species Cyclonaias tuberculata (Rafinesque, 1820): synonymy

Obliquaria (Rotundaria) tuberculata Rafinesque, 1820

(correct name for Potamilus tubercularis Rafinesque, 1818)
Rafinesque, 1818. Amer. Mon. Mag. 3: 355 [as ‘Potamilus tubercularis’]. [nomen nudum]
Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 308 [also as ‘U. tuberculata’].
Rafinesque, 1864. [in] Binney & Tryon, reprint: 23, 52.
Type(s): LECTOTYPE ANSP_20215, Ohio River.

Quadrula (Rotundaria) tuberculata — Simpson (1900), Ortmann (1909), Ortmann (1910), Simpson (1914)
Quadrula tuberculata — Ortmann (1909), Ortmann (1909), Vanatta (1915)
Rotundaria tuberculata — Ortmann (1910), Ortmann (1912), Utterback (1915), Ortmann (1918), Thiele (1934), Campbell & Lydeard (2012), Campbell & Lydeard (2012)
Cyclonaias tuberculata — Ortmann & Walker (1922), Ortmann (1924), Ortmann (1925), F.C. Baker (1928), Haas (1969), ... Hopper et al. (2023), Hauck et al. (2023), Keogh et al. (2024), FMCS (2025), Campbell & Lydeard (2026)
Quadrula (Cyclonaias) tuberculata — Frierson (1927), Haas (1930)
Cyclonaias tuberculata tuberculata — Haas (1969)
“Cyclonaias tuberculata” ** — Serb et al. (2003)
Cyclonaias tuberculate [sic] — Gasparini et al. (2020)

Unio verrucosus Barnes, 1823

Barnes, 1823. Amer. J. Sci. 6 (1-2): 123, pl. 5, figs. 6a-b.

Quadrula (Rotundaria) tuberculata — Simpson (1900), Simpson (1914)
Unio (s.s.) verrucosus — Call (1900)
Unio verrucosus — de Gregorio (1914)
Rotundaria tuberculata — Utterback (1915)
Quadrula (Cyclonaias) tuberculata — Frierson (1927)
Cyclonaias tuberculata — F.C. Baker (1928), Oesch (1984), Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), Lopes-Lima et al. (2019)
Cyclonaias tuberculata tuberculata — Haas (1969)

Unio tuberculosus Valenciennes, 1827

(error for Obliquaria (Rotundaria) tuberculata Rafinesque, 1820)
Valenciennes, 1827. Receuil Observ. Zool. 2(11-12): 232.

Quadrula (Rotundaria) tuberculata — Simpson (1900), Simpson (1914)
Quadrula (Cyclonaias) tuberculata — Frierson (1927)
Cyclonaias tuberculata tuberculata — Haas (1969)
Cyclonaias tuberculata — Parmalee & Bogan (1998), Watters et al. (2009)

Unio verrucosus purpureus Hildreth, 1828

(homonym of Unio purpureus Say, 1817; intro'd as intra-sp. of Unio verrucosus Barnes, 1823)
Hildreth, 1828. Amer. J. Sci. 14: 281.

Quadrula (Rotundaria) tuberculata — Simpson (1900), Simpson (1914)
Cyclonaias tuberculata — Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), Lopes-Lima et al. (2019)

Unio graniferus Lea, 1838

Lea, 1836. Synopsis Fam. Naiades: addenda. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 69, pl. 19, fig. 60.
Type(s): HOLOTYPE USNM_84296, Tuscumbia, Ala.

Quadrula (Rotundaria) granifera — Simpson (1900), Simpson (1914)
Unio (s.s.) graniferus — Call (1900)
Rotundaria granifera — Utterback (1915)
Cyclonaias tuberculata granifera — Ortmann (1925), F.C. Baker (1928), Haas (1969)
Quadrula (Cyclonaias) tuberculata-granifera — Frierson (1927)
Cyclonaias tuberculata — R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), Lopes-Lima et al. (2019)

Quadrula (sec. Rotundaria) granifera var. pusilla Simpson, 1900

(intro'd as intra-sp. of Unio graniferus Lea, 1838)
Simpson, 1900. Proc. U.S. Nat. Mus. 22: 795.
Simpson, 1914. Descr. Cat. Naiades: 906.
Type(s): LECTOTYPE MCZ_20176 (by R.I. Johnson, 1956: 131), Green River, Kentucky.

Quadrula (Rotundaria) granifera var. pusilla — Simpson (1900), Simpson (1914)
Quadrula (Cyclonaias) tuberculata-pusilla — Frierson (1927)
Cyclonaias tuberculata pusilla — Haas (1969)
Cyclonaias tuberculata — R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), Lopes-Lima et al. (2019)

Quadrula (Cyclonaias) tuberculata utterbackiana Frierson, 1927

(intro'd as intra-sp. of Obliquaria (Rotundaria) tuberculata Rafinesque, 1820)
‘Lea’ Utterback, 1915. Amer. Midl. Nat. 4(5): 183, pl. 20, fig. 57a-b [as ‘Plethobasus cooperianus’].
Frierson, 1927. Check List N. Amer. Naiades: 52, sp. 224c.

Quadrula (Cyclonaias) tuberculata-utterbackiana — Frierson (1927)
Cyclonaias tuberculata utterbackiana — Haas (1969)
Cyclonaias tuberculata — R.I. Johnson (1980), Williams et al. (2008), Watters et al. (2009), Lopes-Lima et al. (2019)

Cyclonaias tuberculata compressa F.C. Baker, 1928

(intro'd as intra-sp. of Obliquaria (Rotundaria) tuberculata Rafinesque, 1820)
F.C. Baker, 1928. Bull. Wisconsin Geol. Nat. Hist. Survey 70(2): 107, pl. 75, figs. 1-2.
Type(s): HOLOTYPE (ORIG. DESIG.) INHS_82236, Big Vermilion River, 0.5 mi below junction with Middle Fork, [Vermilion Co.], Illinois.

Cyclonaias tuberculata compressa — F.C. Baker (1928)

Potamilus tubercularis Rafinesque, 1818

(incorrect original spelling of Obliquaria (Rotundaria) tuberculata Rafinesque, 1820)
Rafinesque, 1818. Amer. Mon. Mag. 3: 355. [nomen nudum]
Rafinesque, 1864. [in] Binney & Tryon, reprint: 23.

Plethobasus cooperianus Utterback, 1915

(misidentified reference to Unio cooperianus Lea, 1834)
‘Lea’ Utterback, 1915. Amer. Midl. Nat. 4(5): 183, pl. 20, fig. 57a-b.

Quadrula (Cyclonaias) tuberculata-utterbackiana — Frierson (1927)
Cyclonaias tuberculata utterbackiana — Haas (1969)
Cyclonaias tuberculata — Parmalee & Bogan (1998), Watters et al. (2009)

Cylonaias tuberculate Gasparini et al., 2020

(error for Obliquaria (Rotundaria) tuberculata Rafinesque, 1820)
Gasparini et al., 2020. Environmental DNA 2(3): Table S1.