Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Quadrulini

genus Uniomerus Conrad, 1853: cladomics

4 Recent valid species (5 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (5 total)
analyzed in 49 cladograms in 34 publications
maximum number of terminals in an analysis = 4 (mean = 1.16, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 5
maximum number of included terminals = 4, (mean = 2.6 median = 2)
proportion of cladograms that support basic monophyly = 1

Cladograms

Neemuchwala et al., 2023 [Publ 4292], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4 4 2 4 23 18 2 0 
  +-------------- Megalonaias           
  |                                     
  |     +-------- Uniomerus_carolinianus **
--+  +--+                               
  |  |  |  +----- Uniomerus_columbensis ** 
  |  |  +--+                            
  |  |     |  +-- Uniomerus_tetralasmus ** 
  +--+     +--+                         
     |        +-- Uniomerus_declivis **    
     |                                  
     |  +-------- Theliderma            
     +--+                               
        |  +----- Cyclonaias            
        +--+                            
           |  +-- Quadrula              
           +--+                         
              +-- Pustulosa

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
3 3 0 3 25 15 2 2 
                       +----- Megalonaias           
                       |                            
  +--------------------+  +-- Uniomerus_carolinianus **
  |                    |  |                         
  |                    +--+-- Uniomerus_declivis **    
  |                       |                         
--+                       +-- Uniomerus_tetralasmus ** 
  |                                                 
  |  +----------------------- Quadrula_quadrula     
  |  |                                              
  |  |                    +-- Quadrula_verrucosa    
  +--+  +-----------------+                         
     |  |                 +-- Quadrula_nobilis      
     |  |                                           
     +--+  +----------------- Theliderma            
        |  |                                        
        |  |           +----- Pustulosa_kieneriana  
        +--+  +--------+                            
           |  |        |  +-- Pustulosa_infucata    
           |  |        +--+                         
           +--+           +-- Pustulosa_kleiniana   
              |                                     
              |  +----------- Cyclonaias            
              |  |                                  
              +--+        +-- Pustulosa_pustulosa   
                 |  +-----+                         
                 |  |     +-- Pustulosa_succissa    
                 +--+                               
                    |  +----- Pustulosa_nodulata    
                    +--+                            
                       |  +-- Pustulosa_necki       
                       +--+                         
                          +-- Pustulosa_petrina

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 71 9 1 -- 
                 +-- Uniomerus_declivis **   
  +--------------+                        
  |              +-- Uniomerus_tetralasmus **
  |                                       
--+              +-- Quadrula_verrucosa   
  |  +-----------+                        
  |  |           +-- Quadrula_nobilis     
  |  |                                    
  +--+  +----------- Quadrula_quadrula    
     |  |                                 
     |  +----------- Theliderma           
     +--+                                 
        |        +-- Pustulosa_infucata   
        |  +-----+                        
        |  |     +-- Pustulosa_kleiniana  
        +--+                              
           |  +----- Cyclonaias           
           +--+                           
              |  +-- Pustulosa_pustulosa  
              +--+                        
                 +-- Pustulosa_succissa

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iTtTbiriSoTroTroSiD
2 2 0 2 71 9 1 -- 
                    +-- Uniomerus_declivis **   
  +-----------------+                        
  |                 +-- Uniomerus_tetralasmus **
--+                                          
  |  +----------------- Theliderma           
  |  |                                       
  +--+  +-------------- Quadrula_verrucosa   
     |  |                                    
     |  |           +-- Quadrula_quadrula    
     +--+  +--------+                        
        |  |        +-- Quadrula_nobilis     
        |  |                                 
        +--+        +-- Pustulosa_infucata   
           |  +-----+                        
           |  |     +-- Pustulosa_kleiniana  
           +--+                              
              |  +----- Cyclonaias           
              +--+                           
                 |  +-- Pustulosa_pustulosa  
                 +--+                        
                    +-- Pustulosa_succissa

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
2 2 0 2 102 5 1 -- 
     +-- Uniomerus_declivis **    
  +--+                         
  |  +-- Uniomerus_tetralasmus ** 
  |                            
  |  +-- Cyclonaias            
--+  |                         
  |  +-- Pustulosa_pustulosa__1
  |  |                         
  |  +-- Theliderma            
  +--+                         
     +-- Pustulosa_pustulosa__2
     |                         
     +-- Quadrula_quadrula     
     |                         
     +-- Quadrula_verrucosa

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
-- Uniomerus_carolinianus **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 200 3 2 -- 
  +----- Quadrula              
--+                            
  |  +-- Megalonaias           
  +--+                         
     +-- Uniomerus_carolinianus **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 1 1 -- 
-- Uniomerus_carolinianus **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 28 18 2 -- 
  +-------------------------- Uniomerus_tetralasmus **
  |                                                
--+  +----------------------- Theliderma           
  |  |                                             
  +--+  +-------------------- Quadrula             
     |  |                                          
     +--+  +----------------- Pustulosa_kieneriana 
        |  |                                       
        |  |              +-- Pustulosa_infucata   
        +--+  +-----------+                        
           |  |           +-- Pustulosa_kleiniana  
           |  |                                    
           +--+           +-- Pustulosa_pustulosa  
              |  +--------+                        
              |  |        +-- Pustulosa_succissa   
              +--+                                 
                 |  +-------- Cyclonaias           
                 +--+                              
                    |  +----- Pustulosa_necki      
                    +--+                           
                       |  +-- Pustulosa_petrina    
                       +--+                        
                          +-- Pustulosa_nodulata

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Rosenberg et al., 1994 [Publ 2719], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 18 3 2 -- 
-- Uniomerus_tetralasmus **

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 27 3 2 -- 
-- Uniomerus_carolinianus **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 37 5 1 -- 
  +-------------- Uniomerus_declivis ** 
--+                                  
  |  +----------- Quadrula_quadrula  
  +--+                               
     |  +-------- Quadrula_verrucosa 
     +--+                            
        |  +----- Pustulosa_pustulosa
        +--+                         
           |  +-- Pustulosa_petrina  
           +--+                      
              +-- Cyclonaias

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
1 1 0 1 21 3 2 -- 
-- Uniomerus_tetralasmus **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 55 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 18 1 1 -- 
  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Kandl et al., 2001 [Publ 3384], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 5 0 0 -- 
-- Uniomerus_carolinianus **

Perkins et al., 2017 [Publ 3777], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 23 0 0 -- 
-- Uniomerus_declivis **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
1 1 0 1 200 3 2 -- 
-- Uniomerus_carolinianus **

Campbell et al., 2008 [Publ 2675], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Uniomerus_declivis **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 73 1 1 -- 
  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
1 1 0 1 198 3 2 -- 
  +----- Quadrula              
--+                            
  |  +-- Megalonaias           
  +--+                         
     +-- Uniomerus_carolinianus **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 25 15 2 -- 
  +----------------------------- Megalonaias          
  |                                                   
--+  +-------------------------- Uniomerus_tetralasmus **
  |  |                                                
  +--+  +----------------------- Theliderma           
     |  |                                             
     +--+  +-------------------- Quadrula_quadrula    
        |  |                                          
        |  |                 +-- Quadrula_verrucosa   
        +--+  +--------------+                        
           |  |              +-- Quadrula_nobilis     
           |  |                                       
           |  |           +----- Pustulosa_kieneriana 
           +--+  +--------+                           
              |  |        |  +-- Pustulosa_infucata   
              |  |        +--+                        
              |  |           +-- Pustulosa_kleiniana  
              +--+                                    
                 |           +-- Pustulosa_pustulosa  
                 |  +--------+                        
                 |  |        +-- Pustulosa_succissa   
                 +--+                                 
                    |  +-------- Cyclonaias           
                    +--+                              
                       |  +----- Pustulosa_necki      
                       +--+                           
                          |  +-- Pustulosa_petrina    
                          +--+                        
                             +-- Pustulosa_nodulata

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 42 6 1 -- 
  +-------------- Uniomerus_declivis ** 
  |                                  
--+           +-- Pustulosa_infucata 
  |  +--------+                      
  |  |        +-- Pustulosa_kleiniana
  +--+                               
     |  +-------- Quadrula           
     +--+                            
        |  +----- Cyclonaias         
        +--+                         
           |  +-- Pustulosa_succissa 
           +--+                      
              +-- Pustulosa_pustulosa

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 101 10 2 -- 
  +-------------------- Megalonaias         
  |                                         
--+  +----------------- Uniomerus_declivis **  
  |  |                                      
  +--+  +-------------- Theliderma          
     |  |                                   
     +--+  +----------- Quadrula_quadrula   
        |  |                                
        |  |        +-- Quadrula_nobilis    
        +--+  +-----+                       
           |  |     +-- Quadrula_verrucosa  
           |  |                             
           +--+  +----- Pustulosa_kieneriana
              |  |                          
              |  |  +-- Cyclonaias          
              +--+--+                       
                 |  +-- Pustulosa_succissa  
                 |                          
                 |  +-- Pustulosa_infucata  
                 +--+                       
                    +-- Pustulosa_kleiniana

Campbell et al., 2008 [Publ 2675], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 24 0 0 -- 
-- Uniomerus_declivis **

Johnson et al., 2018 [Publ 3844], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 15 13 2 -- 
  +-------------------------- Megalonaias          
  |                                                
--+  +----------------------- Uniomerus_tetralasmus **
  |  |                                             
  +--+  +-------------------- Theliderma           
     |  |                                          
     +--+  +----------------- Quadrula             
        |  |                                       
        |  |           +----- Pustulosa_kieneriana 
        +--+  +--------+                           
           |  |        |  +-- Pustulosa_infucata   
           |  |        +--+                        
           |  |           +-- Pustulosa_kleiniana  
           +--+                                    
              |           +-- Pustulosa_pustulosa  
              |  +--------+                        
              |  |        +-- Pustulosa_succissa   
              +--+                                 
                 |  +-------- Cyclonaias           
                 +--+                              
                    |  +----- Pustulosa_necki      
                    +--+                           
                       |  +-- Pustulosa_nodulata   
                       +--+                        
                          +-- Pustulosa_petrina

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iTtTbiriSoTroTroSiD
1 1 0 1 14 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Pfeiffer et al., 2018 [Publ 3835], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 3 3 2 -- 
  +----- Quadrula          
--+                        
  |  +-- Uniomerus_declivis **
  +--+                     
     +-- Megalonaias

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Uniomerus_tetralasmus **
--+                        
  +-- Megalonaias

Lydeard et al., 1996 [Publ 2656], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 26 3 2 -- 
-- Uniomerus_carolinianus **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
1 1 0 1 21 3 2 -- 
-- Uniomerus_tetralasmus **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 53 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 32 3 2 -- 
-- Uniomerus_carolinianus **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 101 10 2 -- 
              +-- Megalonaias         
  +-----------+                       
  |           +-- Uniomerus_declivis **  
--+                                   
  |  +----------- Theliderma          
  |  |                                
  +--+  +-------- Quadrula            
     |  |                             
     +--+  +----- Pustulosa_kieneriana
        |  |                          
        |  |  +-- Cyclonaias          
        +--+--+                       
           |  +-- Pustulosa_succissa  
           |                          
           |  +-- Pustulosa_infucata  
           +--+                       
              +-- Pustulosa_kleiniana

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 30 4 2 -- 
  +----------- Megalonaias       
--+                              
  |  +-------- Uniomerus_declivis **
  +--+                           
     |  +----- Quadrula_verrucosa
     +--+                        
        |  +-- Cyclonaias        
        +--+                     
           +-- Quadrula_quadrula

Burlakova et al., 2019 [Publ 3899], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 32 1 1 -- 
  +-- Quadrula          
--+                     
  +-- Uniomerus_declivis **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **