MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Mon 29 June 2026
Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina
genus Alasmidonta Say, 1818: cladomics
11 Recent valid species (11 total species)
Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.
The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!
Cladomics Summary
number of species = 11 (11 total)
analyzed in 67 cladograms in 45 publications
maximum number of terminals in an analysis = 7 (mean = 1.46, median = 1)
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 9
maximum number of included terminals = 7, (mean = 4.44 median = 5)
proportion of cladograms that support basic monophyly = 0.89
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Cladograms
Whelan et al., 2023 [Publ 4287], fig. 2
** Top Monophyly Cladogram **
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 7 | 34 | 26 | 13 | ** |
+-- Utterbackia
+-----------------------+
| +-- Pyganodon
|
| +-------- Utterbackiana
| |
--+ +--------------+ +-- Lasmigona
| | | +--+
| | | | +-- Pseudodontoideus
| | +--+
| | | +-- Pterosyna
| | +--+
+--+ +-- Arcidens
|
| +----- Sulcularia
| +--------------+
| | | +-- Anodontoides
| | +--+
| | +-- Strophitus
+--+
| +-- Prolasmidonta
| +--------------+
| | +-- Platynaias
| |
+--+ +-------------- Alasminota
| |
| | +-- Alasmidonta_raveneliana **
+--+ +--------+
| | +-- Alasmidonta_marginata **
| |
+--+ +-- Alasmidonta_varicosa **
| +-----+
| | +-- Alasmidonta_uwharriensis **
+--+
| +----- Alasmidonta_undulata **
+--+
| +-- Alasmidonta_triangulata **
+--+
+-- Alasmidonta_arcula **
Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS
** Top Sister-Group Cladogram **
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 6 | 6 | 4 | 6 | 30 | 27 | 14 | 1.0 |
+-------------- Utterbackiana
+-----+
| | +----------- Alasminota
| +--+
| | +-------- Pseudodontoideus
| +--+
| | +----- Lasmigona
| +--+
| | +-- Pterosyna
| +--+
--+ +-- Arcidens
|
| +-- Anodontoides
| +-----+
| | +-- Strophitus
| +--------+
| | | +----- Sulcularia
| | +--+
| | | +-- Pyganodon
| | +--+
+--+ +-- Utterbackia
|
| +-- Prolasmidonta
| +-----------+
| | +-- Platynaias
| |
+--+ +----------- Ligodonta
| |
| | +-- Alasmidonta_marginata **
+--+ +-----+
| | +-- Alasmidonta_raveneliana **
| |
+--+ +-- Alasmidonta_varicosa **
| +--+
| | +-- Alasmidonta_uwharriensis **
+--+
| +-- Alasmidonta_arcula **
+--+
+-- Alasmidonta_triangulata **
Whelan et al., 2023 [Publ 4287], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 7 | 7 | 5 | 7 | 21 | 20 | 11 | 0.0 |
+----------------------- Utterbackiana | | +-- Utterbackia | +-----------------+ --+ | +-- Pyganodon | | | | +----------- Alasminota | | | | | +-----+ +-- Anodontoides +--+ | | +-----+ | | | | +-- Strophitus | | +--+ | | | +----- Sulcularia | | +--+ | | | +-- Platynaias +--+ +--+ | +-- Prolasmidonta | | +-- Arcidens | +-----------+ | | +-- Pseudodontoideus | | +--+ +-- Alasmidonta_marginata ** | +--------+ | | +-- Alasmidonta_raveneliana ** | | +--+ +-- Alasmidonta_varicosa ** | +-----+ | | +-- Alasmidonta_uwharriensis ** +--+ | +----- Alasmidonta_undulata ** +--+ | +-- Alasmidonta_triangulata ** +--+ +-- Alasmidonta_arcula **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.933 | 6 | 6 | 4 | 6 | 627 | 31 | 13 | 1.0 |
+-- Utterbackia
+--------------------+
| +-- Pyganodon
|
--+ +-------------------- Utterbackiana
| |
| | +----------------- Simpsonaias
+--+ |
| | +-- Prolasmidonta
| | +--+
+--+ | +-- Platynaias
| +--------+
| | | +-- Anodontoides
| | +--+
| | +-- Strophitus
+--+
| +----------- Alasminota
| |
| | +-- Alasmidonta_marginata **
+--+ +-----+
| | +-- Alasmidonta_raveneliana **
| |
+--+ +-- Alasmidonta_arcula **
| +--+
| | +-- Alasmidonta_undulata **
+--+
| +-- Alasmidonta_uwharriensis **
+--+
+-- Alasmidonta_varicosa **
Whelan et al., 2023 [Publ 4287], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 5 | 5 | 2 | 5 | 22 | 20 | 10 | 3.0 |
+-------------------- Sulcularia | | +-------- Utterbackiana | +--------+ | | | +----- Alasminota --+ | +--+ | | | +-- Arcidens | | +--+ | | +-- Pseudodontoideus | | +--+ +----- Alasmidonta_raveneliana ** | | | | +-- Alasmidonta_uwharriensis ** | +--------+--+ | | | +-- Alasmidonta_varicosa ** | | | | | | +-- Alasmidonta_arcula ** +--+ +--+ | +-- Alasmidonta_triangulata ** | | +----------- Platynaias +--+ | +-------- Strophitus +--+ | +----- Pyganodon +--+ | +-- Utterbackia +--+ +-- Anodontoides
Inoue et al., 2014 [Publ 3635], fig. 3A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 3 | 3 | 1 | 3 | 17 | 11 | 8 | 4.0 |
+-------- Pseudodontoideus
+--------+
| | +----- Lasmigona
| +--+
| | +-- Pterosyna
--+ +--+
| +-- Arcidens
|
| +-------------- Strophitus
| |
+--+ +-- Pyganodon
| +--------+
| | +-- Utterbackia
+--+
| +-------- Platynaias
+--+
| +----- Alasmidonta_marginata **
+--+
| +-- Alasmidonta_undulata **
+--+
+-- Alasmidonta_varicosa **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.455 | 2 | 20 | 0 | 2 | 103 | 14 | 9 | 24.0 |
+-- Alasmidonta_marginata **
|
+-- Prolasmidonta
|
+-- Alasmidonta_undulata **
|
+-- Beringiana
|
+-- Brachyanodon_kennerlyi
|
+-- Brachyanodon_nuttalliana
|
+-- Brachyanodon_oregonensis
|
+-- Arcidens_confragosus
|
+-- Arcidens_wheeleri
|
+-- Pterosyna
|
+--------------------------------------------------+-- Platynaias_compressa
| |
| +-- Lasmigona
| |
| +-- Platynaias_decorata
| |
| +-- Platynaias_subviridis
| |
| +-- Pyganodon_grandis
| |
| +-- Pyganodon_lacustris
| |
| +-- Pseudodontoideus
| |
| +-- Strophitus
| |
--+ +-- Utterbackia_imbecillis
| |
| +-- Utterbackia_peninsularis__1
| |
| +-- Utterbackia_peninsularis__2
|
| +-------------------------------------------------- Gonidea
| |
| | +----------------------------------------------- Reginaia
| | |
| | +----------------------------------------------- Amblema
| | |
| | | +----- Popenaias
| | | |
| | | | +-- Dromus
| | | | |
| | | | +-- Epioblasma
+--+ | | |
| +-----------------------------------------+ +-- Glebula
| | | |
| | | +-- Lampsilis_ovata
| | | |
| | | +-- Lampsilis_virescens
| | | |
| | | +-- Lemiox
| | +--+
| | +-- Obliquaria
| | |
+--+ +-- Obovaria_olivaria
| |
| +-- Obovaria_unicolor
| |
| +-- Plectomerus
| |
| +-- Potamilus_alatus
| |
| +-- Potamilus_capax
|
| +----- Uniomerus
| |
| | +-- Cyclonaias
| +--------------------------------------+ |
| | | +-- Pustulosa_pustulosa__1
| | | |
| | | +-- Theliderma
| | +--+
| | +-- Pustulosa_pustulosa__2
| | |
+--+ +-- Quadrula_quadrula
| |
| +-- Quadrula_verrucosa
|
| +----------------------------------------- Elliptoideus
| |
| | +-------------------------------------- Parvaspina
| | |
+--+ | +----------------------------------- Plethobasus
| | |
| | +----------------------------------- Eurynia
| | |
| | +----------------------------------- Hemistena
+--+ |
| +----------------------------------- Elliptio
| |
| | +-------------------------------- Pleuronaia
| | |
| | | +-- Pleurobema_cordatum
| | | +--------------------------+
+--+ | | +-- Fusconaia
| | |
| | | +-------------------------- Pleurobema_plenum
| | | |
| | | | +----------------------- Pleurobema_stabile
| | | | |
| | | | | +----- Pleurobema_pyriforme
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum__2
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni
| | | | | | |
+--+ | +--+ | | +----- Pleurobema_decisum
| | | | | |
| | | | +--+ +-- Pleurobema_georgianum__1
| | | | | |
| | | | +--+-- Pleurobema_hanleyianum__1
| | | | |
| | | | +-- Pleurobema_hanleyianum__2
| | | |
| | +--+ +-------------- Pleurobema_beadleianum__2
| | | |
| | | +--+ +----------- Pleurobema_strodeanum__2
| | | | | |
| | | | +--+----------- Pleurobema_rubellum
| | | | |
| | | | | +-------- Pleurobema_beadleianum__1
| | | | +--+
+--+ | | | +----- Pleurobema_taitianum
| | | +--+
| | | | +-- Pleurobema_perovatum
| +--+ +--+
| | +-- Pleurobema_strodeanum__1
| |
| | +----- Pleurobema_oviforme__6
| | |
| | +--+----- Pleurobema_clava__3
| | | |
| | | | +-- Pleurobema_oviforme__5
| | | +--+
| +--------+ +-- Pleurobema_clava__2
| |
| | +----- Pleurobema_oviforme__2
| | |
| | +----- Pleurobema_oviforme__3
| +--+
| +----- Pleurobema_oviforme__4
| |
| | +-- Pleurobema_clava__1
| +--+
| +-- Pleurobema_oviforme__1
|
| +----------------- Pleurobema_riddellii
| |
| | +-------------- Pleurobema_sintoxia__34
| | |
| | | +----------- Pleurobema_sintoxia__31
| | | |
+-----------+ | +----------- Pleurobema_sintoxia__32
| | |
| | +----------- Pleurobema_sintoxia__33
| | |
| | | +-------- Pleurobema__4
| | | |
| | | +-------- Pleurobema__5
+--+ | |
| | +-------- Pleurobema_sintoxia__2
| | |
| | +-------- Pleurobema_sintoxia__3
| | |
| | +-------- Pleurobema__8
| | |
| | +-------- Pleurobema__9
| | |
| | +-------- Pleurobema__10
| | |
| | +-------- Pleurobema_sintoxia__5
+--+ |
| +-------- Pleurobema_sintoxia__6
| |
| +-------- Pleurobema_sintoxia__7
| |
| +-------- Pleurobema_sintoxia__8
| |
| +-------- Pleurobema_sintoxia__9
| |
| +-------- Pleurobema_sintoxia__10
| |
| +-------- Pleurobema_sintoxia__11
| |
| +-------- Pleurobema_sintoxia__12
| |
| +-------- Pleurobema_sintoxia__13
| |
| +-------- Pleurobema_sintoxia__14
| |
| +-------- Pleurobema_sintoxia__15
| |
| +-------- Pleurobema_sintoxia__16
+--+
+-------- Pleurobema_sintoxia__17
|
+-------- Pleurobema_sintoxia__18
|
+-------- Pleurobema_sintoxia__19
|
+-------- Pleurobema_sintoxia__20
|
+-------- Pleurobema_sintoxia__21
|
+-------- Pleurobema_sintoxia__22
|
+-------- Pleurobema_sintoxia__23
|
+-------- Pleurobema_sintoxia__24
|
+-------- Pleurobema_sintoxia__25
|
+-------- Pleurobema_sintoxia__26
|
+-------- Pleurobema_sintoxia__27
|
+-------- Pleurobema_sintoxia__28
|
+-------- Pleurobema_sintoxia__29
|
+-------- Pleurobema_sintoxia__30
|
| +-- Pleurobema__6
+-----+
| +-- Pleurobema_sintoxia__1
|
| +-- Pleurobema__7
+-----+
| +-- Pleurobema_sintoxia__4
|
| +----- Pleurobema__3
+--+
| +-- Pleurobema__1
+--+
+-- Pleurobema__2
Breton et al., 2011 [Publ 3376], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 36 | 9 | 6 | -- |
+-- Pterosyna
+-----------+
| +-- Lasmigona
--+
| +----------- Strophitus
| |
+--+ +-- Pyganodon
| +-----+
| | +-- Utterbackia
+--+
| +----- Platynaias
+--+
| +-- Alasmidonta_marginata **
+--+
+-- Alasmidonta_undulata **
Chase et al., 2018 [Publ 3847], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 2 | 2 | 0 | 2 | 37 | 9 | 6 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
|
--+ +-- Lasmigona
| +-----+
| | +-- Pterosyna
| |
+--+ +-- Alasmidonta_undulata **
| +--+
| | +-- Alasmidonta_marginata **
+--+
| +-- Strophitus
+--+
+-- Platynaias
Boyer et al., 2011 [Publ 2718], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 37 | 8 | 8 | -- |
+-------------- Strophitus | | +----- Pterosyna --+ +-----+ | | | +-- Lasmigona | | +--+ +--+ +-- Arcidens | | +-------- Utterbackia | | +--+-------- Alasmidonta_marginata ** | | +----- Pyganodon +--+ | +-- Anodontoides +--+ +-- Platynaias
Krebs et al., 2003 [Publ 2691], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.889 | 1 | 1 | 0 | 1 | 32 | 8 | 7 | -- |
+----------- Utterbackiana | +----------- Platynaias | +----------- Alasmidonta_triangulata ** --+ | +-- Pterosyna +--------+ | +-- Lasmigona | | +-------- Strophitus +--+ | +----- Utterbackia +--+ | +-- Pyganodon +--+ +-- Anodonta
Chong et al., 2008 [Publ 2668], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.889 | 1 | 1 | 0 | 1 | 17 | 7 | 7 | -- |
+-- Pseudodontoideus | --+-- Alasmidonta_marginata ** | +-- Pterosyna
Smith et al., 2018 [Publ 3790], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 17 | 16 | 7 | -- |
+-- Utterbackia
+--------+
| +-- Pyganodon
|
--+ +-- Anodontoides
| +-----+
| | +-- Strophitus
| |
+--+ +-- Alasmidonta_triangulata **
| +--+
| | +-- Alasminota
+--+
| +-- Pseudodontoideus
+--+
+-- Utterbackiana
Boyer et al., 2011 [Publ 2718], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 34 | 5 | 5 | -- |
+----- Alasmidonta_marginata ** | +----- Pyganodon | --+ +-- Pterosyna +--+ | +-- Lasmigona | | +-- Platynaias +--+ +-- Strophitus
Huang et al., 2019 [Publ 3843], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 1 | 1 | 0 | 1 | 200 | 6 | 5 | -- |
+-------- Alasmidonta_marginata ** | +-------- Lasmigona | +-------- Pseudanodonta | +-------- Simpsonaias --+ +-------- Strophitus | | +-- Pyganodon +-----+ | +-- Utterbackia | | +----- Anodonta_anatina +--+ | +-- Anodonta_cygnea +--+ +-- Brachyanodon
Huang et al., 2019 [Publ 3843], fig. S3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 1 | 1 | 0 | 1 | 198 | 6 | 5 | -- |
+-- Alasmidonta_marginata ** --+ +-- Lasmigona
Huang et al., 2019 [Publ 3843], fig. S4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.857 | 1 | 1 | 0 | 1 | 200 | 6 | 5 | -- |
+-------- Acuticosta | +-------- Alasmidonta_marginata ** | +-------- Anodonta_anatina | +-------- Anemina | +-------- Anodonta_cygnea | +-------- Cristaria | +-------- Brachyanodon | +-------- Lasmigona | +-------- Lepidodesma | +-------- Pseudanodonta | +-------- Pyganodon | --+-------- Simpsonaias | +-------- Strophitus | +-------- Utterbackia | +-------- Schistodesmus | +-------- Unio | +-------- Scriptolamprotula | +-------- Lanceolaria | | +-- Pletholophus +-----+ | +-- Sinanodonta | | +----- Cuneopsis_celtiformis | | | +----- Tchangsinaia | | +--+----- Cuneopsis_rufescens | +----- Nodularia | | +-- Cuneopsis_szechenyii +--+ +-- Cuneopsis_heudei
Stewart et al., 2021 [Publ 4451], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 15 | 9 | 5 | -- |
+-- Alasmidonta_marginata **
+-----+
| +-- Pterosyna
|
--+ +-- Strophitus
| +--+
| | +-- Platynaias
+--+
| +-- Pyganodon
+--+
+-- Utterbackia
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 94 | 6 | 5 | -- |
+-- Pyganodon
+-----+
| +-- Utterbackia
|
--+ +-- Alasmidonta_varicosa **
| +--+
| | +-- Platynaias
+--+
| +-- Prolasmidonta
+--+
+-- Pterosyna
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 94 | 6 | 5 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 90 | 6 | 5 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 90 | 6 | 5 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Wu et al., 2024 [Publ 4379], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 6 | 5 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Hou et al., 2024 [Publ 4400], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 74 | 6 | 5 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Hou et al., 2025 [Publ 4445], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 51 | 6 | 5 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Platynaias
+--+
+-- Alasmidonta_varicosa **
Wang et al., 2025 [Publ 4488], fig. 3B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 63 | 6 | 5 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 139 | 7 | 5 | -- |
+-- Pyganodon
+-----+
| +-- Utterbackia
|
--+ +-- Pterosyna
| +--+
| | +-- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Wu et al., 2026 [Publ 4811], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 94 | 6 | 5 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Wu et al., 2026 [Publ 4825], fig. 6B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 64 | 6 | 5 | -- |
+-- Pyganodon
+--------+
| +-- Utterbackia
--+
| +-------- Pterosyna
+--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 26 | 5 | 4 | -- |
+-- Utterbackia
+-----+
| +-- Utterbackiana
--+
| +----- Strophitus
+--+
| +-- Lasmigona
+--+
+-- Alasmidonta_triangulata **
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.833 | 1 | 1 | 0 | 1 | 27 | 4 | 4 | -- |
+----- Strophitus --+ | +-- Lasmigona +--+ +-- Alasmidonta_triangulata **
Huang et al., 2002 [Publ 2657], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 33 | 4 | 4 | -- |
+----------- Utterbackiana --+ | +-------- Utterbackia +--+ | +----- Strophitus +--+ | +-- Alasmidonta_triangulata ** +--+ +-- Lasmigona
Huang et al., 2002 [Publ 2657], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.833 | 1 | 1 | 0 | 1 | 33 | 4 | 4 | -- |
+----- Utterbackia | --+----- Utterbackiana | | +-- Alasmidonta_triangulata ** +--+ +-- Lasmigona
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 80 | 4 | 4 | -- |
+-- Pyganodon
+-----+
| +-- Simpsonaias
--+
| +----- Strophitus
+--+
| +-- Alasmidonta_marginata **
+--+
+-- Lasmigona
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 138 | 4 | 4 | -- |
+-- Lasmigona
+-----+
| +-- Alasmidonta_marginata **
--+
| +----- Simpsonaias
+--+
| +-- Pyganodon
+--+
+-- Strophitus
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 138 | 4 | 4 | -- |
+----- Pyganodon | +----- Simpsonaias --+ +----- Strophitus | | +-- Alasmidonta_marginata ** +--+ +-- Lasmigona
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.833 | 1 | 1 | 0 | 1 | 124 | 4 | 4 | -- |
+-- Alasmidonta_marginata ** | +-- Anodonta_anatina | +-- Anodonta_cygnea | +-- Brachyanodon | +-- Lasmigona | +-- Pseudanodonta | +-- Pyganodon | +-- Simpsonaias | +-- Strophitus --+ +-- Sinanodonta_lauta | +-- Cristaria_plicata | +-- Cristaria_radiata | +-- Pletholophus | +-- Sinanodonta_lucida | +-- Sinanodonta_jourdyi | +-- Lanceolaria_gladiola | +-- Lanceolaria_grayii | +-- Lanceolaria_danzhouensis
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 81 | 4 | 4 | -- |
+----------- Pyganodon --+ | +-------- Simpsonaias +--+ | +----- Strophitus +--+ | +-- Alasmidonta_marginata ** +--+ +-- Lasmigona
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 121 | 4 | 4 | -- |
+----- Pyganodon | +----- Simpsonaias --+ +----- Strophitus | | +-- Alasmidonta_marginata ** +--+ +-- Lasmigona
Sano et al., 2020 [Publ 4138], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 68 | 5 | 4 | -- |
+-- Pyganodon
+-----+
| +-- Utterbackia
--+
| +----- Alasmidonta_varicosa **
+--+
| +-- Prolasmidonta
+--+
+-- Platynaias
Sano et al., 2020 [Publ 4138], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 68 | 5 | 4 | -- |
+-- Pyganodon
+-----+
| +-- Utterbackia
--+
| +----- Alasmidonta_varicosa **
+--+
| +-- Platynaias
+--+
+-- Prolasmidonta
Sano et al., 2020 [Publ 4138], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 68 | 5 | 4 | -- |
+-- Pyganodon
+-----+
| +-- Utterbackia
--+
| +----- Alasmidonta_varicosa **
+--+
| +-- Prolasmidonta
+--+
+-- Platynaias
Chen et al., 2020 [Publ 4170], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 0.667 | 1 | 1 | 0 | 1 | 33 | 5 | 4 | -- |
+-- Alasmidonta_varicosa ** --+ +-- Anemina
Wu et al., 2021 [Publ 4366], fig. 3-BI
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 72 | 6 | 4 | -- |
+-- Pyganodon
+-----+
| +-- Utterbackia
--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Wu et al., 2021 [Publ 4366], fig. 3-ML
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 72 | 6 | 4 | -- |
+-- Pyganodon
+-----+
| +-- Utterbackia
--+
| +----- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
+--+
+-- Platynaias
Wu et al., 2021 [Publ 4366], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 67 | 5 | 4 | -- |
+-- Pyganodon
+-----+
| +-- Utterbackia
--+
| +----- Prolasmidonta
+--+
| +-- Platynaias
+--+
+-- Alasmidonta_varicosa **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 75 | 4 | 4 | -- |
+-- Pyganodon
+-----+
| +-- Simpsonaias
--+
| +----- Strophitus
+--+
| +-- Alasmidonta_marginata **
+--+
+-- Lasmigona
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 75 | 4 | 4 | -- |
+-- Alasmidonta_marginata **
+-----+
| +-- Lasmigona
--+
| +----- Pyganodon
+--+
| +-- Strophitus
+--+
+-- Simpsonaias
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 29 | 4 | 3 | -- |
+-- Alasmidonta_marginata **
+--+
| +-- Lasmigona
--+
| +-- Strophitus
+--+
+-- Pyganodon
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 29 | 4 | 3 | -- |
+-- Alasmidonta_marginata **
+--+
| +-- Lasmigona
--+
| +-- Strophitus
+--+
+-- Pyganodon
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 46 | 3 | 3 | -- |
+-------- Lasmigona --+ | +----- Alasmidonta_marginata ** +--+ | +-- Pyganodon +--+ +-- Strophitus
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 46 | 3 | 3 | -- |
+-------- Lasmigona --+ | +----- Alasmidonta_marginata ** +--+ | +-- Strophitus +--+ +-- Pyganodon
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 45 | 3 | 3 | -- |
+-------- Lasmigona --+ | +----- Alasmidonta_marginata ** +--+ | +-- Strophitus +--+ +-- Pyganodon
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 45 | 3 | 3 | -- |
+-------- Lasmigona --+ | +----- Alasmidonta_marginata ** +--+ | +-- Strophitus +--+ +-- Pyganodon
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 24 | 3 | 3 | -- |
+----- Lasmigona | --+ +-- Strophitus | | +--+-- Alasmidonta_marginata ** | +-- Pyganodon
Wu et al., 2018 [Publ 3836], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 4 | 3 | -- |
+----- Strophitus | --+ +-- Platynaias | | +--+-- Alasmidonta_marginata ** | +-- Pyganodon
Aunins et al., 2018 [Publ 3863], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 39 | 4 | 3 | -- |
+-- Prolasmidonta
+--+
| +-- Alasmidonta_varicosa **
--+
| +-- Pyganodon
+--+
+-- Utterbackia
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 53 | 3 | 3 | -- |
+-- Pyganodon
+--+
| +-- Utterbackia
--+
| +-- Alasmidonta_varicosa **
+--+
+-- Prolasmidonta
Graf, 2002 [Publ 2651], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 17 | 1 | 1 | -- |
+-- Alasmidonta_marginata ** --+ +-- Pyganodon
Araujo et al., 2018 [Publ 3779], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 17 | 1 | 1 | -- |
+-- Pyganodon --+ +-- Alasmidonta_marginata **
Chen et al., 2023 [Publ 4302], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- |
+-- Alasmidonta_marginata ** --+ +-- Lasmigona
Chen et al., 2023 [Publ 4302], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- |
+-- Alasmidonta_marginata ** --+ +-- Lasmigona
Wu et al., 2024 [Publ 4388], fig. 6
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 52 | 1 | 1 | -- |
+-- Alasmidonta_varicosa ** --+ +-- Utterbackia
Dai et al., 2025 [Publ 4440], fig. 1
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 43 | 1 | 1 | -- |
+-- Alasmidonta_marginata ** --+ +-- Lasmigona
Dai et al., 2025 [Publ 4440], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 43 | 1 | 1 | -- |
+-- Alasmidonta_marginata ** --+ +-- Lasmigona
Chen et al., 2025 [Publ 4487], fig. 2
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- |
+-- Alasmidonta_marginata ** --+ +-- Lasmigona
Chen et al., 2025 [Publ 4487], fig. 3
| iT | tT | bi | riS | oT | roT | roS | iD | |
| 1.000 | 1 | 1 | 0 | 1 | 40 | 1 | 1 | -- |
+-- Alasmidonta_marginata ** --+ +-- Lasmigona
