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Data last updated
Mon 1 Dec 2025
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Bivalvia
| Unionoida
| Unionoidea
| Unionidae
| Unioninae
| Unionini
| Unio
species Unio crassus Philipsson in Retzius, 1788: synonymy98 specimens + 264 Genbank vouchers
Synonymy
Schröter, 1779. Gesch. Flußconch.: 182, pl. 2, fig. 2 [Mya testa crassa]. [non-binomial] Philipsson in Retzius, 1788. Diss. Nat.-Hist. Test. Gen.: 17.
Margaritana crassa — Simpson (1900), Simpson (1914) Unio crassus — Kobelt & Haas (1910), Ortmann (1912), Ortmann (1914), Germain (1931), Zhadin (1965), ... Dai et al. (2025), Wu et al. (2025), Chen et al. (2025), Hou et al. (2025), Wang et al. (2025) Unio (s.s.) crassus — Zhadin (1938) Unio crassus crassus — Haas (1940), Haas (1969), Fauna Europaea (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017), Klishko et al. (2017), Bogatov (2022) Crassiana crassa crassa — Akromovski (1976), Kantor & Sysoev (2005), Kantor et al. (2010) Batavusiana (Crassunio) crassa — Zatravkin (1983), Stadnichenko (1984), Zatravkin (1987) Unio (Crassunio) crassus — Falkner (1994) Batavusiana (Crassiana) crassus — Korniushin (1998) Unio (Crassiana) crassus crassus — Falkner et al. (2001), Falkner et al. (2002) Crassiana (s.s.) crassa — Starobogatov et al. (2004) Unio (Crassiana) crassus — Vinarski & Kantor (2016) Unio c. crassus — Príe et al. (2021)
Lister, 1678. Hist. Anim. Anglic.: pl. 2, fig. 30; [app.] pl. 1, fig. 4. [pre-linnaean] Lister, 1685. Conch.: pl. 146, fig. 1; pl. 147, figs. 2-3. [pre-linnaean] Schröter, 1779. Gesch. Flußconch.: pl. 2, fig. 2; pl. 3, figs. 2-5; pl. 4, fig. 6.
Martini & Chemnitz, 1782. Conchyl. Cab. 6: 19, pl. 1, fig. 6. Gmelin, 1791. Syst. Nat. 1(6): 3218, sp. 3. (misidentified reference to Mya pictorum Linnaeus, 1758)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Spengler, 1793. Skr. Naturhist. Selsk. 3: 67, no. 32.
Unio musivus — Simpson (1900) Unio crassus musivus — Ortmann (1912), Bogatov (2022) Unio (sec. Lymnium) batavus — Simpson (1914) Unio crassus crassus — Haas (1940), Haas (1969), Fauna Europaea (2012), Klishko et al. (2017) Batavusiana (s.s.) musiva — Zatravkin (1983), Zatravkin (1987) Batavusiana (s.s.) musivus [??] — Korniushin (1998) Crassiana (Bataviana) musiva — Starobogatov et al. (2004) Crassiana musiva musiva — Kantor & Sysoev (2005), Kantor et al. (2010) Unio crassus — Graf (2007), Bogatov et al. (2018), Anisimova et al. (2018), Lopes-Lima et al. (2024) Unio (Crassiana) musivus — Vinarski & Kantor (2016) Unio crassus cf. courtillieri — Klishko et al. (2017)
Schröter, 1779. Gesch. Flußconch.: pl. 3, figs. 2, 4-5. Martini & Chemnitz, 1782. Conchyl. Cab. 6: 19, pl. 1, fig. 6. Bruguière, 1797. Encycl. Méth. 2: pl. 248, fig. 3. Maton & Rackett, 1807. Trans. Linn. Soc. London 8: 37.
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914) Unio (Batavusiana) batavus — Germain (1904) Unio (Groupe des Batavusiana) batavus — Caziot (1907) Unio batavus — Kobelt & Haas (1910), Haas (1924), Germain (1931) Unio crassus musivus — Ortmann (1912) Unio crassus batavus — Haas (1940), Haas (1969), Fauna Europaea (2012), Lopes-Lima et al. (2017) Batavusiana (s.s.) musiva — Zatravkin (1983), Zatravkin (1987) Unio (Crassiana) crassus — Vinarski & Kantor (2016) Unio crassus — Araujo et al. (2018)
Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): ??. (intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio pusillus Lea, 1840)
Scholtz, 1853. unknown publication: ??. (homonym of Unio ventricosus Barnes, 1823)
‘St. Germain, in litt.’ Moquin-Tandon, 1855. Hist. Natur. Moll. France 2: 571. (intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 287, pl. 10, fig. 1. (homonym of Unio acutus Rossmässler, 1837)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: 288, pl. 10, fig. 3. (intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio longus Philippi, 1869)
Unio batavus — Germain (1931)
Brot, 1867. Assoc. Zool. Léman Ann. (1866): pl. 9, fig. 1. (intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; misidentified reference to Unio batavus var. squamosus Charpentier, 1837)
Unio consentaneus — Germain (1931)
Malzine, 1867. Essai faun. malac. Belg.: 32. (homonym of Unio cuneatus Jacquemin, 1835; intro'd as intra-sp. of Unio ryckholtii Malzine, 1867)
Unio batavus — Germain (1931)
Colbeau, 1868. Ann. Soc. Malac. Belgique 3: 106, pl. 4, fig. 3. (intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)
Blanc in Westerlund & Blanc, 1879. Aperçu Faune Malac. Grèce: 145. (intro'd as intra-sp. of Unio vescoi Bourguignat, 1856)
Servain, 1888. Bull. Soc. Malac. France 5: 317. [nomen nudum] Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 141 [35], 206 [100]. ‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 172, fig. 186 [as ‘besnardinus’]. (renamed Unio besnardi Germain, 1904) Type(s): LECTOTYPE MNHN_22133 (by Graf, 2011: 463), La Loire, à Saint-Gemmes (Maine-et-Loire).
Unio (Batavusiana) besnardi — Germain (1904) Unio batavus — Germain (1931) Unio crassus batavus — Haas (1940), Haas (1969) Unio crassus — Lopes-Lima et al. (2024)
‘Linn.’ Draparnaud, 1806. Hist. Moll. France: 131, pl. 11, figs. 3 [as ‘Unio pictorum var. B’]. Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 145 [39], 208 [102]. ‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174, fig. 187 [as ‘senauxi’]. Type(s): LECTOTYPE MNHN_22523 (by Graf, 2011: 503), La Loire à Orléans.
Unio (Batavusiana) seneauxi — Germain (1904) Unio batavus — Germain (1931) Unio crassus batavus — Haas (1940), Haas (1969) Unio crassus — Lopes-Lima et al. (2024)
Locard, 1889. Contr. Faune Malacol. Franc. 13: 135 [29]. (homonym of Unio curtus Lea, 1859; intro'd as intra-sp. of Unio subrobustus Servain, 1885)
Kobelt, 1886. Icon. Land- und Süssw.-Moll. 2: pl. 53, fig. 286 [as ‘U. sieversi’]. Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 151. (intro'd as intra-sp. of Unio sieversi Drouët, 1881; homonym of Unio kobelti Neumayr, 1899)
‘Malzine’ of Authors. Locard, 1893. Coqu. Eau Douce Saumâtre France: 177. Germain, 1931. Faune de France 22: 732. Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 130. (error for Unio lambottei Malzine, 1867)
Drouët, 1898. Mém. Acad. Sci. Dijon ??: 61. (intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio (Anodontes) sirionos var. major d’Orbigny, 1835)
Unio batavus var. baraceus — Germain (1931)
Bourguignat in Locard, 1889. Contr. Faune Malacol. Franc. 13: 140 [34]. [nomen nudum] Bourguignat in Locard, 1899. Conchyliol. Portug.: 218.
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 261 [23]. (new name for Unio besnardianus Locard, 1889)
Unio (Batavusiana) besnardi — Germain (1904)
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 260 [22]. (intro'd as intra-sp. of Unio batavellus Locard, 1885)
Unio (Batavusiana) batavellus perconvexus — Germain (1904)
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 257 [19]. (intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio (Elliptio) viridis var. radiata Rafinesque, 1820)
Unio (Batavusiana) batavus radiata — Germain (1904)
Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 259 [21]. (intro'd as intra-sp. of Unio ligericus Locard, 1882)
Unio (Batavusiana) ligericus subsinuatus — Germain (1904)
‘Bourguignat in Locard’ of Authors. Germain, 1931. Faune de France 22: 724. Prie, 2012. MalaCo 8: 430. (error for Unio lagnisicus Locard, 1882)
‘Kobelt’ Zhadin, 1938. Faune USSR Mollusques 4(1): 89. (new name for Unio kungurensis chlebnikowi Kobelt, 1912)
‘Hattemann’ of Authors. Locard, 1889. Contr. Faune Malacol. Franc. 13: 132 [26]. ‘Hattemann’ Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 256 [18]. ‘Hattmann’ Germain, 1931. Faune de France 22: 724, 727. Lopes-Lima et al., 2024. MPE 195: suppl. append. 2, p. 3. (error for Unio courtillieri Hattemann, 1859)
Ziégl. in sched. of Authors.
‘Linn.’ Draparnaud, 1806. Hist. Moll. France: pl. 11, fig. 5. (misidentified reference to Mya margaritifera Linnaeus, 1758)
Unio consentaneus — Germain (1931)
‘Lamarck’ Nilsson, 1822. Hist. Moll. Svec.: 112. (reference to Mya batava Maton & Rackett, 1807)
‘Daud.’ Kickx, 1830. Synopsin Molluscorum Brabantiae: 82. (error for Unio lemovicincae Férussac, 1807)
‘Zieg.’ Lea, 1836. Synopsis Fam. Naiades: 27. (intro'd as syn of Mya batava Maton & Rackett, 1807)
‘Mühl. Pfeif.’ Lea, 1836. Synopsis Fam. Naiades: 27. (intro'd as syn of Mya batava Maton & Rackett, 1807)
Rossmässler, 1837. Icon. Land- und Süssw.-Moll. 1 (5-6): 56, pl 29, fig. 413 [as ‘Unio an n. sp.?’]. [nomen nudum]
‘Ziegl.’ Schlüter, 1838. Kurzgef. Syst. Verz. Conch. Samml.: 33. [infra-subsp.] (intro'd as intra-sp. of Unio fusculus Rossmässler, 1836)
Schlüter, 1838. Kurzgef. Syst. Verz. Conch. Samml.: 32. [infra-subsp.] (intro'd as intra-sp. of Unio reniformis Rossmässler, 1836)
‘Mich.’ Rossmässler, 1844. Icon. Land- und Süssw.-Moll. 2 (6): 29, pl. 57, fig. 757-761. (misidentified reference to Unio requienii Michaud, 1831)
Unio (sec. Lymnium) batavus — Simpson (1900) Unio (sec. Lymnium) batavus var. requieni — Simpson (1914)
J.F. Schmidt, 1847. Conch. Krain: 27. (misidentified reference to Unio piscinalis Rossmässler, 1836)
‘Küst.’ Lea, 1852. Synopsis Fam. Naiades: 30. (intro'd as syn of Mya batava Maton & Rackett, 1807)
‘Dup.’ Lea, 1852. Synopsis Fam. Naiades: 39. (error for Unio drouetii Dupuy, 1849)
‘Küst.’ Lea, 1852. Synopsis Fam. Naiades: 30. (intro'd as syn of Mya batava Maton & Rackett, 1807)
Drouët, 1852. Etudes Nayades de la France: pl. 6, fig. 3. (misidentified reference to Unio moquinianus Dupuy, 1843)
Unio consentaneus — Germain (1931)
‘Retz.’ Küster, 1854. Conchyl. Cab. 9(2): 113, pl. 31, fig. 5. [in part] (reference to Unio crassus Retzius, 1788)
Lampsilis (sec. Eurynia) radiatus — Simpson (1900) Lampsilis (Eurynia) radiata — Simpson (1914)
‘Nilss.’ Moquin-Tandon, 1855. Hist. Natur. Moll. France 2: 570. (reference to Unio ater Nilsson, 1822)
‘Nilss.’ Drouët, 1857. Étud. Naïades. France, Unio: 72, pl. 4, fig. 1. (reference to Unio ater Nilsson, 1822)
Unio consentaneus — Germain (1931)
‘Dup.’ Drouët, 1857. Étud. Naïades. France, Unio: 100, pl. 9, fig. 1. (misidentified reference to Unio platyrinchoideus Dupuy, 1849)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 2. [in part] (reference to Mya batava Maton & Rackett, 1807)
Unio (Batavusiana) batavus radiata — Germain (1904)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 3. (reference to Unio courtillieri Hattemann, 1859; error for Unio courtillieri Hattemann, 1859)
Unio (Batavusiana) batavus courtilieri — Germain (1904) Unio batavus — Germain (1931)
de Joannis, 1859. Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 10. [in part] (misidentified reference to Unio nana Lamarck, 1819)
Unio (Rhomboïdiana) nana — Germain (1904)
‘Parreis in sched.’ Küster, 1862. Conchyl. Cab. 9(2): 267. (intro'd as syn of Unio kotschyi Küster, 1862)
‘Lecomte’ Colbeau, 1868. Ann. Soc. Malac. Belgique 3: 106. [nomen nudum] (intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)
‘Bourg.’ Lea, 1870. Synopsis Fam. Naiades: 48. (error for Unio maccarthyanus Bourguignat, 1866; misidentified reference to Unio maccarthyanus Bourguignat, 1866)
‘Held’ Drouët, 1881. J. de Conch. 29: 27. (intro'd as syn of Unio batavus var. squamosus Charpentier, 1837)
Locard, 1882. Prodr. Malac. Franç. 1: 292. (misidentified reference to Unio sandrii Rossmässler, 1844)
Kobelt, 1886. Icon. Land- und Süssw.-Moll. 2: pl. 53, fig. 286. (misidentified reference to Unio sieversi Drouët, 1881)
‘Bourguignat’ Servain, 1888. Bull. Soc. Malac. France 5: 316, footnote. [nomen nudum] Locard, 1889. Contr. Faune Malacol. Franc. 13: 136 [30], footnote. [nomen nudum]
‘Thièse’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 141 [35], footnote. [nomen nudum]
Locard, 1889. Contr. Faune Malacol. Franc. 13: 129 [23]. [nomen nudum] (intro'd as intra-sp. of Unio pilloti Locard, 1882)
Locard, 1889. Contr. Faune Malacol. Franc. 13: 127 [21]. [nomen nudum] Westerlund, 1890. Fauna Paläarct. Binnenconch. 2(7): 84. [nomen nudum] (intro'd as intra-sp. of Unio suborbicularis Drouët, 1888)
‘Servain’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 172. (error for Unio besnardianus Locard, 1889)
Unio (Batavusiana) besnardi — Germain (1904)
‘Bourguignat’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174. (error for Unio seneauxi Locard, 1889)
‘Servain in Locard’ Locard, 1893. Coqu. Eau Douce Saumâtre France: 174. (error for Unio surraulti Locard, 1889)
‘Spengler’ Simpson, 1900. Proc. U.S. Nat. Mus. 22: 744. (error for Unio musivus Spengler, 1793)
Unio (sec. Lymnium) batavus — Simpson (1914)
‘Bourguignat’ Germain, 1904. Bull. Soc. Sci. Nat. Ouest France 4: 265 [27]. (error for Unio hattemani Locard, 1889)
‘Pacôme’ Germain, 1931. Faune de France 22: 728. (error for Unio caroliensis Locard, 1889)
‘Servain’ Germain, 1931. Faune de France 22: 728. (error for Unio visurgisinus Servain, 1888)
‘Bourguignat’ Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 131. (error for Unio hattemani Locard, 1889)
‘Kobelt’ Haas, 1940. Zool. Ser. Field Mus. Nat. Hist. 24: 132. ‘Kobelt’ Haas, 1969. Das Tierreich 88: 53. Vinarski & Kantor, 2016. Analyt. Cat. Fr. Brack. Moll. Russia: 25. (error for Unio kungurensis sylwensis Kobelt, 1912)
‘Locard’ Haas, 1969. Das Tierreich 88: 51. (error for Unio lancelevei Locard, 1893)
‘Lindholm’ Zatravkin, 1987. Byull. Mosk. Obshch.. Ispyt. Prir. 92(4): 57. (error for Unio ater var. irgizlaicus Lindholm, 1904)
Taxonomic History (since 1900)Lampsilis (sec. Eurynia) radiatus — Simpson (1900) Quadrula (sec. Fusconaia) coccinea — Simpson (1900), Simpson (1914) Margaritana crassa — Simpson (1900), Simpson (1914) Unio (sec. Lymnium) littoralis — Simpson (1900), Simpson (1914) Unio (sec. Lymnium) pseudonymus — Simpson (1900), Simpson (1914) Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914) Unio musivus — Simpson (1900) Unio baudoni — Kobelt (1902) Unio lagnysicus [sic] — Rochebrune (1903), Kobelt & Haas (1910) Unio carantoni — Rochebrune (1903) Unio crassatellus — Rochebrune (1903) Unio materniacus — Rochebrune (1903) Unio batavellus — Rochebrune (1903), Kobelt & Haas (1910) Unio (Rhomboïdiana) nana — Germain (1904) Unio (Melasiana) melas — Germain (1904) Unio (Mancusiana) mancus — Germain (1904) Unio (Fusculusiana) fusculus — Germain (1904) Unio (Socardiana) feliciani — Germain (1904) Unio (Batavusiana) batavus — Germain (1904) Unio (Batavusiana) batavus radiata — Germain (1904) Unio (Batavusiana) batavus courtilieri — Germain (1904) Unio (Batavusiana) andegavensis — Germain (1904) Unio (Batavusiana) diptychus — Germain (1904) Unio (Batavusiana) ligericus — Germain (1904) Unio (Batavusiana) ligericus subsinuatus — Germain (1904) Unio (Batavusiana) batavellus — Germain (1904) Unio (Batavusiana) batavellus perconvexus — Germain (1904) Unio (Batavusiana) arenarum — Germain (1904) Unio (Batavusiana) besnardi — Germain (1904) Unio (Batavusiana) ryckholtii — Germain (1904) Unio (Batavusiana) visurgicus [sic] — Germain (1904) Unio (Batavusiana) ingrandiensis — Germain (1904) Unio (Batavusiana) seneauxi — Germain (1904) Unio (Batavusiana) surraulti — Germain (1904) Unio (Batavusiana) gliscerus — Germain (1904) Unio (Batavusiana) matronicus — Germain (1904) Unio (Batavusiana) nicolloni — Germain (1904) Unio (Batavusiana) ampullaceus — Germain (1904) Unio (Lemotheuxiana) adonus — Germain (1904) Unio (Lemotheuxiana) lemotheuxi — Germain (1904), Caziot (1910) Unio (Lemotheuxiana) hattemanni [sic] — Germain (1904) Unio (Ateriana) lambottei — Germain (1904) Unio (Ateriana) brevieri [sic] — Germain (1904) Unio (Ateriana) marcellinus — Germain (1904) Unio (Groupe des Ninusiana) [sic] rayi — Caziot (1907) Unio (Groupe des Mancusiana) subtilis — Caziot (1907) Unio (Groupe des Batavusiana) batavus — Caziot (1907) Unio (Groupe des Ateriana) brevierei — Caziot (1907) Unio subamnicus — Kobelt & Haas (1910) Unio consentaneus var. — Kobelt & Haas (1910) Unio nana — Kobelt & Haas (1910) Unio rayi — Kobelt & Haas (1910) Unio pilloti — Kobelt & Haas (1910) Unio batavus — Kobelt & Haas (1910), Haas (1924), Germain (1931) Unio crassus — Kobelt & Haas (1910), Ortmann (1912), Ortmann (1914), Germain (1931), Zhadin (1965), ... Dai et al. (2025), Wu et al. (2025), Chen et al. (2025), Hou et al. (2025), Wang et al. (2025) Unio crassus musivus — Ortmann (1912), Bogatov (2022) Lampsilis (Eurynia) radiata — Simpson (1914) Unio (sec. Lymnium) batavus var. requieni — Simpson (1914) Quadrula (Obliquata) coccinea — Frierson (1927) Margaritana auricularia — Germain (1931) Unio crassus var. piscinalis — Germain (1931) Unio mancus — Germain (1931) Unio mancus var. macrorhynchus — Germain (1931) Unio mancus var. nanus — Germain (1931) Unio mancus var. amnicus — Germain (1931) Unio mancus var. amnicus form riparius — Germain (1931) Unio mancus var. elongatulus — Germain (1931) Unio batavus var. baraceus — Germain (1931) Unio consentaneus — Germain (1931) Unio consentaneus var. oxyrhynchus — Germain (1931) Unio consentaneus var. moquini — Germain (1931) Unio (s.s.) crassus — Zhadin (1938) Unio (s.s.) crassus var. pseudolitoralis — Zhadin (1938) Unio (s.s.) crassus kungurensis — Zhadin (1938) Unio (s.s.) crassus var. chlebnikovi [sic] — Zhadin (1938) Unio (s.s.) crassus var. okae — Zhadin (1938) Unio (s.s.) crassus var. ishmensis — Zhadin (1938) Unio (s.s.) crassus var. roseni — Zhadin (1938) Unio (s.s.) ater — Zhadin (1938) Unio crassus crassus — Haas (1940), Haas (1969), Fauna Europaea (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017), Klishko et al. (2017), Bogatov (2022) Unio crassus cytherea — Haas (1940), Haas (1969) Unio crassus batavus — Haas (1940), Haas (1969), Fauna Europaea (2012), Lopes-Lima et al. (2017) Unio crassus gontieri — Haas (1940), Haas (1969), Graf (2007) Unio crassus var. pseudolitoralis — Zhadin (1965) Crassiana crassa crassa — Akromovski (1976), Kantor & Sysoev (2005), Kantor et al. (2010) Batavusiana (s.s.) musiva — Zatravkin (1983), Zatravkin (1987) Batavusiana (s.s.) irenjensis — Zatravkin (1983), Stadnichenko (1984), Zatravkin (1987) Batavusiana (Crassunio) crassa — Zatravkin (1983), Stadnichenko (1984), Zatravkin (1987) Unio (Tumidusiana) tumidus var. minor — Stadnichenko (1984) Batavusiana (s.s.) irgislaica [sic] — Zatravkin (1987) Unio (Crassunio) crassus — Falkner (1994) Pleurobema sintoxia — Parmalee & Bogan (1998), Williams et al. (2008) Batavusiana (s.s.) musivus [??] — Korniushin (1998) Batavusiana (Crassiana) crassus — Korniushin (1998) Unio (Crassiana) crassus crassus — Falkner et al. (2001), Falkner et al. (2002) Unio (Crassiana) crassus courtillieri — Falkner et al. (2001), Falkner et al. (2002) Crassiana (s.s.) crassa — Starobogatov et al. (2004) Crassiana (Bataviana) irgizlaica — Starobogatov et al. (2004) Crassiana (Bataviana) musiva — Starobogatov et al. (2004) Crassiana (Bataviana) cyprinorum — Starobogatov et al. (2004) Crassiana cyprinorum — Kantor & Sysoev (2005), Kantor et al. (2010) Crassiana irenjensis — Kantor & Sysoev (2005), Kantor et al. (2010) Crassiana irgizlaica — Kantor & Sysoev (2005), Kantor et al. (2010) Crassiana musiva musiva — Kantor & Sysoev (2005), Kantor et al. (2010) Unio mancus mancus — Prie (2012) Unio mancus moquinianus — Prie (2012) Unio crassus courtillieri — Prié & Puillandre (2014), Lopes-Lima et al. (2017) Unio (Crassiana) crassus — Vinarski & Kantor (2016) Unio (Crassiana) cyprinorum — Vinarski & Kantor (2016) Unio (Crassiana) irenjensis — Vinarski & Kantor (2016) Unio (Crassiana) irgizlaicus — Vinarski & Kantor (2016) Unio (Crassiana) musivus — Vinarski & Kantor (2016) Unio crassus cf. courtillieri — Klishko et al. (2017) Unio crassus courtillierii [sic] — Araujo et al. (2018) Unio c. crassus — Príe et al. (2021) Unio c. courtillieri — Príe et al. (2021) Unio cf. courtillieri — Lopes-Lima et al. (2021)
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