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Data last updated
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genus Uniomerus Conrad, 1853: cladomics

4 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (5 total)
analyzed in 49 cladograms in 34 publications
maximum number of terminals in an analysis = 4 (mean = 1.16, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 5
maximum number of included terminals = 4, (mean = 2.6 median = 2)
proportion of cladograms that support basic monophyly = 1

Cladograms

Neemuchwala et al., 2023 [Publ 4292], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	23	18	2	0

  +-------------- Megalonaias           
  |                                     
  |     +-------- Uniomerus_carolinianus **
--+  +--+                               
  |  |  |  +----- Uniomerus_columbensis ** 
  |  |  +--+                            
  |  |     |  +-- Uniomerus_tetralasmus ** 
  +--+     +--+                         
     |        +-- Uniomerus_declivis **    
     |                                  
     |  +-------- Theliderma            
     +--+                               
        |  +----- Cyclonaias            
        +--+                            
           |  +-- Quadrula              
           +--+                         
              +-- Pustulosa

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	0	3	25	15	2	2

                       +----- Megalonaias           
                       |                            
  +--------------------+  +-- Uniomerus_carolinianus **
  |                    |  |                         
  |                    +--+-- Uniomerus_declivis **    
  |                       |                         
--+                       +-- Uniomerus_tetralasmus ** 
  |                                                 
  |  +----------------------- Quadrula_quadrula     
  |  |                                              
  |  |                    +-- Quadrula_verrucosa    
  +--+  +-----------------+                         
     |  |                 +-- Quadrula_nobilis      
     |  |                                           
     +--+  +----------------- Theliderma            
        |  |                                        
        |  |           +----- Pustulosa_kieneriana  
        +--+  +--------+                            
           |  |        |  +-- Pustulosa_infucata    
           |  |        +--+                         
           +--+           +-- Pustulosa_kleiniana   
              |                                     
              |  +----------- Cyclonaias            
              |  |                                  
              +--+        +-- Pustulosa_pustulosa   
                 |  +-----+                         
                 |  |     +-- Pustulosa_succissa    
                 +--+                               
                    |  +----- Pustulosa_nodulata    
                    +--+                            
                       |  +-- Pustulosa_necki       
                       +--+                         
                          +-- Pustulosa_petrina

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	71	9	1	--

                 +-- Uniomerus_declivis **   
  +--------------+                        
  |              +-- Uniomerus_tetralasmus **
  |                                       
--+              +-- Quadrula_verrucosa   
  |  +-----------+                        
  |  |           +-- Quadrula_nobilis     
  |  |                                    
  +--+  +----------- Quadrula_quadrula    
     |  |                                 
     |  +----------- Theliderma           
     +--+                                 
        |        +-- Pustulosa_infucata   
        |  +-----+                        
        |  |     +-- Pustulosa_kleiniana  
        +--+                              
           |  +----- Cyclonaias           
           +--+                           
              |  +-- Pustulosa_pustulosa  
              +--+                        
                 +-- Pustulosa_succissa

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	71	9	1	--

                    +-- Uniomerus_declivis **   
  +-----------------+                        
  |                 +-- Uniomerus_tetralasmus **
--+                                          
  |  +----------------- Theliderma           
  |  |                                       
  +--+  +-------------- Quadrula_verrucosa   
     |  |                                    
     |  |           +-- Quadrula_quadrula    
     +--+  +--------+                        
        |  |        +-- Quadrula_nobilis     
        |  |                                 
        +--+        +-- Pustulosa_infucata   
           |  +-----+                        
           |  |     +-- Pustulosa_kleiniana  
           +--+                              
              |  +----- Cyclonaias           
              +--+                           
                 |  +-- Pustulosa_pustulosa  
                 +--+                        
                    +-- Pustulosa_succissa

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	102	5	1	--

     +-- Uniomerus_declivis **    
  +--+                         
  |  +-- Uniomerus_tetralasmus ** 
  |                            
  |  +-- Cyclonaias            
--+  |                         
  |  +-- Pustulosa_pustulosa__1
  |  |                         
  |  +-- Theliderma            
  +--+                         
     +-- Pustulosa_pustulosa__2
     |                         
     +-- Quadrula_quadrula     
     |                         
     +-- Quadrula_verrucosa

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	25	15	2	--

  +----------------------------- Megalonaias          
  |                                                   
--+  +-------------------------- Uniomerus_tetralasmus **
  |  |                                                
  +--+  +----------------------- Theliderma           
     |  |                                             
     +--+  +-------------------- Quadrula_quadrula    
        |  |                                          
        |  |                 +-- Quadrula_verrucosa   
        +--+  +--------------+                        
           |  |              +-- Quadrula_nobilis     
           |  |                                       
           |  |           +----- Pustulosa_kieneriana 
           +--+  +--------+                           
              |  |        |  +-- Pustulosa_infucata   
              |  |        +--+                        
              |  |           +-- Pustulosa_kleiniana  
              +--+                                    
                 |           +-- Pustulosa_pustulosa  
                 |  +--------+                        
                 |  |        +-- Pustulosa_succissa   
                 +--+                                 
                    |  +-------- Cyclonaias           
                    +--+                              
                       |  +----- Pustulosa_necki      
                       +--+                           
                          |  +-- Pustulosa_petrina    
                          +--+                        
                             +-- Pustulosa_nodulata

Pfeiffer et al., 2018 [Publ 3835], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	3	3	2	--

  +----- Quadrula          
--+                        
  |  +-- Uniomerus_declivis **
  +--+                     
     +-- Megalonaias

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	1	1	--

  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	198	3	2	--

  +----- Quadrula              
--+                            
  |  +-- Megalonaias           
  +--+                         
     +-- Uniomerus_carolinianus **

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	1	1	--

  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	200	3	2	--

-- Uniomerus_carolinianus **

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Huang et al., 2002 [Publ 2657], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	1	1	--

-- Uniomerus_carolinianus **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	28	18	2	--

  +-------------------------- Uniomerus_tetralasmus **
  |                                                
--+  +----------------------- Theliderma           
  |  |                                             
  +--+  +-------------------- Quadrula             
     |  |                                          
     +--+  +----------------- Pustulosa_kieneriana 
        |  |                                       
        |  |              +-- Pustulosa_infucata   
        +--+  +-----------+                        
           |  |           +-- Pustulosa_kleiniana  
           |  |                                    
           +--+           +-- Pustulosa_pustulosa  
              |  +--------+                        
              |  |        +-- Pustulosa_succissa   
              +--+                                 
                 |  +-------- Cyclonaias           
                 +--+                              
                    |  +----- Pustulosa_necki      
                    +--+                           
                       |  +-- Pustulosa_petrina    
                       +--+                        
                          +-- Pustulosa_nodulata

Smith, 2021 [Publ 4331], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	1	1	--

  +-- Uniomerus_tetralasmus **
--+                        
  +-- Megalonaias

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	87	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	200	3	2	--

  +----- Quadrula              
--+                            
  |  +-- Megalonaias           
  +--+                         
     +-- Uniomerus_carolinianus **

Rosenberg et al., 1994 [Publ 2719], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	18	3	2	--

-- Uniomerus_tetralasmus **

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	53	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	53	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Campbell et al., 2005 [Publ 2660], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	101	10	2	--

              +-- Megalonaias         
  +-----------+                       
  |           +-- Uniomerus_declivis **  
--+                                   
  |  +----------- Theliderma          
  |  |                                
  +--+  +-------- Quadrula            
     |  |                             
     +--+  +----- Pustulosa_kieneriana
        |  |                          
        |  |  +-- Cyclonaias          
        +--+--+                       
           |  +-- Pustulosa_succissa  
           |                          
           |  +-- Pustulosa_infucata  
           +--+                       
              +-- Pustulosa_kleiniana

Pfeiffer et al., 2019 [Publ 3866], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	4	2	--

  +----------- Megalonaias       
--+                              
  |  +-------- Uniomerus_declivis **
  +--+                           
     |  +----- Quadrula_verrucosa
     +--+                        
        |  +-- Cyclonaias        
        +--+                     
           +-- Quadrula_quadrula

Burlakova et al., 2019 [Publ 3899], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	32	1	1	--

  +-- Quadrula          
--+                     
  +-- Uniomerus_declivis **

Huang et al., 2019 [Publ 3843], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Pfeiffer et al., 2016 [Publ 3660], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	5	1	--

  +-------------- Uniomerus_declivis ** 
--+                                  
  |  +----------- Quadrula_quadrula  
  +--+                               
     |  +-------- Quadrula_verrucosa 
     +--+                            
        |  +----- Pustulosa_pustulosa
        +--+                         
           |  +-- Pustulosa_petrina  
           +--+                      
              +-- Cyclonaias

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Lydeard et al., 1996 [Publ 2656], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	3	2	--

-- Uniomerus_carolinianus **

Perkins et al., 2017 [Publ 3777], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	23	0	0	--

-- Uniomerus_declivis **

Huang et al., 2002 [Publ 2657], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	1	1	--

-- Uniomerus_carolinianus **

Campbell et al., 2008 [Publ 2675], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Uniomerus_declivis **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	85	1	1	--

  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	42	6	1	--

  +-------------- Uniomerus_declivis ** 
  |                                  
--+           +-- Pustulosa_infucata 
  |  +--------+                      
  |  |        +-- Pustulosa_kleiniana
  +--+                               
     |  +-------- Quadrula           
     +--+                            
        |  +----- Cyclonaias         
        +--+                         
           |  +-- Pustulosa_succissa 
           +--+                      
              +-- Pustulosa_pustulosa

Kandl et al., 2001 [Publ 3384], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Uniomerus_carolinianus **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	3	2	--

-- Uniomerus_tetralasmus **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	1	1	--

  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula

Rosenberg et al., 1997 [Publ 2720], fig. 2A

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	3	2	--

-- Uniomerus_tetralasmus **

Campbell et al., 2008 [Publ 2675], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Uniomerus_declivis **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	18	1	1	--

  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula

Krebs et al., 2003 [Publ 2691], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	32	3	2	--

-- Uniomerus_carolinianus **

Lydeard et al., 1996 [Publ 2656], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	26	3	2	--

-- Uniomerus_carolinianus **

Campbell et al., 2005 [Publ 2660], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	101	10	2	--

  +-------------------- Megalonaias         
  |                                         
--+  +----------------- Uniomerus_declivis **  
  |  |                                      
  +--+  +-------------- Theliderma          
     |  |                                   
     +--+  +----------- Quadrula_quadrula   
        |  |                                
        |  |        +-- Quadrula_nobilis    
        +--+  +-----+                       
           |  |     +-- Quadrula_verrucosa  
           |  |                             
           +--+  +----- Pustulosa_kieneriana
              |  |                          
              |  |  +-- Cyclonaias          
              +--+--+                       
                 |  +-- Pustulosa_succissa  
                 |                          
                 |  +-- Pustulosa_infucata  
                 +--+                       
                    +-- Pustulosa_kleiniana

Johnson et al., 2018 [Publ 3844], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	15	13	2	--

  +-------------------------- Megalonaias          
  |                                                
--+  +----------------------- Uniomerus_tetralasmus **
  |  |                                             
  +--+  +-------------------- Theliderma           
     |  |                                          
     +--+  +----------------- Quadrula             
        |  |                                       
        |  |           +----- Pustulosa_kieneriana 
        +--+  +--------+                           
           |  |        |  +-- Pustulosa_infucata   
           |  |        +--+                        
           |  |           +-- Pustulosa_kleiniana  
           +--+                                    
              |           +-- Pustulosa_pustulosa  
              |  +--------+                        
              |  |        +-- Pustulosa_succissa   
              +--+                                 
                 |  +-------- Cyclonaias           
                 +--+                              
                    |  +----- Pustulosa_necki      
                    +--+                           
                       |  +-- Pustulosa_nodulata   
                       +--+                        
                          +-- Pustulosa_petrina

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