species Pustulosa nodulata (Rafinesque, 1820): synonymy

Obliquaria (Quadrula) nodulata Rafinesque, 1820

Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 307, pl. 81, figs. 17-18 [also as ‘Unio nodulata’].
Rafinesque, 1864. [in] Binney & Tryon, reprint: 51.
Type(s): LECTOTYPE ANSP_20225, Kentucky River.

Quadrula nodulata — Utterback (1915), Vanatta (1915), Ortmann & Walker (1922), Murray & Leonard (1962), Burch (1975), ... Boyer et al. (2011), ITIS (2012), Szumowski et al. (2012), Haag (2012), McMurray et al. (2012)
Quadrula (Bullata) nodulata — Frierson (1927)
Quadrula (s.s.) nodulata — F.C. Baker (1928)
Quadrula (Pustulosa) nodulata — Haas (1969)
Obliquaria nodulata — Burch (1975)
Amphinaias nodulata — Graf & Cummings (2007), Cummings & Graf (2010), IUCN (2011)
Cyclonaias nodulata — Williams et al. (2017), Burlakova et al. (2018), Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021), de Moulpied et al. (2022), Hopper et al. (2023)
Pustulosa nodulata — Neemuchwala et al. (2023), FMCS (2025), Campbell & Lydeard (2026)

Unio pustulatus Lea, 1831

Lea, 1831. Trans. Amer. Phil. Soc. 4 [O. 1]: 79 [89], pl. 7, fig. 9.
Type(s): SYNTYPE USNM_84200, Cincinnati, Ohio.

Quadrula (sec. Theliderma) pustulata — Simpson (1900), Simpson (1914)
Unio (s.s.) pustulatus — Call (1900)
Quadrula pustulata — Ortmann (1914)
Unio pustulatus, var. — de Gregorio (1914)
Quadrula nodulata — Utterback (1915), Vanatta (1915), Ortmann & Walker (1922), Murray & Leonard (1962), Burch (1975), R.I. Johnson (1980), Oesch (1984), Vidrine (1993), Howells et al. (1996), Strayer & Jirka (1997), Parmalee & Bogan (1998), Watters et al. (2009)
Quadrula (Bullata) nodulata — Frierson (1927)
Quadrula (s.s.) nodulata — F.C. Baker (1928)
Quadrula (Pustulosa) nodulata — Haas (1969)
Cyclonaias nodulata — Lopes-Lima et al. (2019)

Unio nodulatus ‘Rafinesque’ of Authors

(reference to Obliquaria (Quadrula) nodulata Rafinesque, 1820)
‘Rafinesque’ of Authors.
Say, 1834. Amer. Conch. 6: ??.
Conrad, 1837. Monogr. Fam. Unionidae 1(9): 80, pl. 45, fig. 1.
Reeve, 1864. Conch. Icon. 16: pl. 13, sp. 51.

Quadrula (sec. Theliderma) pustulata — Simpson (1900), Simpson (1914)
Quadrula nodulata — Utterback (1915), Watters et al. (2009)

Unio undulatus Lea, 1852

(error for Unio nodulatus ‘Rafinesque’ of Authors)
‘Con.’ Lea, 1852. Synopsis Fam. Naiades: 22.

Unio modulatus Küster, 1862

(error for Obliquaria (Quadrula) nodulata Rafinesque, 1820)
‘Raf.’ Küster, 1862. Conchyl. Cab. 9(2): 311.