste 10: 273, 4 text figs.
(homonym of Unio baudoni Folin & Bérillon, 1874)

Unio baudoniKobelt (1902)
Unio crassus batavusHaas (1940), Haas (1969)

image of type lot

Unio crassulus Drouët, 1888

Drouët, 1888. J. de Conch. 36: 106.
Drouët, 1889. Mém. Acad. Sci. Dijon 1: 66, pl. 2, fig. 5.
(homonym of Unio requienii crassulus Drouët, 1867)
Type(s): SYNTYPE MNHN_MP_0027, Saone.

Unio consentaneusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio hamburgiensis Servain, 1888

Servain, 1888. Bull. Soc. Malac. France 5: 315.

Unio crassusGermain (1931), Lopes-Lima et al. (2024)
Unio crassus crassusHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)

Unio minutulus Servain, 1888

‘Ray’ Locard, 1882. Prodr. Malac. Franç. 1: 290 [as ‘Unio minutus’]. [nomen nudum]
‘Ray’ Servain, 1888. Bull. Soc. Malac. France 5: 316.

Unio mancus var. amnicusGermain (1931)
Unio crassus batavusHaas (1940)

Unio vegesackensis Servain, 1888

Bourguignat in Servain, 1888. Bull. Soc. Malac. France 5: 317.

Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio visurgisinus Servain, 1888

Servain, 1888. Bull. Soc. Malac. France 5: 316.

Unio (Batavusiana) visurgicus [sic]Germain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio adonus Locard, 1889

‘Servain’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 146 [40], 211 [105].
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 175, fig. 188.
Type(s): LECTOTYPE MNHN_22085 (by Graf, 2011: 456), La Loire a Basse Indre (Loire-Inf.).

Unio (Lemotheuxiana) adonusGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio andeliacus Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 138 [32], 196 [90].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 165.
Type(s): LECTOTYPE MNHN_22101 (by Graf, 2011: 458), le Gouet, à Saint-Brieue (Côtes-du-Nord).

Unio mancus var. elongatulusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio aturicus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 129 [23], 188 [82].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 157.
Type(s): HOLOTYPE (MONOTYPY) MNHN_22118, Bahus (Landes).

Unio mancus var. amnicus form ripariusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)

image of type lot

Unio balbignyanus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 147 [41], 217 [111].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 178.
Type(s): LECTOTYPE MNHN_22122 (by Graf, 2011: 462), La Loire, à Balbigny (Loire).

Unio consentaneusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio besnardianus Locard, 1889

Servain, 1888. Bull. Soc. Malac. France 5: 317. [nomen nudum]
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 141 [35], 206 [100].
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 172, fig. 186 [as ‘besnardinus’].
(renamed Unio besnardi Germain, 1904)
Type(s): LECTOTYPE MNHN_22133 (by Graf, 2011: 463), La Loire, à Saint-Gemmes (Maine-et-Loire).

Unio (Batavusiana) besnardiGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio bouchardi Locard, 1889

‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 149 [43].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 180.
(new name for Unio arcuata Bouchard-Chantereaux, 1838)
Type(s): LECTOTYPE FIGURED SPECIMEN (by Graf, 2011: 464), ditches of Saint-Omer, fed by the Aa.

Unio consentaneus var. moquiniGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus moquinianusPrie (2012)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio caroliensis Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 159 [53], 237 [131].
‘F. Pacome’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 172.
Type(s): LECTOTYPE MNHN_22167 (by Graf, 2011: 467), Charolles (Saône-et-Loire).

Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio catalaunicus Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 133 [27], 192 [86].
‘Coutagne’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 160.
Type(s): LECTOTYPE MNHN_22171 (by Graf, 2011: 467), La Marne, à Charenton (Seine).

Unio crassus var. piscinalisGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio caumonti Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 145 [39], 207 [101].
Bourguignat in Locard, 1893. Coqu. Eau Douce Saumâtre France: 171.

Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio diptychus Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 143 [37], 201 [95].
‘Surault’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 169.
Type(s): POSSIBLE SYNTYPE MNHN_22192, La Loire à Ingrandes.

Unio (Batavusiana) diptychusGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio financei Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 143 [37], 205 [99].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 171.
Type(s): IDIOTYPE MNHN_22217, La Marne à Châlons (Marne).

Unio batavusGermain (1931)

image of type lot

Unio hattemani Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 146 [40], 212 [106].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 176.
Type(s): SYNTYPE MNHN_22254, La Loire, à Basse-Indre.

Unio (Lemotheuxiana) hattemanni [sic]Germain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio ignariformis Locard, 1889

‘Dup.’ Drouët, 1857. Étud. Naïades. France, Unio: 100, pl. 9, fig. 1 [as ‘U. platyrhynchoideus’].
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 147 [41], 214 [108].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 177.

Unio consentaneusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)

image of type lot

Unio ingrandiensis Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 143 [37], 202 [96].
‘Sarault’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 173.
Type(s): POSSIBLE SYNTYPE MNHN_22282, Ingrandes (Maine-et-Loire).

Unio (Batavusiana) ingrandiensisGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio lagnysicus Locard, 1889

‘Bourguignat’ of Authors.
Locard, 1889. Contr. Faune Malacol. Franc. 13: 127 [21].
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 84.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 154.
Dumas, 1895. Rev. Sci. Bourbonnais, Suppl.: 47.
Rochebrune, 1903. Bull. Mus. Hist. Nat. 9: 389.
Kobelt & Haas, 1910. Icon. Land- und Süssw.-Moll. 17: 37.
(error for Unio lagnisicus Locard, 1882)

image of type lot

Unio lemotheuxi Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 145 [39], 208 [102].
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 175, fig. 189.
Type(s): LECTOTYPE MNHN_22306 (by Graf, 2011: 484), le Reyran (Var).

Unio (Lemotheuxiana) lemotheuxiGermain (1904), Caziot (1910)
Unio mancus var. elongatulusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio mariae Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 159 [53], 236 [130] [as ‘mariæ’].
‘F. Pacome’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 172 [as ‘mariæ’].
Type(s): IDIOTYPE MNHN_22342, La Loire à Nantes (Loire-Infereur).

Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio materniacus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 143 [37], 203 [97].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 168, fig. 182.
Type(s): LECTOTYPE MNHN_22347 (by Graf, 2011: 488), la Somme à Abbeville.

Unio materniacusRochebrune (1903)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio melantatus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 147 [41], 216 [110].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 177.
Type(s): LECTOTYPE MNHN_22360 (by Graf, 2011: 489), La Loire à Balbigny (Loire).

Unio consentaneusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio nicolloni Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 141 [35], 200 [94].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 173.
Type(s): HOLOTYPE (MONOTYPY) MNHN_22404, Nantes.

Unio (Batavusiana) nicolloniGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio occidentalis Locard, 1889

‘Bourguignat’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 149 [43], 220 [114].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 180.
‘Bourguignat’ Locard, 1899. Conchyliol. Portug.: 220.
(homonym of Unio occidentalis Conrad, 1836)
Type(s): LECTOTYPE MNHN_22424 (by Graf, 2011: 494), Balbigny (Loire).

Unio consentaneusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio seneauxi Locard, 1889

‘Linn.’ Draparnaud, 1806. Hist. Moll. France: 131, pl. 11, figs. 3 [as ‘Unio pictorum var. B’].
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 145 [39], 208 [102].
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174, fig. 187 [as ‘senauxi’].
Type(s): LECTOTYPE MNHN_22523 (by Graf, 2011: 503), La Loire à Orléans.

Unio (Batavusiana) seneauxiGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio subamnicus Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 137 [31], 194 [88].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 163.
Type(s): SYNTYPE MNHN_22559, La Saône, à Auxonne (Côte-d’Or).

Unio subamnicusKobelt & Haas (1910)
Unio mancus var. amnicusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio surraulti Locard, 1889

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 144 [38], 204 [98].
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174 [as ‘suraulti’].
Type(s): SYNTYPE MNHN_22569, La Loire à Ingrandes.

Unio (Batavusiana) surraultiGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio visurgicus Locard, 1889

‘Servain’ of Authors.
Locard, 1889. Contr. Faune Malacol. Franc. 13: 141 [35].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 173.
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 262.
(error for Unio visurgisinus Servain, 1888)

Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio subrobustus var. curta Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 135 [29].
(homonym of Unio curtus Lea, 1859; intro'd as intra-sp. of Unio subrobustus Servain, 1885)

Unio sieversi var. kobelti Westerlund, 1890

Kobelt, 1886. Icon. Land- und Süssw.-Moll. 2: pl. 53, fig. 286 [as ‘U. sieversi’].
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 151.
(intro'd as intra-sp. of Unio sieversi Drouët, 1881; homonym of Unio kobelti Neumayr, 1899)

image of type lot

Unio ampullaceus Locard, 1893

Locard, 1893. Coqu. Eau Douce Saumâtre France: 174.
(homonym of Unio ampullaceus Lea, 1866)
Type(s): LECTOTYPE MNHN_22098 (by Graf, 2011: 458), Le Mans (Sarthe).

Unio (Batavusiana) ampullaceusGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio chardoni Locard, 1893

‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 173.

Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

image of type lot

Unio gliscerus Locard, 1893

Locard, 1893. Coqu. Eau Douce Saumâtre France: 168.
Type(s): LECTOTYPE MNHN_22241 (by Graf, 2011: 477), La Loire à Nantes (Loire-Infereur).

Unio (Batavusiana) gliscerusGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio lamboltei Locard, 1893

‘Malzine’ of Authors.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 177.
Germain, 1931. Faune de France 22: 732.
Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 130.
(error for Unio lambottei Malzine, 1867)

image of type lot

Unio lancelevei Locard, 1893

Locard, 1893. Bull. Soc. Sci. Nat. Elbeuf 12: 1.
Locard, 1893. Coqu. Eau Douce Saumâtre France: 170.
Type(s): SYNTYPE MNHN_22305, le Seine au nord d. Rouen.

Unio batavus var. baraceusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio lepidulus Drouët, 1898

Drouët, 1898. Mém. Acad. Sci. Dijon ??: 60, pl. 1, fig. 7.

Unio mancus var. elongatulusGermain (1931)

Unio batavus var. major Drouët, 1898

Drouët, 1898. Mém. Acad. Sci. Dijon ??: 61.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio (Anodontes) sirionos var. major d’Orbigny, 1835)

Unio batavus var. baraceusGermain (1931)

Unio ellipsopsis Locard, 1899

Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 140 [34]. [nomen nudum]
Bourguignat in Locard, 1899. Conchyliol. Portug.: 218.

Unio besnardi Germain, 1904

Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 261 [23].
(new name for Unio besnardianus Locard, 1889)

Unio (Batavusiana) besnardiGermain (1904)

Unio ater var. irgizlaicus Lindholm, 1904

Lindholm, 1904. Ann. Mus. St. Pétersbourg 8(3-4): 343.
(intro'd as intra-sp. of Unio ater Nilsson, 1822)

Unio crassus crassusHaas (1969), Klishko et al. (2017)
Batavusiana (s.s.) irgislaica [sic]Zatravkin (1987)
Crassiana (Bataviana) irgizlaicaStarobogatov et al. (2004)
Crassiana irgizlaicaKantor & Sysoev (2005), Kantor et al. (2010)
Unio crassusGraf (2007), Lopes-Lima et al. (2024)
Unio (Crassiana) irgizlaicusVinarski & Kantor (2016)
Unio crassus musivusBogatov (2022)

Unio batavellus var. perconvexus Germain, 1904

Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 260 [22].
(intro'd as intra-sp. of Unio batavellus Locard, 1885)

Unio (Batavusiana) batavellus perconvexusGermain (1904)

Unio batavus var. radiata Germain, 1904

Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 257 [19].
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio (Elliptio) viridis var. radiata Rafinesque, 1820)

Unio (Batavusiana) batavus radiataGermain (1904)

Unio ligericus var. subsinuatus Germain, 1904

Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 259 [21].
(intro'd as intra-sp. of Unio ligericus Locard, 1882)

Unio (Batavusiana) ligericus subsinuatusGermain (1904)

Unio pseudolittoralis [sic] var. curonicus Riemenschneider, 1907

Riemenschneider, 1907. Nachr. Bl. Dtsch. Malak. Ges. 39: 150.
(intro'd as intra-sp. of Unio pseudolitoralis Clessin, 1877)

Unio crassus crassusHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio crassus var. okae Kobelt & Haas, 1910

Kobelt & Haas, 1910. Icon. Land- und Süssw.-Moll. 17: 58, index, pl. 480, fig. 2567.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Unio (s.s.) crassus var. okaeZhadin (1938)
Unio crassus gontieriHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio crassus heimburgi Haas, 1911

Haas, 1911. Verh. Naturhist. Ver. Preuß. Rheinl. 68: 527, pl. 6, fig. 20.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Unio crassus crassusHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio kungurensis Kobelt, 1912

Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 24, pl. 490, figs. 2622-2623.

Unio (s.s.) crassus kungurensisZhadin (1938)
Unio crassus gontieriHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio roseni Kobelt, 1912

Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 27, pl. 492, fig. 2627.

Unio (s.s.) crassus var. roseniZhadin (1938)
Unio crassus gontieriHaas (1940), Haas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio kungurensis chlebnikowi Kobelt, 1912

Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 26, pl. 491, fig. 2624.
(intro'd as intra-sp. of Unio kungurensis Kobelt, 1912; renamed Unio crassus var. chlebnikovi Zhadin, 1938)

Unio (s.s.) crassus var. chlebnikovi [sic]Zhadin (1938)
Unio crassus gontieriHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio kungurensis irenjensis Kobelt, 1912

Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 26, pl. 491, fig. 2626.
(intro'd as intra-sp. of Unio kungurensis Kobelt, 1912)

Unio (s.s.) crassus var. chlebnikovi [sic]Zhadin (1938)
Unio crassus gontieriHaas (1940), Haas (1969), Graf (2007)
Batavusiana (s.s.) irenjensisZatravkin (1983), Stadnichenko (1984), Zatravkin (1987)
Crassiana irenjensisKantor & Sysoev (2005), Kantor et al. (2010)
Unio (Crassiana) irenjensisVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio kungurensis sylwensis Kobelt, 1912

Kobelt, 1912. Icon. Land- und Süssw.-Moll. 18: 26, pl. 491, fig. 2625.
(intro'd as intra-sp. of Unio kungurensis Kobelt, 1912)

Unio (s.s.) crassus var. chlebnikovi [sic]Zhadin (1938)
Unio crassus gontieriHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio crassus thuringiacus Israel, 1913

Israel, 1913. [in] Thüring. Lehrerverein Naturk.: 28, pl. 6, figs. 4-6.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Unio crassus crassusHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio crassus ornatus Polinksi, 1917

Polinksi, 1917. C. R. Soc. Sci. Varsovie 10: 904, pl. ??, fig. 2.
(intro'd as intra-sp. of Unio crassus Retzius, 1788; homonym of Unio ovatus var. ornatus Conrad, 1835)

Unio crassus crassusHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio crassus polonicus Polinksi, 1917

Polinksi, 1917. C. R. Soc. Sci. Varsovie 10: 902, pl. ??, figs. 1, 3.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Unio crassus crassusHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio cantianus Kennard & Woodward, 1924

Kennard & Woodward, 1924. Proc. Malac. Soc. London 16: 136, text figs. 8-9.

Unio crassus batavusHaas (1940), Haas (1969)

Unio lagnysiacus Germain, 1931

‘Bourguignat in Locard’ of Authors.
Germain, 1931. Faune de France 22: 724.
Prie, 2012. MalaCo 8: 430.
(error for Unio lagnisicus Locard, 1882)

Unio crassus var. chlebnikovi Zhadin, 1938

‘Kobelt’ Zhadin, 1938. Faune USSR Mollusques 4(1): 89.
(new name for Unio kungurensis chlebnikowi Kobelt, 1912)

Unio crassus var. ishmensis Zhadin, 1938

Zhadin, 1938. Faune USSR Mollusques 4(1): 160.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Unio (s.s.) crassus var. ishmensisZhadin (1938)
Unio crassus crassusHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusLopes-Lima et al. (2024)

Unio pseudolittoralis Riemenschneider, 1907

‘Clessin’ of Authors.
Riemenschneider, 1907. Nachr. Bl. Dtsch. Malak. Ges. 39: 150.
Vinarski & Kantor, 2016. Analyt. Cat. Fr. Brack. Moll. Russia: 25.
(error for Unio pseudolitoralis Clessin, 1877)

Unio courtilieri Locard, 1889

‘Hattemann’ of Authors.
Locard, 1889. Contr. Faune Malacol. Franc. 13: 132 [26].
‘Hattemann’ Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 256 [18].
‘Hattmann’ Germain, 1931. Faune de France 22: 724, 727.
Lopes-Lima et al., 2024. MPE 195: suppl. append. 2, p. 3.
(error for Unio courtillieri Hattemann, 1859)

Unio coactatus of Authors, unknown year

Ziégl. in sched. of Authors.

Unio inflatus of Authors, unknown year

Locard 1881 of Authors.

Unio margaritifera Draparnaud, 1806

‘Linn.’ Draparnaud, 1806. Hist. Moll. France: pl. 11, fig. 5.
(misidentified reference to Mya margaritifera Linnaeus, 1758)

Unio consentaneusGermain (1931)

Unio pictorum Draparnaud, 1806

‘Linn.’ Draparnaud, 1806. Hist. Moll. France: 131, pl. 11, figs. 3 [var. B]. [in part]
(misidentified reference to Mya pictorum Linnaeus, 1758)

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio (Batavusiana) seneauxiGermain (1904)
Unio batavusGermain (1931)

Unio batavus Nilsson, 1822

‘Lamarck’ Nilsson, 1822. Hist. Moll. Svec.: 112.
(reference to Mya batava Maton & Rackett, 1807)

Unio crassus Nilsson, 1822

‘Retzius’ Nilsson, 1822. Hist. Moll. Svec.: 108.
(reference to Unio crassus Retzius, 1788)

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)

Unio lemovicensis Kickx, 1830

‘Daud.’ Kickx, 1830. Synopsin Molluscorum Brabantiae: 82.
(error for Unio lemovicincae Férussac, 1807)

Unio rubens Menke, 1830

Menke, 1830. Syn. Method. Moll.: 149.
(reference to Unio rubens Menke, 1828)

Unio (sec. Lymnium) littoralisSimpson (1900), Simpson (1914)

Unio fuscus Lea, 1836

‘Zieg.’ Lea, 1836. Synopsis Fam. Naiades: 27.
(intro'd as syn of Mya batava Maton & Rackett, 1807)

Unio gibba Lea, 1836

‘Mühl. Pfeif.’ Lea, 1836. Synopsis Fam. Naiades: 27.
(intro'd as syn of Mya batava Maton & Rackett, 1807)

Unio acutus Rossmässler, 1837

Rossmässler, 1837. Icon. Land- und Süssw.-Moll. 1 (5-6): 56, pl 29, fig. 413 [as ‘Unio an n. sp.?’]. [nomen nudum]

Mysca batava var. fuscula subvar. pallipes Schlüter, 1838

‘Ziegl.’ Schlüter, 1838. Kurzgef. Syst. Verz. Conch. Samml.: 33. [infra-subsp.]
(intro'd as intra-sp. of Unio fusculus Rossmässler, 1836)

Mysca reniformis subvar. kokeilli Schlüter, 1838

Schlüter, 1838. Kurzgef. Syst. Verz. Conch. Samml.: 32. [infra-subsp.]
(intro'd as intra-sp. of Unio reniformis Rossmässler, 1836)

Unio gargottae Rossmässler, 1838

Rossmässler, 1838. Icon. Land- und Süssw.-Moll. 2 (1-2): 26, pl. 35, fig. 493.
(misidentified reference to Unio gargottae Philippi, 1836)

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)

Unio requieni Rossmässler, 1844

‘Mich.’ Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): 29, pl. 57, fig. 757-761.
(misidentified reference to Unio requienii Michaud, 1831)

Unio (sec. Lymnium) batavusSimpson (1900)
Unio (sec. Lymnium) batavus var. requieniSimpson (1914)

Unio glaucinus Stabile, 1845

Stabile, 1845. Faun. Lug.: f. 75.
(misidentified reference to Unio glaucinus Porro, 1838)

Unio (sec. Lymnium) batavusSimpson (1900)

Unio piscinalis J.F. Schmidt, 1847

J.F. Schmidt, 1847. Conch. Krain: 27.
(misidentified reference to Unio piscinalis Rossmässler, 1836)

Unio retzii C.B. Adams, 1847

‘Küst.’ C.B. Adams, 1847. Cat. Gen. Sp. Adams: 14. [nomen nudum]

Unio moquinianus Sandberger & Koch, 1851

Sandberger & Koch, 1851. Jb. Nassau. Ver. Naturk. 7: 245.
(misidentified reference to Unio moquinianus Dupuy, 1843)

Unio argens Lea, 1852

‘Küst.’ Lea, 1852. Synopsis Fam. Naiades: 30.
(intro'd as syn of Mya batava Maton & Rackett, 1807)

Unio drouatii Lea, 1852

‘Dup.’ Lea, 1852. Synopsis Fam. Naiades: 39.
(error for Unio drouetii Dupuy, 1849)

Unio kurrii Lea, 1852

‘Küst.’ Lea, 1852. Synopsis Fam. Naiades: 30.
(intro'd as syn of Mya batava Maton & Rackett, 1807)

Unio moquinianus Drouët, 1852

Drouët, 1852. Etudes Nayades de la France: pl. 6, fig. 3.
(misidentified reference to Unio moquinianus Dupuy, 1843)

Unio consentaneusGermain (1931)

Unio nanus Dupuy, 1852

‘Lamarck’ Dupuy, 1852. Hist. Natur. Moll. France (6): pl. 25, fig. 16.
(misidentified reference to Unio nana Lamarck, 1819)

Unio (sec. Lymnium) batavusSimpson (1900)
Unio (Rhomboïdiana) nanaGermain (1904)
Unio nanaKobelt & Haas (1910)
Unio (sec. Lymnium) batavus var. requieniSimpson (1914)
Unio mancus var. nanusGermain (1931)

Unio ovalis Dupuy, 1852

Dupuy, 1852. Hist. Natur. Moll. France (6): pl. 25, fig. 13.
(misidentified reference to Mya ovalis Montagu, 1803)

Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio (Batavusiana) besnardiGermain (1904)
Unio batavusGermain (1931)

Unio crassus Küster, 1854

specimen image from publication‘Retz.’ Küster, 1854. Conchyl. Cab. 9(2): 113, pl. 31, fig. 5. [in part]
(reference to Unio crassus Retzius, 1788)

Lampsilis (sec. Eurynia) radiatusSimpson (1900)
Lampsilis (Eurynia) radiataSimpson (1914)

Unio ater Moquin-Tandon, 1855

‘Nilss.’ Moquin-Tandon, 1855. Hist. Natur. Moll. France 2: 570.
(reference to Unio ater Nilsson, 1822)

Unio ater Drouët, 1857

‘Nilss.’ Drouët, 1857. Étud. Naïades. France, Unio: 72, pl. 4, fig. 1.
(reference to Unio ater Nilsson, 1822)

Unio consentaneusGermain (1931)

Unio nanus Drouët, 1857

‘Lam.’ Drouët, 1857. Étud. Naïades. France, Unio: pl. 5, fig. 2.
(misidentified reference to Unio nana Lamarck, 1819)

Unio (Rhomboïdiana) nanaGermain (1904)
Unio nanaKobelt & Haas (1910)
Unio mancus var. nanusGermain (1931)

Unio platyrhynchoideus Drouët, 1857

‘Dup.’ Drouët, 1857. Étud. Naïades. France, Unio: 100, pl. 9, fig. 1.
(misidentified reference to Unio platyrinchoideus Dupuy, 1849)

Unio batavus de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 2. [in part]
(reference to Mya batava Maton & Rackett, 1807)

Unio (Batavusiana) batavus radiataGermain (1904)

Unio batavus var. courtilieri de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 3.
(reference to Unio courtillieri Hattemann, 1859; error for Unio courtillieri Hattemann, 1859)

Unio (Batavusiana) batavus courtilieriGermain (1904)
Unio batavusGermain (1931)

Unio batavus var. nanus de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 8. [in part]
(misidentified reference to Unio nana Lamarck, 1819)

Unio (Melasiana) melasGermain (1904)
Unio mancus var. amnicusGermain (1931)

Unio batavus var. nanus de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 10. [in part]
(misidentified reference to Unio nana Lamarck, 1819)

Unio (Rhomboïdiana) nanaGermain (1904)

Unio batavus var. ovalis de Joannis, 1859

de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 10, fig. 2.
(misidentified reference to Mya ovalis Montagu, 1803)

Unio (Batavusiana) ligericusGermain (1904)
Unio batavusGermain (1931)

Unio orientalis Küster, 1862

‘Parreis in sched.’ Küster, 1862. Conchyl. Cab. 9(2): 267.
(intro'd as syn of Unio kotschyi Küster, 1862)

Unio ater Bourguignat, 1864

Bourguignat, 1864. Malac. Aix-les-Baines: 73.
(reference to Unio ater Nilsson, 1822)

Unio mancus var. macrorhynchusGermain (1931)

Unio batavus var. fontainei Colbeau, 1868

‘Lecomte’ Colbeau, 1868. Ann. Soc. Malac. Belgique 3: 106. [nomen nudum]
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio maceerthyanus Lea, 1870

‘Bourg.’ Lea, 1870. Synopsis Fam. Naiades: 48.
(error for Unio maccarthyanus Bourguignat, 1866; misidentified reference to Unio maccarthyanus Bourguignat, 1866)

Unio batavus Miller, 1873

Miller, 1873. Schalth. Bodensees: pl. 2, fig. 8.
(reference to Mya batava Maton & Rackett, 1807)

Unio crassusGermain (1931)

Unio diagonalis Drouët, 1881

‘Held’ Drouët, 1881. J. de Conch. 29: 27.
(intro'd as syn of Unio batavus var. squamosus Charpentier, 1837)

Unio minutus Locard, 1882

‘Ray’ Locard, 1882. Prodr. Malac. Franç. 1: 290. [nomen nudum]

Unio mancus var. amnicusGermain (1931)
Unio crassus batavusHaas (1969)
Unio crassusLopes-Lima et al. (2024)

Unio sandrii Locard, 1882

Locard, 1882. Prodr. Malac. Franç. 1: 292.
(misidentified reference to Unio sandrii Rossmässler, 1844)

Unio hueti Kobelt, 1886

‘Bourg.’ Kobelt, 1886. Icon. Land- und Süssw.-Moll. 2: 22, pl. 41, fig. 255.
(misidentified reference to Unio hueti Bourguignat, 1855)

Unio (sec. Lymnium) pseudonymusSimpson (1900), Simpson (1914)

Unio sieversi Kobelt, 1886

Kobelt, 1886. Icon. Land- und Süssw.-Moll. 2: pl. 53, fig. 286.
(misidentified reference to Unio sieversi Drouët, 1881)

Unio dilophius Servain, 1888

‘Bourguignat’ Servain, 1888. Bull. Soc. Malac. France 5: 316, footnote. [nomen nudum]
Locard, 1889. Contr. Faune Malacol. Franc. 13: 136 [30], footnote. [nomen nudum]

Unio sperchinus Locard, 1889

‘Thièse’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 141 [35], footnote. [nomen nudum]

Unio pilloti var. minor Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 129 [23]. [nomen nudum]
(intro'd as intra-sp. of Unio pilloti Locard, 1882)

Unio suborbicularis var. minor Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 127 [21]. [nomen nudum]
Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 84. [nomen nudum]
(intro'd as intra-sp. of Unio suborbicularis Drouët, 1888)

Unio besnardinus Locard, 1893

‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 172.
(error for Unio besnardianus Locard, 1889)

Unio (Batavusiana) besnardiGermain (1904)

Unio senauxi Locard, 1893

‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174.
(error for Unio seneauxi Locard, 1889)

Unio suraulti Locard, 1893

‘Servain in Locard’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174.
(error for Unio surraulti Locard, 1889)

Unio musicus Simpson, 1900

‘Spengler’ Simpson, 1900. Proc. U.S. Nat. Mus. 22: 744.
(error for Unio musivus Spengler, 1793)

Unio (sec. Lymnium) batavusSimpson (1914)

Unio hattemanni Germain, 1904

‘Bourguignat’ Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 265 [27].
(error for Unio hattemani Locard, 1889)

Unio carioliensis Germain, 1931

‘Pacôme’ Germain, 1931. Faune de France 22: 728.
(error for Unio caroliensis Locard, 1889)

Unio visursisinus Germain, 1931

‘Servain’ Germain, 1931. Faune de France 22: 728.
(error for Unio visurgisinus Servain, 1888)

Unio hattmanni Haas, 1940

‘Bourguignat’ Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 131.
(error for Unio hattemani Locard, 1889)

Unio kungurensis sylvensis Haas, 1940

‘Kobelt’ Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 132.
‘Kobelt’ Haas, 1969. Das Tierreich 88: 53.
Vinarski & Kantor, 2016. Analyt. Cat. Fr. Brack. Moll. Russia: 25.
(error for Unio kungurensis sylwensis Kobelt, 1912)

Unio lancelevi Haas, 1969

‘Locard’ Haas, 1969. Das Tierreich 88: 51.
(error for Unio lancelevei Locard, 1893)

Unio crassus var. irgislaicus Zatravkin, 1987

‘Lindholm’ Zatravkin, 1987. Byull. Mosk. Obshch.. Ispyt. Prir. 92(4): 57.
(error for Unio ater var. irgizlaicus Lindholm, 1904)

Taxonomic History (since 1900)

Lampsilis (sec. Eurynia) radiatusSimpson (1900)
Quadrula (sec. Fusconaia) coccineaSimpson (1900), Simpson (1914)
Margaritana crassaSimpson (1900), Simpson (1914)
Unio (sec. Lymnium) littoralisSimpson (1900), Simpson (1914)
Unio (sec. Lymnium) pseudonymusSimpson (1900), Simpson (1914)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio musivusSimpson (1900)
Unio baudoniKobelt (1902)
Unio lagnysicus [sic]Rochebrune (1903), Kobelt & Haas (1910)
Unio carantoniRochebrune (1903)
Unio crassatellusRochebrune (1903)
Unio materniacusRochebrune (1903)
Unio batavellusRochebrune (1903), Kobelt & Haas (1910)
Unio (Rhomboïdiana) nanaGermain (1904)
Unio (Melasiana) melasGermain (1904)
Unio (Mancusiana) mancusGermain (1904)
Unio (Fusculusiana) fusculusGermain (1904)
Unio (Socardiana) felicianiGermain (1904)
Unio (Batavusiana) batavusGermain (1904)
Unio (Batavusiana) batavus radiataGermain (1904)
Unio (Batavusiana) batavus courtilieriGermain (1904)
Unio (Batavusiana) andegavensisGermain (1904)
Unio (Batavusiana) diptychusGermain (1904)
Unio (Batavusiana) ligericusGermain (1904)
Unio (Batavusiana) ligericus subsinuatusGermain (1904)
Unio (Batavusiana) batavellusGermain (1904)
Unio (Batavusiana) batavellus perconvexusGermain (1904)
Unio (Batavusiana) arenarumGermain (1904)
Unio (Batavusiana) besnardiGermain (1904)
Unio (Batavusiana) ryckholtiiGermain (1904)
Unio (Batavusiana) visurgicus [sic]Germain (1904)
Unio (Batavusiana) ingrandiensisGermain (1904)
Unio (Batavusiana) seneauxiGermain (1904)
Unio (Batavusiana) surraultiGermain (1904)
Unio (Batavusiana) gliscerusGermain (1904)
Unio (Batavusiana) matronicusGermain (1904)
Unio (Batavusiana) nicolloniGermain (1904)
Unio (Batavusiana) ampullaceusGermain (1904)
Unio (Lemotheuxiana) adonusGermain (1904)
Unio (Lemotheuxiana) lemotheuxiGermain (1904), Caziot (1910)
Unio (Lemotheuxiana) hattemanni [sic]Germain (1904)
Unio (Ateriana) lambotteiGermain (1904)
Unio (Ateriana) brevieri [sic]Germain (1904)
Unio (Ateriana) marcellinusGermain (1904)
Unio (Groupe des Ninusiana) [sic] rayiCaziot (1907)
Unio (Groupe des Mancusiana) subtilisCaziot (1907)
Unio (Groupe des Batavusiana) batavusCaziot (1907)
Unio (Groupe des Ateriana) breviereiCaziot (1907)
Unio subamnicusKobelt & Haas (1910)
Unio consentaneus var.Kobelt & Haas (1910)
Unio nanaKobelt & Haas (1910)
Unio rayiKobelt & Haas (1910)
Unio pillotiKobelt & Haas (1910)
Unio batavusKobelt & Haas (1910), Haas (1924), Germain (1931)
Unio crassusKobelt & Haas (1910), Ortmann (1912), Ortmann (1914), Germain (1931), Zhadin (1965), ... Dai et al. (2025), Wu et al. (2025), Chen et al. (2025), Hou et al. (2025), Wang et al. (2025)
Unio crassus musivusOrtmann (1912), Bogatov (2022)
Lampsilis (Eurynia) radiataSimpson (1914)
Unio (sec. Lymnium) batavus var. requieniSimpson (1914)
Quadrula (Obliquata) coccineaFrierson (1927)
Margaritana auriculariaGermain (1931)
Unio crassus var. piscinalisGermain (1931)
Unio mancusGermain (1931)
Unio mancus var. macrorhynchusGermain (1931)
Unio mancus var. nanusGermain (1931)
Unio mancus var. amnicusGermain (1931)
Unio mancus var. amnicus form ripariusGermain (1931)
Unio mancus var. elongatulusGermain (1931)
Unio batavus var. baraceusGermain (1931)
Unio consentaneusGermain (1931)
Unio consentaneus var. oxyrhynchusGermain (1931)
Unio consentaneus var. moquiniGermain (1931)
Unio (s.s.) crassusZhadin (1938)
Unio (s.s.) crassus var. pseudolitoralisZhadin (1938)
Unio (s.s.) crassus kungurensisZhadin (1938)
Unio (s.s.) crassus var. chlebnikovi [sic]Zhadin (1938)
Unio (s.s.) crassus var. okaeZhadin (1938)
Unio (s.s.) crassus var. ishmensisZhadin (1938)
Unio (s.s.) crassus var. roseniZhadin (1938)
Unio (s.s.) aterZhadin (1938)
Unio crassus crassusHaas (1940), Haas (1969), Fauna Europaea (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017), Klishko et al. (2017), Bogatov (2022)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio crassus batavusHaas (1940), Haas (1969), Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio crassus gontieriHaas (1940), Haas (1969), Graf (2007)
Unio crassus var. pseudolitoralisZhadin (1965)
Crassiana crassa crassaAkromovski (1976), Kantor & Sysoev (2005), Kantor et al. (2010)
Batavusiana (s.s.) musivaZatravkin (1983), Zatravkin (1987)
Batavusiana (s.s.) irenjensisZatravkin (1983), Stadnichenko (1984), Zatravkin (1987)
Batavusiana (Crassunio) crassaZatravkin (1983), Stadnichenko (1984), Zatravkin (1987)
Unio (Tumidusiana) tumidus var. minorStadnichenko (1984)
Batavusiana (s.s.) irgislaica [sic]Zatravkin (1987)
Unio (Crassunio) crassusFalkner (1994)
Pleurobema sintoxiaParmalee & Bogan (1998), Williams et al. (2008)
Batavusiana (s.s.) musivus [??]Korniushin (1998)
Batavusiana (Crassiana) crassusKorniushin (1998)
Unio (Crassiana) crassus crassusFalkner et al. (2001), Falkner et al. (2002)
Unio (Crassiana) crassus courtillieriFalkner et al. (2001), Falkner et al. (2002)
Crassiana (s.s.) crassaStarobogatov et al. (2004)
Crassiana (Bataviana) irgizlaicaStarobogatov et al. (2004)
Crassiana (Bataviana) musivaStarobogatov et al. (2004)
Crassiana (Bataviana) cyprinorumStarobogatov et al. (2004)
Crassiana cyprinorumKantor & Sysoev (2005), Kantor et al. (2010)
Crassiana irenjensisKantor & Sysoev (2005), Kantor et al. (2010)
Crassiana irgizlaicaKantor & Sysoev (2005), Kantor et al. (2010)
Crassiana musiva musivaKantor & Sysoev (2005), Kantor et al. (2010)
Unio mancus mancusPrie (2012)
Unio mancus moquinianusPrie (2012)
Unio crassus courtillieriPrié & Puillandre (2014), Lopes-Lima et al. (2017)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio (Crassiana) cyprinorumVinarski & Kantor (2016)
Unio (Crassiana) irenjensisVinarski & Kantor (2016)
Unio (Crassiana) irgizlaicusVinarski & Kantor (2016)
Unio (Crassiana) musivusVinarski & Kantor (2016)
Unio crassus cf. courtillieriKlishko et al. (2017)
Unio crassus courtillierii [sic]Araujo et al. (2018)
Unio c. crassusPríe et al. (2021)
Unio c. courtillieriPríe et al. (2021)
Unio cf. courtillieriLopes-Lima et al. (2021)

 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."