Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Quadrulini

genus Uniomerus Conrad, 1853: cladomics

4 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (5 total)
analyzed in 51 cladograms in 35 publications
maximum number of terminals in an analysis = 4 (mean = 1.2, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 6
maximum number of included terminals = 4, (mean = 2.67 median = 2.5)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Uniomerus cladogram distances
PCoA Ordination

NJ Tree of Uniomerus cladogram distances
NJ Tree

Cladograms

Neemuchwala et al., 2023 [Publ 4292], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 23 18 2 **
Cladomics cladogram Megalonaias Uniomerus carolinianus Uniomerus columbensis Uniomerus tetralasmus Uniomerus declivis Theliderma Cyclonaias Quadrula Pustulosa

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 3 3 0 3 25 15 2 2.0
Cladomics cladogram Megalonaias Uniomerus carolinianus Uniomerus declivis Uniomerus tetralasmus Quadrula quadrula Quadrula verrucosa Quadrula nobilis Theliderma Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa Pustulosa nodulata Pustulosa necki Pustulosa petrina

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.625 3 3 2 3 630 18 2 2.0
Cladomics cladogram Megalonaias Uniomerus carolinianus Uniomerus declivis 2 Uniomerus declivis 1 Uniomerus tetralasmus Theliderma johnsoni Theliderma metanevra Quadrula nobilis Quadrula verrucosa Theliderma cylindrica Quadrula fragosa Quadrula quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa Pustulosa nodulata Pustulosa necki Pustulosa petrina

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.714 2 2 0 2 71 9 1 --
Cladomics cladogram Uniomerus declivis Uniomerus tetralasmus Theliderma Quadrula verrucosa Quadrula quadrula Quadrula nobilis Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

cst iT tT bi riS oT roT roS iD
0.714 2 2 0 2 71 9 1 --
Cladomics cladogram Uniomerus declivis Uniomerus tetralasmus Quadrula verrucosa Quadrula nobilis Quadrula quadrula Theliderma Pustulosa infucata Pustulosa kleiniana Cyclonaias Pustulosa pustulosa Pustulosa succissa

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

cst iT tT bi riS oT roT roS iD
0.714 2 2 0 2 103 5 1 --
Cladomics cladogram Uniomerus declivis Uniomerus tetralasmus Cyclonaias Pustulosa pustulosa 1 Theliderma Pustulosa pustulosa 2 Quadrula quadrula Quadrula verrucosa

Rosenberg et al., 1994 [Publ 2719], fig. 3

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 18 3 2 --
Cladomics cladogram Pyganodon cataracta Pyganodon grandis Utterbackia Lampsilis teres Lampsilis straminea Megalonaias Obliquaria Theliderma Quadrula Uniomerus tetralasmus

Lydeard et al., 1996 [Publ 2656], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 26 3 2 --
Cladomics cladogram Eurynia Pleurobema Uniomerus carolinianus

Lydeard et al., 1996 [Publ 2656], fig. 4

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 27 3 2 --
Cladomics cladogram Uniomerus carolinianus Eurynia Pleurobema

Rosenberg et al., 1997 [Publ 2720], fig. 2A

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 21 3 2 --
Cladomics cladogram Gonidea Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus tetralasmus Amblema Elliptio Fusconaia Plectomerus Pleurobema Unio

Rosenberg et al., 1997 [Publ 2720], fig. 2B

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 21 3 2 --
Cladomics cladogram Gonidea Pyganodon grandis Utterbackia Pyganodon cataracta Lampsilis straminea Lampsilis teres Megalonaias Obliquaria Theliderma Quadrula Uniomerus tetralasmus Amblema Elliptio Fusconaia Plectomerus Scriptolamprotula Unio

Krebs et al., 2003 [Publ 2691], fig. 3

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 32 3 2 --
Cladomics cladogram Uniomerus carolinianus Fusconaia Eurynia

Campbell et al., 2005 [Publ 2660], fig. 1

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 101 10 2 --
Cladomics cladogram Megalonaias Uniomerus declivis Theliderma Quadrula Pustulosa kieneriana Cyclonaias Pustulosa succissa Pustulosa infucata Pustulosa kleiniana

Campbell et al., 2005 [Publ 2660], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 101 10 2 --
Cladomics cladogram Megalonaias Uniomerus declivis Theliderma Quadrula quadrula Quadrula nobilis Quadrula verrucosa Pustulosa kieneriana Cyclonaias Pustulosa succissa Pustulosa infucata Pustulosa kleiniana

Pfeiffer et al., 2018 [Publ 3835], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 3 2 --
Cladomics cladogram Quadrula Uniomerus declivis Megalonaias

Johnson et al., 2018 [Publ 3844], fig. 2

cst iT tT bi riS oT roT roS iD
0.875 1 1 0 1 15 13 2 --
Cladomics cladogram Megalonaias Uniomerus tetralasmus Theliderma Quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Pustulosa pustulosa Pustulosa succissa Cyclonaias Pustulosa necki Pustulosa nodulata Pustulosa petrina

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 200 3 2 --
Cladomics cladogram Quadrula Megalonaias Uniomerus carolinianus

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 198 3 2 --
Cladomics cladogram Quadrula Megalonaias Uniomerus carolinianus

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 200 3 2 --
Cladomics cladogram Chamberlainia Megalonaias Uniomerus carolinianus Quadrula Sinohyriopsis Lamprotula Potomida Pronodularia Indopseudodon Nyeinchanconcha Thaiconcha Sundadontina cumingii Sundadontina tumida 1 Sundadontina tumida 2 Pseudodon Pilsbryoconcha Bineurus Sinosolenaia carinata Gonidea Leguminaia Microcondylaea Ptychorhynchus Parvasolenaia Koreosolenaia Sinosolenaia recognita Sinosolenaia iridinea Sinosolenaia oleivora Physunio modelli Physunio Physunio superbus Yaukthwa Lens Hyriopsis desowitzi Hyriopsis myersiana Hyriopsis panhai Hyriopsis bialata Hyriopsis khoratensis Rectidens Ensidens Amblema Toxolasma Obliquaria Plectomerus Elliptoideus Elliptio Eurynia Fusconaia Pleurobema Ellipsaria Potamilus leptodon Medionidus Potamilus alatus Ptychobranchus Truncilla Venustaconcha Cambarunio Leaunio Ortmanniana Lampsilis Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus Unio Scriptolamprotula Lanceolaria Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Pfeiffer et al., 2019 [Publ 3866], fig. 1

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 30 4 2 --
Cladomics cladogram Megalonaias Uniomerus declivis Quadrula verrucosa Cyclonaias Quadrula quadrula

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 28 18 2 --
Cladomics cladogram Uniomerus tetralasmus Theliderma Quadrula Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Pustulosa pustulosa Pustulosa succissa Cyclonaias Pustulosa necki Pustulosa petrina Pustulosa nodulata

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 25 15 2 --
Cladomics cladogram Megalonaias Uniomerus tetralasmus Theliderma Quadrula quadrula Quadrula verrucosa Quadrula nobilis Pustulosa kieneriana Pustulosa infucata Pustulosa kleiniana Pustulosa pustulosa Pustulosa succissa Cyclonaias Pustulosa necki Pustulosa petrina Pustulosa nodulata

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 33 1 1 --
Cladomics cladogram Uniomerus carolinianus Pleurobema

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 33 1 1 --
Cladomics cladogram Uniomerus carolinianus Pleurobema

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

cst iT tT bi riS oT roT roS iD
0.833 1 1 0 1 42 6 1 --
Cladomics cladogram Uniomerus declivis Pustulosa infucata Pustulosa kleiniana Quadrula Cyclonaias Pustulosa succissa Pustulosa pustulosa

Pfeiffer et al., 2016 [Publ 3660], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 37 5 1 --
Cladomics cladogram Uniomerus declivis Quadrula quadrula Quadrula verrucosa Pustulosa pustulosa Pustulosa petrina Cyclonaias

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Burlakova et al., 2019 [Publ 3899], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 32 1 1 --
Cladomics cladogram Quadrula Uniomerus declivis

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 53 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 1 1 --
Cladomics cladogram Uniomerus tetralasmus Quadrula

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 1 1 --
Cladomics cladogram Uniomerus tetralasmus Megalonaias

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 1 1 --
Cladomics cladogram Uniomerus tetralasmus Quadrula

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 1 1 --
Cladomics cladogram Uniomerus tetralasmus Quadrula

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 1 1 --
Cladomics cladogram Uniomerus tetralasmus Quadrula

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 14 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 139 1 1 --
Cladomics cladogram Quadrula Uniomerus tetralasmus

Kandl et al., 2001 [Publ 3384], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Elliptio Uniomerus carolinianus Pleurobema

Campbell et al., 2008 [Publ 2675], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram Uniomerus declivis Plectomerus Cyrtonaias Eurynia Fusconaia cerina Fusconaia flava Hemistena Plethobasus Parvaspina Pleurobema cordatum Pleuronaia Pleurobema oviforme 2 Pleurobema stabile Elliptio Pleurobema decisum Pleurobema sintoxia Pleurobema clava Pleurobema oviforme 1 Pleurobema beadleianum Pleurobema pyriforme Pleurobema rubellum Pleurobema hanleyianum 2 Pleurobema georgianum 2 Pleurobema hanleyianum 1 Pleurobema georgianum 1 Pleurobema athearni

Campbell et al., 2008 [Publ 2675], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 0 0 --
Cladomics cladogram Uniomerus declivis Plectomerus Elliptio arctata Elliptio crassidens Pleurobema decisum Pleurobema rubellum Pleurobema georgianum 2 Pleurobema stabile Pleurobema pyriforme Pleurobema oviforme 2 Pleurobema hanleyianum 2 Pleuronaia Pleurobema georgianum 3 Pleurobema cordatum Pleurobema beadleianum Hemistena Fusconaia flava Fusconaia cerina Eurynia Elliptio arca Cyrtonaias Plethobasus Parvaspina Pleurobema sintoxia Pleurobema oviforme 1 Pleurobema clava Pleurobema athearni Pleurobema georgianum 1 Pleurobema hanleyianum 1

Perkins et al., 2017 [Publ 3777], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 23 0 0 --
Cladomics cladogram Plectomerus Uniomerus declivis Parvaspina Elliptio arca Hemistena Elliptio arctata Elliptio crassidens Eurynia Pleurobema oviforme Pleurobema clava Pleurobema pyriforme Pleurobema georgianum Pleurobema beadleianum Pleurobema hanleyianum Pleurobema decisum Pleurobema sintoxia Pleurobema rubellum Elliptio spinosa Fusconaia escambia Fusconaia flava Fusconaia cerina Pleuronaia