MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Tue 02 June 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina
genus Utterbackia F.C. Baker, 1927: cladomics
3 Recent valid species (3 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 3 (3 total)
analyzed in 117 cladograms in 68 publications
maximum number of terminals in an analysis = 3 (mean = 1.55, median = 2)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 63
maximum number of included terminals = 3, (mean = 2.02 median = 2)
proportion of cladograms that support basic monophyly = 0.95
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Hoeh et al., 1995 [Publ 3391], fig. 2
** Top Monophyly Cladogram **
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 4 | 4 | 1 | ** |
+-------- Pyganodon --+ | +----- Utterbackia_peggyae ** +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS
** Top Sister-Group Cladogram **
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 32 | 14 | -- |
+-------------- Utterbackiana
+--+
| | +----------- Alasminota
| +--+
| | +-------- Pseudodontoideus
| +--+
| | +----- Lasmigona
| +--+
| | +-- Pterosyna
| +--+
--+ +-- Arcidens
|
| +-- Prolasmidonta
| +--+
| | +-- Platynaias
| +-----+
| | | +-- Ligodonta
| | +--+
| | +-- Alasmidonta
+-----+
| +-- Anodontoides
| +-----+
| | +-- Strophitus
+--+
| +----- Sulcularia
+--+
| +-- Pyganodon
+--+
+-- Utterbackia_peninsularis **
Whelan et al., 2023 [Publ 4287], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 1 | 2 | 39 | 31 | 13 | -- |
+-------- Pyganodon
+-----+
| | +----- Utterbackia_peninsularis__2 **
| +--+
| | +-- Utterbackia_peninsularis__1 **
| +--+
| +-- Utterbackia_imbecillis **
|
--+ +-------- Utterbackiana
| |
| +--+ +-- Lasmigona
| | | +--+
| | | | +-- Pseudodontoideus
| | +--+
| | | +-- Pterosyna
| | +--+
+--+ +-- Arcidens
|
| +----- Sulcularia
| +--+
| | | +-- Anodontoides
| | +--+
| | +-- Strophitus
+--+
| +-- Prolasmidonta
| +--+
| | +-- Platynaias
+--+
| +-- Alasminota
+--+
+-- Alasmidonta
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.933 | 2 | 2 | 1 | 2 | 630 | 35 | 13 | -- |
+-------- Pyganodon
+--------+
| | +----- Utterbackia_peninsularis__2 **
| +--+
| | +-- Utterbackia_imbecillis **
--+ +--+
| +-- Utterbackia_peninsularis__1 **
|
| +-------------- Utterbackiana
| |
+--+ +----------- Simpsonaias
| |
| | +-- Alasminota
+--+ +-----+
| | +-- Alasmidonta
| |
+--+ +-- Prolasmidonta
| +--+
| | +-- Platynaias
+--+
| +-- Anodontoides
+--+
+-- Strophitus
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.455 | 2 | 20 | 0 | 2 | 103 | 14 | 9 | 20.0 |
+-- Alasmidonta_marginata
|
+-- Prolasmidonta
|
+-- Alasmidonta_undulata
|
+-- Beringiana
|
+-- Brachyanodon_kennerlyi
|
+-- Brachyanodon_nuttalliana
|
+-- Brachyanodon_oregonensis
|
+-- Arcidens_confragosus
|
+-- Arcidens_wheeleri
|
+-- Pterosyna
|
+--------------------------------------------------+-- Platynaias_compressa
| |
| +-- Lasmigona
| |
| +-- Platynaias_decorata
| |
| +-- Platynaias_subviridis
| |
| +-- Pyganodon_grandis
| |
| +-- Pyganodon_lacustris
| |
| +-- Pseudodontoideus
| |
| +-- Strophitus
| |
--+ +-- Utterbackia_imbecillis **
| |
| +-- Utterbackia_peninsularis__1 **
| |
| +-- Utterbackia_peninsularis__2 **
|
| +-------------------------------------------------- Gonidea
| |
| | +----------------------------------------------- Reginaia
| | |
| | +----------------------------------------------- Amblema
| | |
| | | +----- Popenaias
| | | |
| | | | +-- Dromus
| | | | |
| | | | +-- Epioblasma
+--+ | | |
| +-----------------------------------------+ +-- Glebula
| | | |
| | | +-- Lampsilis_ovata
| | | |
| | | +-- Lampsilis_virescens
| | | |
| | | +-- Lemiox
| | +--+
| | +-- Obliquaria
| | |
+--+ +-- Obovaria_olivaria
| |
| +-- Obovaria_unicolor
| |
| +-- Plectomerus
| |
| +-- Potamilus_alatus
| |
| +-- Potamilus_capax
|
| +----- Uniomerus
| |
| | +-- Cyclonaias
| +--------------------------------------+ |
| | | +-- Pustulosa_pustulosa__1
| | | |
| | | +-- Theliderma
| | +--+
| | +-- Pustulosa_pustulosa__2
| | |
+--+ +-- Quadrula_quadrula
| |
| +-- Quadrula_verrucosa
|
| +----------------------------------------- Elliptoideus
| |
| | +-------------------------------------- Parvaspina
| | |
+--+ | +----------------------------------- Plethobasus
| | |
| | +----------------------------------- Eurynia
| | |
| | +----------------------------------- Hemistena
+--+ |
| +----------------------------------- Elliptio
| |
| | +-------------------------------- Pleuronaia
| | |
| | | +-- Pleurobema_cordatum
| | | +--------------------------+
+--+ | | +-- Fusconaia
| | |
| | | +-------------------------- Pleurobema_plenum
| | | |
| | | | +----------------------- Pleurobema_stabile
| | | | |
| | | | | +----- Pleurobema_pyriforme
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum__2
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni
| | | | | | |
+--+ | +--+ | | +----- Pleurobema_decisum
| | | | | |
| | | | +--+ +-- Pleurobema_georgianum__1
| | | | | |
| | | | +--+-- Pleurobema_hanleyianum__1
| | | | |
| | | | +-- Pleurobema_hanleyianum__2
| | | |
| | +--+ +-------------- Pleurobema_beadleianum__2
| | | |
| | | +--+ +----------- Pleurobema_strodeanum__2
| | | | | |
| | | | +--+----------- Pleurobema_rubellum
| | | | |
| | | | | +-------- Pleurobema_beadleianum__1
| | | | +--+
+--+ | | | +----- Pleurobema_taitianum
| | | +--+
| | | | +-- Pleurobema_perovatum
| +--+ +--+
| | +-- Pleurobema_strodeanum__1
| |
| | +----- Pleurobema_oviforme__6
| | |
| | +--+----- Pleurobema_clava__3
| | | |
| | | | +-- Pleurobema_oviforme__5
| | | +--+
| +--------+ +-- Pleurobema_clava__2
| |
| | +----- Pleurobema_oviforme__2
| | |
| | +----- Pleurobema_oviforme__3
| +--+
| +----- Pleurobema_oviforme__4
| |
| | +-- Pleurobema_clava__1
| +--+
| +-- Pleurobema_oviforme__1
|
| +----------------- Pleurobema_riddellii
| |
| | +-------------- Pleurobema_sintoxia__34
| | |
| | | +----------- Pleurobema_sintoxia__31
| | | |
+-----------+ | +----------- Pleurobema_sintoxia__32
| | |
| | +----------- Pleurobema_sintoxia__33
| | |
| | | +-------- Pleurobema__4
| | | |
| | | +-------- Pleurobema__5
+--+ | |
| | +-------- Pleurobema_sintoxia__2
| | |
| | +-------- Pleurobema_sintoxia__3
| | |
| | +-------- Pleurobema__8
| | |
| | +-------- Pleurobema__9
| | |
| | +-------- Pleurobema__10
| | |
| | +-------- Pleurobema_sintoxia__5
+--+ |
| +-------- Pleurobema_sintoxia__6
| |
| +-------- Pleurobema_sintoxia__7
| |
| +-------- Pleurobema_sintoxia__8
| |
| +-------- Pleurobema_sintoxia__9
| |
| +-------- Pleurobema_sintoxia__10
| |
| +-------- Pleurobema_sintoxia__11
| |
| +-------- Pleurobema_sintoxia__12
| |
| +-------- Pleurobema_sintoxia__13
| |
| +-------- Pleurobema_sintoxia__14
| |
| +-------- Pleurobema_sintoxia__15
| |
| +-------- Pleurobema_sintoxia__16
+--+
+-------- Pleurobema_sintoxia__17
|
+-------- Pleurobema_sintoxia__18
|
+-------- Pleurobema_sintoxia__19
|
+-------- Pleurobema_sintoxia__20
|
+-------- Pleurobema_sintoxia__21
|
+-------- Pleurobema_sintoxia__22
|
+-------- Pleurobema_sintoxia__23
|
+-------- Pleurobema_sintoxia__24
|
+-------- Pleurobema_sintoxia__25
|
+-------- Pleurobema_sintoxia__26
|
+-------- Pleurobema_sintoxia__27
|
+-------- Pleurobema_sintoxia__28
|
+-------- Pleurobema_sintoxia__29
|
+-------- Pleurobema_sintoxia__30
|
| +-- Pleurobema__6
+-----+
| +-- Pleurobema_sintoxia__1
|
| +-- Pleurobema__7
+-----+
| +-- Pleurobema_sintoxia__4
|
| +----- Pleurobema__3
+--+
| +-- Pleurobema__1
+--+
+-- Pleurobema__2
Inoue et al., 2014 [Publ 3635], fig. 3A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 1 | 2 | 18 | 12 | 8 | -- |
+-------- Pseudodontoideus
+--------+
| | +----- Lasmigona
| +--+
| | +-- Pterosyna
--+ +--+
| +-- Arcidens
|
| +-------------- Strophitus
| |
+--+ +-- Platynaias
| +--------+
| | +-- Alasmidonta
+--+
| +-------- Pyganodon
+--+
| +----- Utterbackia_peninsularis__2 **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis__1 **
Breton et al., 2011 [Publ 3376], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 1 | 2 | 36 | 9 | 6 | -- |
+-- Pterosyna
+--------------+
| +-- Lasmigona
--+
| +-------------- Strophitus
| |
+--+ +-- Platynaias
| +--------+
| | +-- Alasmidonta
+--+
| +-------- Pyganodon
+--+
| +----- Utterbackia_peninsularis__2 **
+--+
| +-- Utterbackia_peninsularis__1 **
+--+
+-- Utterbackia_imbecillis **
Chase et al., 2018 [Publ 3847], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 1 | 2 | 37 | 9 | 6 | -- |
+-------- Pyganodon
+--+
| | +----- Utterbackia_peninsularis__2 **
| +--+
| | +-- Utterbackia_peninsularis__1 **
| +--+
--+ +-- Utterbackia_imbecillis **
|
| +-- Lasmigona
| +-----+
| | +-- Pterosyna
+--+
| +----- Alasmidonta
+--+
| +-- Strophitus
+--+
+-- Platynaias
Hoeh, 1990 [Publ 2717], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 2 | 2 | 0 | 2 | 13 | 11 | 5 | -- |
+----- Pyganodon --+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peggyae **
Hoeh, 1990 [Publ 2717], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.429 | 2 | 15 | 0 | 2 | 13 | 11 | 5 | 15.0 |
+----- Pyganodon_fragilis | +----- Utterbackia_imbecillis ** | +----- Utterbackia_peggyae ** | +----- Anodonta | +----- Brachyanodon | +----- Utterbackiana_couperiana__1 | --+----- Utterbackiana_couperiana__2 | +----- Utterbackiana_suborbiculata | | +-- Lasmigona +--+ | +-- Utterbackiana_implicata | | +-- Pterosyna | | | +-- Platynaias | | | +-- Pyganodon_cataracta +--+ +-- Pyganodon_gibbosa | +-- Pyganodon_grandis | +-- Pyganodon_lacustris
Hoeh, 1990 [Publ 2717], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 2 | 2 | 0 | 2 | 13 | 11 | 5 | -- |
+----- Pyganodon --+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peggyae **
Zanatta et al., 2007 [Publ 2698], fig. 1A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 2 | 2 | 0 | 2 | 14 | 12 | 5 | -- |
+----- Pyganodon --+ | +-- Utterbackia_peggyae ** +--+ +-- Utterbackia_imbecillis **
Zanatta et al., 2007 [Publ 2698], fig. 1B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 2 | 2 | 0 | 2 | 14 | 12 | 5 | -- |
+----- Pyganodon --+ | +-- Utterbackia_peggyae ** +--+ +-- Utterbackia_imbecillis **
Huang et al., 2019 [Publ 3843], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 2 | 2 | 0 | 2 | 199 | 5 | 5 | -- |
+----- Pyganodon --+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Huang et al., 2019 [Publ 3843], fig. S3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 2 | 2 | 0 | 2 | 197 | 5 | 5 | -- |
+----- Pyganodon --+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Huang et al., 2019 [Publ 3843], fig. S4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 2 | 2 | 0 | 2 | 199 | 5 | 5 | -- |
+-------- Acuticosta | +-------- Alasmidonta | +-------- Anodonta_anatina | +-------- Anemina | +-------- Anodonta_cygnea | +-------- Cristaria | +-------- Brachyanodon | +-------- Lasmigona | +-------- Lepidodesma | +-------- Pseudanodonta | +-------- Pyganodon | +-------- Simpsonaias --+ +-------- Strophitus | +-------- Schistodesmus | +-------- Unio | +-------- Scriptolamprotula | +-------- Lanceolaria | | +-- Utterbackia_imbecillis ** +-----+ | +-- Utterbackia_peninsularis ** | | +-- Pletholophus +-----+ | +-- Sinanodonta | | +----- Cuneopsis_celtiformis | | | +----- Tchangsinaia | | +--+----- Cuneopsis_rufescens | +----- Nodularia | | +-- Cuneopsis_szechenyii +--+ +-- Cuneopsis_heudei
Stewart et al., 2021 [Publ 4451], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 1 | 2 | 14 | 8 | 5 | -- |
+-- Alasmidonta
+-----------+
| +-- Pterosyna
|
--+ +-- Strophitus
| +--------+
| | +-- Platynaias
+--+
| +-------- Pyganodon
+--+
| +----- Utterbackia_peninsularis__2 **
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis__1 **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 93 | 5 | 5 | -- |
+----- Pyganodon
+--+
| | +-- Utterbackia_peninsularis **
| +--+
| +-- Utterbackia_imbecillis **
--+
| +-- Alasmidonta
| +--+
| | +-- Platynaias
+--+
| +-- Prolasmidonta
+--+
+-- Pterosyna
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 93 | 5 | 5 | -- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 89 | 5 | 5 | -- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 89 | 5 | 5 | -- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Wu et al., 2024 [Publ 4379], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 51 | 5 | 5 | -- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Hou et al., 2024 [Publ 4400], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 73 | 5 | 5 | -- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Hou et al., 2025 [Publ 4445], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 50 | 5 | 5 | -- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Platynaias
+--+
+-- Alasmidonta
Wang et al., 2025 [Publ 4488], fig. 3B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 62 | 5 | 5 | -- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 138 | 6 | 5 | -- |
+----- Pyganodon
+--+
| | +-- Utterbackia_imbecillis **
| +--+
| +-- Utterbackia_peninsularis **
--+
| +-- Pterosyna
| +--+
| | +-- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Wu et al., 2026 [Publ 4811], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 93 | 5 | 5 | -- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Wu et al., 2026 [Publ 4825], fig. 6B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 63 | 5 | 5 | -- |
+----- Pyganodon
+-----+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Sano et al., 2020 [Publ 4138], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 67 | 4 | 4 | -- |
+----- Alasmidonta
+--+
| | +-- Prolasmidonta
| +--+
--+ +-- Platynaias
|
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Sano et al., 2020 [Publ 4138], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 67 | 4 | 4 | -- |
+----- Pyganodon
+--+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +----- Alasmidonta
+--+
| +-- Platynaias
+--+
+-- Prolasmidonta
Sano et al., 2020 [Publ 4138], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 67 | 4 | 4 | -- |
+----- Alasmidonta
+--+
| | +-- Prolasmidonta
| +--+
--+ +-- Platynaias
|
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Chen et al., 2020 [Publ 4170], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 2 | 27 | 0 | 2 | 32 | 4 | 4 | 26.0 |
+----- Prolasmidonta
+--+
| | +-- Alasmidonta
| +--+
| +-- Anemina
+-----------------+
| | +-- Lamprotula_caveata
| | +--+
| | | +-- Chamberlainia
| +--+
| | +-- Pseudodon
| +--+
| +-- Pilsbryoconcha
|
| +----- Schistodesmus
| +-----+
| | | +-- Lepidodesma
--+ | +--+
| | +-- Scriptolamprotula
| +-----------+
| | | +----- Platynaias
| | | +--+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Pyganodon
| | |
| | | +----- Sinanodonta
| | +--+
| | | +-- Cristaria
+--+ +--+
| +-- Anodonta
|
| +-- Utterbackia_peninsularis **
| +--------+
| | +-- Potamilus_alatus
| +--------+
| | | +-------- Lamprotula_leaii
| | +--+
| | | +----- Sinohyriopsis
| | +--+
+--+ | +-- Parvasolenaia
| +--+
| +-- Sinosolenaia
|
| +----------------- Gibbosula
+--+
| +-------------- Quadrula
+--+
| +----------- Toxolasma
+--+
| +-------- Lampsilis_ornata
+--+
| +----- Potamilus_leptodon
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Wu et al., 2021 [Publ 4366], fig. 3-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 71 | 5 | 4 | -- |
+----- Pyganodon
+--+
| | +-- Utterbackia_peninsularis **
| +--+
--+ +-- Utterbackia_imbecillis **
|
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Wu et al., 2021 [Publ 4366], fig. 3-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 71 | 5 | 4 | -- |
+----- Pyganodon
+--+
| | +-- Utterbackia_peninsularis **
| +--+
--+ +-- Utterbackia_imbecillis **
|
| +----- Prolasmidonta
+--+
| +-- Alasmidonta
+--+
+-- Platynaias
Wu et al., 2021 [Publ 4366], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 66 | 4 | 4 | -- |
+----- Pyganodon
+--+
| | +-- Utterbackia_imbecillis **
| +--+
--+ +-- Utterbackia_peninsularis **
|
| +----- Prolasmidonta
+--+
| +-- Platynaias
+--+
+-- Alasmidonta
Amaro et al., 2017 [Publ 3791], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 14 | 3 | 3 | -- |
+-- Lasmigona
+-----+
| +-- Platynaias
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
+--+
+-- Utterbackia_imbecillis **
Aunins et al., 2018 [Publ 3863], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 38 | 3 | 3 | -- |
+-- Prolasmidonta
+-----+
| +-- Alasmidonta
--+
| +----- Pyganodon
+--+
| +-- Utterbackia_imbecillis **
+--+
+-- Utterbackia_peninsularis **
Huang et al., 2013 [Publ 3629], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 14 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Zhou et al., 2016 [Publ 3710], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 18 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 29 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 23 | 3 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 23 | 3 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Amaro et al., 2017 [Publ 3791], fig. 3A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 10 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Amaro et al., 2017 [Publ 3791], fig. 3B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 10 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 30 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 21 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Huang et al., 2018 [Publ 3828], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 40 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Cha et al., 2018 [Publ 3916], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 36 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Huang et al., 2019 [Publ 3843], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 54 | 2 | 2 | -- |
+-------- Lasmigona --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Wu et al., 2019 [Publ 3864], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 44 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Froufe et al., 2019 [Publ 3936], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 42 | 2 | 2 | -- |
+-------- Pterosyna --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Froufe et al., 2019 [Publ 3936], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 42 | 2 | 2 | -- |
+-------- Pterosyna --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Guerra et al., 2019 [Publ 4168], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 39 | 3 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Guerra et al., 2019 [Publ 4168], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 39 | 3 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Riccardi et al., 2020 [Publ 4089], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 17 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Wu et al., 2020 [Publ 4121], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 46 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 46 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Wu et al., 2022 [Publ 4238], fig. 2d
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 36 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Wu et al., 2022 [Publ 4238], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 71 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Dai et al., 2025 [Publ 4440], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 85 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Chen et al., 2025 [Publ 4448], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 83 | 2 | 2 | -- |
+-------- Platynaias --+ | +----- Pyganodon +--+ | +-- Utterbackia_imbecillis ** +--+ +-- Utterbackia_peninsularis **
Wu et al., 2016 [Publ 3711], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 19 | 1 | 1 | -- |
+----- Pyganodon --+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Wu et al., 2018 [Publ 4143], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 20 | 1 | 1 | -- |
+----- Pyganodon --+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Wu et al., 2020 [Publ 4147], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 28 | 1 | 1 | -- |
+----- Pyganodon --+ | +-- Utterbackia_peninsularis ** +--+ +-- Utterbackia_imbecillis **
Whelan et al., 2023 [Publ 4287], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 27 | 26 | 11 | -- |
+-------------------- Utterbackiana | | +-- Utterbackia_peninsularis ** --+ +--------------+ | | +-- Pyganodon | | +--+ +----- Alasmidonta | +--------+ | | | +-- Arcidens | | +--+ +--+ +-- Pseudodontoideus | | +----------- Alasminota | | +--+ +-- Anodontoides | +-----+ | | +-- Strophitus +--+ | +----- Sulcularia +--+ | +-- Platynaias +--+ +-- Prolasmidonta
Whelan et al., 2023 [Publ 4287], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 26 | 24 | 10 | -- |
+-------------------- Sulcularia | | +-------- Utterbackiana --+ +--------+ | | | +----- Alasminota | | +--+ | | | +-- Arcidens +--+ +--+ | +-- Pseudodontoideus | | +-------------- Alasmidonta +--+ | +----------- Platynaias +--+ | +-------- Strophitus +--+ | +----- Pyganodon +--+ | +-- Utterbackia_peninsularis ** +--+ +-- Anodontoides
Boyer et al., 2011 [Publ 2718], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 37 | 8 | 8 | -- |
+-------------- Strophitus | | +----- Pterosyna --+ +-----+ | | | +-- Lasmigona | | +--+ +--+ +-- Arcidens | | +-------- Utterbackia_imbecillis ** | | +--+-------- Alasmidonta | | +----- Pyganodon +--+ | +-- Anodontoides +--+ +-- Platynaias
Krebs et al., 2003 [Publ 2691], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.889 | 1 | 1 | 0 | 1 | 32 | 8 | 7 | -- |
+----- Utterbackia_imbecillis ** --+ | +-- Pyganodon +--+ +-- Anodonta
Smith et al., 2018 [Publ 3790], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 17 | 16 | 7 | -- |
+-- Utterbackia_peggyae **
+--------+
| +-- Pyganodon
|
--+ +-- Anodontoides
| +-----+
| | +-- Strophitus
| |
+--+ +-- Alasmidonta
| +--+
| | +-- Alasminota
+--+
| +-- Pseudodontoideus
+--+
+-- Utterbackiana
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 26 | 5 | 4 | -- |
+-- Utterbackia_imbecillis **
+-----+
| +-- Utterbackiana
--+
| +----- Strophitus
+--+
| +-- Lasmigona
+--+
+-- Alasmidonta
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.833 | 1 | 1 | 0 | 1 | 27 | 4 | 4 | -- |
+----- Utterbackia_imbecillis ** --+ | +-- Utterbackiana +--+ +-- Anodonta
Huang et al., 2002 [Publ 2657], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 33 | 4 | 4 | -- |
+----------- Utterbackiana --+ | +-------- Utterbackia_imbecillis ** +--+ | +----- Strophitus +--+ | +-- Alasmidonta +--+ +-- Lasmigona
Huang et al., 2002 [Publ 2657], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.833 | 1 | 1 | 0 | 1 | 33 | 4 | 4 | -- |
+----- Utterbackia_imbecillis ** | --+----- Utterbackiana | | +-- Alasmidonta +--+ +-- Lasmigona
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 53 | 3 | 3 | -- |
+-- Pyganodon
+--+
| +-- Utterbackia_peninsularis **
--+
| +-- Alasmidonta
+--+
+-- Prolasmidonta
Amaro et al., 2017 [Publ 3791], fig. 2-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.750 | 1 | 1 | 0 | 1 | 9 | 2 | 2 | -- |
+-- Lasmigona | --+-- Sinanodonta | +-- Utterbackia_peninsularis **
Dai et al., 2023 [Publ 4285], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 2 | 2 | -- |
+----- Platynaias --+ | +-- Pyganodon +--+ +-- Utterbackia_imbecillis **
Dai et al., 2024 [Publ 4358], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 2 | 2 | -- |
+----- Platynaias --+ | +-- Pyganodon +--+ +-- Utterbackia_imbecillis **
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.333 | 1 | 1 | 0 | 1 | 18 | 2 | 1 | -- |
+-- Pyganodon_cataracta | +-- Pyganodon_grandis | +-- Utterbackia_imbecillis ** | +-- Lampsilis_teres | +-- Lampsilis_straminea --+ +-- Megalonaias | +-- Obliquaria | +-- Theliderma | +-- Quadrula | +-- Uniomerus
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.333 | 1 | 1 | 0 | 1 | 21 | 2 | 1 | -- |
+-- Gonidea | +-- Pyganodon_grandis | +-- Utterbackia_imbecillis ** | +-- Pyganodon_cataracta | +-- Lampsilis_straminea | +-- Lampsilis_teres | +-- Megalonaias | +-- Obliquaria | --+-- Theliderma | +-- Quadrula | +-- Uniomerus | +-- Amblema | +-- Elliptio | +-- Fusconaia | +-- Plectomerus | +-- Pleurobema | +-- Unio
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.333 | 1 | 1 | 0 | 1 | 21 | 2 | 1 | -- |
+-- Gonidea | +-- Pyganodon_grandis | +-- Utterbackia_imbecillis ** | +-- Pyganodon_cataracta | +-- Lampsilis_straminea | +-- Lampsilis_teres | +-- Megalonaias | +-- Obliquaria | --+-- Theliderma | +-- Quadrula | +-- Uniomerus | +-- Amblema | +-- Elliptio | +-- Fusconaia | +-- Plectomerus | +-- Scriptolamprotula | +-- Unio
Huang et al., 2013 [Publ 3629], fig. 3-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- |
+-- Pyganodon --+ +-- Utterbackia_peninsularis **
Froufe et al., 2015 [Publ 3664], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Froufe et al., 2015 [Publ 3664], fig. 1-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 7 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Fonseca et al., 2016 [Publ 3775], fig. 1-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 8 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Feng et al., 2016 [Publ 3800], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 13 | 1 | 1 | -- |
+-- Anodonta --+ +-- Utterbackia_peninsularis **
Guerra et al., 2017 [Publ 3772], fig. 2a-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 7 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Guerra et al., 2017 [Publ 3772], fig. 2b-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 7 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 12 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Burzynski et al., 2017 [Publ 3774], fig. 1-top
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 12 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- |
+-- Pyganodon --+ +-- Utterbackia_peninsularis **
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Wen et al., 2017 [Publ 3795], fig. 3-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- |
+-- Pyganodon --+ +-- Utterbackia_peninsularis **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- |
+-- Pyganodon --+ +-- Utterbackia_peninsularis **
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 16 | 1 | 1 | -- |
+-- Pyganodon --+ +-- Utterbackia_peninsularis **
Chase et al., 2018 [Publ 3861], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 10 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Chase et al., 2018 [Publ 3861], fig. 1-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 10 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Burzynski & Soroka, 2018 [Publ 3862], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 3 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- |
+-- Pyganodon --+ +-- Utterbackia_peninsularis **
Froufe et al., 2019 [Publ 3936], fig. 3-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Froufe et al., 2019 [Publ 3936], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Smith, 2021 [Publ 4331], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- |
+-- Pyganodon --+ +-- Utterbackia_peninsularis **
Wu et al., 2022 [Publ 4238], fig. 2e
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 17 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Dai et al., 2024 [Publ 4386], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- |
+-- Pyganodon --+ +-- Utterbackia_peninsularis **
Wu et al., 2024 [Publ 4388], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 1 | 1 | -- |
+-- Alasmidonta --+ +-- Utterbackia_imbecillis **
Klymus et al., 2025 [Publ 4814], fig. 3A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Klymus et al., 2025 [Publ 4814], fig. 3B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Utterbackia_peninsularis ** --+ +-- Pyganodon
Campbell & Lydeard, 2026 [Publ 4810], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- |
+-- Pyganodon --+ +-- Utterbackia_peninsularis **
Mulvey et al., 1997 [Publ 3390], fig. 1_3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- |
+----- Utterbackia_imbecillis ** --+ | +-- Megalonaias +--+ +-- Amblema
Campbell, 2000 [Publ 2724], fig. 1-5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- |
+-- Utterbackia_imbecillis ** --+ +-- Elliptio
Chase et al., 2018 [Publ 3847], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
+-- Anodonta --+ +-- Utterbackia_peninsularis **
