MUSSELpdb
Taxonomy
Species & Genus Richness of
freshwater mussels
by Taxonomy
Species & Genus Richness of
freshwater bivalves
by Family
Geography
Species Richness & Endemism of
freshwater mussels
by Geography
Data last updated
Wed 04 February 2026
Bivalvia | ... | Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina | Pustulosa
species Pustulosa kleiniana (Lea, 1852): synonymy
1 specimen + 7 Genbank vouchers
Synonymy
Unio kleinianus Lea, 1852
Lea, 1852. Trans. Amer. Phil. Soc. 10 [O. 5]: 265 [21], pl. 17, fig. 18.
Lea, 1852. Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_84041, Suwanee R., Fla.
Quadrula (sec. Theliderma) kleiniana — Simpson (1900), Simpson (1914)
Quadrula kleiniana — Ortmann (1910), Bogan (2010), Williams et al. (2011), Williams et al. (2014)
Quadrula (Quincuncina) securiformis-kleiniana — Frierson (1927)
Quincuncina infucata — Clench & Turner (1956), Burch (1975), Brim Box & Williams (2000)
Quincuncina securiformis kleiniana — Haas (1969)
Quincuncina kleiniana — Campbell et al. (2005)
Rotundaria kleiniana — Campbell & Lydeard (2012), Campbell & Lydeard (2012)
Amphinaias kleiniana — ITIS (2013)
Cyclonaias kleiniana — Williams et al. (2017), Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023)
Pustulosa kleiniana — Neemuchwala et al. (2023), FMCS (2025)
Unio sparsus Küster, 1861
‘Lea Mss.’ Küster, 1861. Conchyl. Cab. 9(2): 252, pl. 85, fig. 2.
(homonym of Unio sparsus Lea, 1841)
Quadrula (sec. Theliderma) kleiniana — Simpson (1900), Simpson (1914)
Quadrula (Quincuncina) securiformis — Frierson (1927)
Cyclonaias kleiniana — Lopes-Lima et al. (2019)
Unio liratus Shuttleworth, 1852
Shuttleworth, 1852. unknown publication: ??.
Quadrula (Quincuncina) securiformis — Frierson (1927)
Cyclonaias kleiniana — Lopes-Lima et al. (2019)
Taxonomic History (since 1900)
Quadrula (sec. Theliderma) kleiniana — Simpson (1900), Simpson (1914)
Quadrula kleiniana — Ortmann (1910), Bogan (2010), Williams et al. (2011), Williams et al. (2014)
Quadrula (Quincuncina) securiformis — Frierson (1927)
Quadrula (Quincuncina) securiformis-kleiniana — Frierson (1927)
Quincuncina infucata — Clench & Turner (1956), Burch (1975), Brim Box & Williams (2000)
Quincuncina securiformis kleiniana — Haas (1969)
Quincuncina kleiniana — Campbell et al. (2005)
Rotundaria kleiniana — Campbell & Lydeard (2012), Campbell & Lydeard (2012)
Amphinaias kleiniana — ITIS (2013)
Cyclonaias kleiniana — Williams et al. (2017), Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023)
Pustulosa kleiniana — Neemuchwala et al. (2023), FMCS (2025)
