species Pseudunio auricularius (Spengler, 1793): synonymy

Mya margaritifera Gmelin, 1791

(misidentified reference to Mya margaritifera Linnaeus, 1758)
Lister, 1678. Hist. Anim. Anglic.: 15, pl. 1, fig. 1. [pre-linnaean]
Lister, 1685. Conch.: pl. 149, fig. 4. [pre-linnaean]
Klein, 1753. Tent. Method. Ostrac., etc.: pl. 10, fig. 47. [pre-linnaean]
Knorr, 1757. Vergnügen 2(4): pl. 25, fig. 2.
Schröter, 1779. Gesch. Flußconch.: pl. 4, fig. 1.
Martini & Chemnitz, 1782. Conchyl. Cab. 6: 15, pl. 1, fig. 5.
Gmelin, 1791. Syst. Nat. 1(6): 3219, sp. 4.

Unio auricularius Spengler, 1793

Lister, 1685. Conch.: pl. 149, fig. 4. [pre-linnaean]
Spengler, 1793. Skr. Naturhist. Selsk. 3: 54, no. 21.
LECTOTYPE ZMUC_BIV-315 (by Falkner, 1994: __), Berlingen near Unterseen, Switzerland.

Margaritana crassa — Simpson (1900)
Margaritana margaritifera — Preston (1912)
Margaritana (Group of M. margaritifera) crassa — Simpson (1914)
Margaritana auricularia — Haas (1924), Germain (1931)
Margaritana (Potomida) auricularius — Thiele (1934)
Margaritifera auricularia auricularia — Haas (1940)
Margaritifera (Pseudunio) auricularia auricularia — Haas (1969)
Margaritifera (Pseudunio) auricularia — Haas (1969), Van Damme (1984), Bolotov et al. (2016)
Margaritifera auricularia — Mandahl-Barth (1988), Ziuganov et al. (1994), Daget (1998), Nagel et al. (1998), Bába (2000), ... Huang et al. (2018), Prié et al. (2018), Huang et al. (2019), Múrria et al. (2020), Sano et al. (2020)
Pseudunio auricularius — Falkner (1994), Lopes-Lima et al. (2018), Lyubas et al. (2019), Vikhrev et al. (2019), Guerra et al. (2019), ... Gomes-dos-Santos et al. (2023), Zieritz et al. (2024), Nakamura et al. (2024), Chen et al. (2025), Campbell & Lydeard (2026)
Pseudunio auricularia — D.G. Smith (2001), Graf & Cummings (2007), Wu et al. (2020)
Pseudunio auricularius auricularius — Falkner et al. (2001), Falkner et al. (2002)

Unio rugosa Poiret, 1801

(homonym of Mya rugosa Gmelin, 1791)
Poiret, 1801. Coq. Fluv., et Terr. de l’Aisne: 105.

Margaritana crassa — Simpson (1900)
Margaritana (Group of M. margaritifera) crassa — Simpson (1914)
Margaritana auricularia — Germain (1931)
Margaritifera auricularia — Ziuganov et al. (1994)

Unio sinuata Lamarck, 1819

Lister, 1685. Conch.: pl. 149, fig. 3 [sic]. [pre-linnaean]
Knorr, 1757. Vergnügen 2(4): pl. 25, fig. 2. [pre-linnaean]
Schröter, 1779. Gesch. Flußconch.: pl. 4, fig. 1.
Bruguière, 1797. Encycl. Méth. 2: pl. 248, figs. 1a-b [as Unio sp.].
Lamarck, 1819. Hist. Natur. Anim. sans Vert. 6: 70, sp. 1.
SYNTYPES MHNG_XXX, the Rhine, the Loire and other rivers of continental Europe.

Margaritana crassa — Simpson (1900)
Unio sinuatus — Rochebrune (1903), Kobelt (1903)
Unio (Baryana) sinuatus — Germain (1904)
Unio (Groupe Baryana) sinuatus — Caziot (1907)
Pseudunio sinuatus — Haas (1910), Gozhik (2006)
Margaritana sinuata — Ortmann (1912), Ortmann (1913)
Margaritana (Group of M. margaritifera) crassa — Simpson (1914)
Margaritana auricularia — Haas (1924), Germain (1931)
Margaritana (Potomida) auricularius — Thiele (1934)
Margaritifera auricularia auricularia — Haas (1940)
Unio crassus — R.I. Johnson (1953)
Margaritifera (Pseudunio) auricularia auricularia — Haas (1969)
Margaritifera (Pseudunio) auricularia — Haas (1969)
Margaritifera auricularia — Ziuganov et al. (1994), Nagel et al. (1998), Graf & Cummings (2009), Araujo et al. (2009), IUCN (2010), Fauna Europaea (2012)
Pseudunio auricularia — D.G. Smith (2001)
Pseudunio auricularius — Lopes-Lima et al. (2018), Lyubas et al. (2019), Graf & Cummings (2021)

Unio crassissima Férussac, 1827

(homonym of Unio crassissimus J. de C. Sowerby, 1817)
‘ex Des Moulins’ Férussac, 1827. Moll. Gir.: 42.

Margaritana auricularia — Germain (1931)

Unio sinuatus compressus Moquin-Tandon, 1855

(homonym of Unio compressus J. de C. Sowerby, 1828; intro'd as intra-sp. of Unio sinuata Lamarck, 1819)
Moquin-Tandon, 1855. Hist. Natur. Moll. France 2: 567.

Unio margaritanopsis Locard, 1889

Locard, 1889. Contr. Faune Malacol. Franc. 13: 123 [17], 182 [76].
Locard, 1893. Coqu. Eau Douce Saumâtre France: 151, fig. 163.

Margaritana margaritanopsis — Germain (1931)
Margaritifera auricularia auricularia — Haas (1940)
Margaritifera (Pseudunio) auricularia auricularia — Haas (1969)
Margaritifera auricularia — Ziuganov et al. (1994), Araujo et al. (2009), IUCN (2010)

Unio araris Barbié of Authors

Barbié of Authors.

Margaritana auricularia — Germain (1931)

Unio garumnae de Grateloup of Authors

de Grateloup of Authors.

Margaritana auricularia — Germain (1931)

Unio margaritacea ‘Drap.’ of Authors

(error for Unio margaritifera Draparnaud, 1801)
‘Drap.’ of Authors.
Lea, 1838. Synopsis Fam. Naiades: 26.

Dichonca crassissima Klein, 1753

Klein, 1753. Tent. Method. Ostrac., etc.: 128, pl. 10, fig. 47. [pre-linnaean]

Margaritifera margaritifera — Watters et al. (2009)

Unio margaritifera Draparnaud, 1801

(misidentified reference to Mya margaritifera Linnaeus, 1758)
Draparnaud, 1801. Tab. Moll. Fr.: 107.
‘Linn.’ Draparnaud, 1806. Hist. Moll. France: 132, pl. 10, figs. 17-19.

Margaritana margaritifera — Preston (1912)
Margaritana auricularia — Germain (1931)
Margaritifera auricularia — IUCN (2010)

Unio margaritifera Millet, 1813

(misidentified reference to Mya margaritifera Linnaeus, 1758)
Millet, 1813. Moll. Terr. Fluv. Maine et Loire: ??.

Unio (Baryana) sinuatus — Germain (1904)

Margarita (Unio) crassissimus Lea, 1836

(reference to Unio crassissima Férussac, 1827)
‘Ferussac’ Lea, 1836. Synopsis Fam. Naiades: 40.
‘Fer. Des Moul. Grat.’ Lea, 1838. Synopsis Fam. Naiades: 26.

Margaritana margaritifera — Simpson (1900), Preston (1912)
Margaritana (Group of M. margaritifera) margaritifera — Simpson (1914)
Margaritifera auricularia — Ziuganov et al. (1994), IUCN (2010)

Unio auricularis Lea, 1836

(error for Unio auricularius Spengler, 1793)
‘Speng.’ Lea, 1836. Synopsis Fam. Naiades: 40.

Unio sinuosa Swainson, 1840

(error for Unio sinuata Lamarck, 1819)
‘Lamarck’ Swainson, 1840. Treat. Malacol.: 262.

Unio sinuata Sganzin, 1841

(reference to Unio sinuata Lamarck, 1819)
‘Lamarck’ Sganzin, 1841. Mém. Soc. Hist. Nat. Strasbourg 3: 8.

Margaritifera auricularia — Graf & Cummings (2009)

Unio baryus Locard, 1889

‘Bourg.’ Locard, 1889. Contr. Faune Malacol. Franc. 13: 124 [18], footnote. [nomen nudum]

Margarita (Unio) crassiamus Preston, 1912

(error for Margarita (Unio) crassissimus Lea, 1836)
‘Lea’ Preston, 1912. Rec. Ind. Mus. 7: 286.