species Cyclonaias tuberculata (Rafinesque, 1820): synonymy

Obliquaria (Rotundaria) tuberculata Rafinesque, 1820

Rafinesque, 1818. Amer. Mon. Mag. 3: 355 [as ‘Potamilus tubercularis’]. [nomen nudum]
Rafinesque, 1820. Ann. Gén. Sci. Phys. 5 (13): 308 [also as ‘U. tuberculata’].
Rafinesque, 1864. [in] Binney & Tryon, reprint: 23, 52.
(correct name for Potamilus tubercularis Rafinesque, 1818)
Type(s): LECTOTYPE ANSP_20215, Ohio River.

Quadrula (Rotundaria) tuberculata — Simpson (1900), Ortmann (1909), Ortmann (1910), Simpson (1914)
Quadrula tuberculata — Ortmann (1909), Ortmann (1909), Vanatta (1915)
Rotundaria tuberculata — Ortmann (1910), Ortmann (1912), Utterback (1915), Ortmann (1918), Thiele (1934), Campbell & Lydeard (2012), Campbell & Lydeard (2012)
Cyclonaias tuberculata — Ortmann & Walker (1922), Ortmann (1924), Ortmann (1925), F.C. Baker (1928), Haas (1969), ... Neemuchwala et al. (2023), Hopper et al. (2023), Hauck et al. (2023), Keogh et al. (2024), FMCS (2025)
Quadrula (Cyclonaias) tuberculata — Frierson (1927), Haas (1930)
Cyclonaias tuberculata tuberculata — Haas (1969)
Cyclonaias tuberculate [sic] — Gasparini et al. (2020)

Unio verrucosus Barnes, 1823

Barnes, 1823. Amer. J. Sci. 6 (1-2): 123, pl. 5, figs. 6a-b.

Quadrula (Rotundaria) tuberculata — Simpson (1900), Simpson (1914)
Unio (s.s.) verrucosus — Call (1900)
Unio verrucosus — de Gregorio (1914)
Rotundaria tuberculata — Utterback (1915)
Quadrula (Cyclonaias) tuberculata — Frierson (1927)
Cyclonaias tuberculata — F.C. Baker (1928), Oesch (1984), Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), Lopes-Lima et al. (2019)
Cyclonaias tuberculata tuberculata — Haas (1969)

Unio tuberculosus Valenciennes, 1827

Valenciennes, 1827. Receuil Observ. Zool. 2(11-12): 232.
(error for Obliquaria (Rotundaria) tuberculata Rafinesque, 1820)

Quadrula (Rotundaria) tuberculata — Simpson (1900), Simpson (1914)
Quadrula (Cyclonaias) tuberculata — Frierson (1927)
Cyclonaias tuberculata tuberculata — Haas (1969)
Cyclonaias tuberculata — Parmalee & Bogan (1998), Watters et al. (2009)

Unio verrucosus purpureus Hildreth, 1828

Hildreth, 1828. Amer. J. Sci. 14: 281.
(homonym of Unio purpureus Say, 1817; intro'd as intra-sp. of Unio verrucosus Barnes, 1823)

Quadrula (Rotundaria) tuberculata — Simpson (1900), Simpson (1914)
Cyclonaias tuberculata — Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), Lopes-Lima et al. (2019)

Unio graniferus Lea, 1838

Lea, 1836. Synopsis Fam. Naiades: addenda. [nomen nudum]
Lea, 1838. Trans. Amer. Phil. Soc. 6 [O. 2]: 69, pl. 19, fig. 60.
Type(s): HOLOTYPE USNM_84296, Tuscumbia, Ala.

Quadrula (Rotundaria) granifera — Simpson (1900), Simpson (1914)
Unio (s.s.) graniferus — Call (1900)
Rotundaria granifera — Utterback (1915)
Cyclonaias tuberculata granifera — Ortmann (1925), F.C. Baker (1928), Haas (1969)
Quadrula (Cyclonaias) tuberculata-granifera — Frierson (1927)
Cyclonaias tuberculata — R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), Lopes-Lima et al. (2019)

Quadrula (sec. Rotundaria) granifera var. pusilla Simpson, 1900

Simpson, 1900. Proc. U.S. Nat. Mus. 22: 795.
Simpson, 1914. Descr. Cat. Naiades: 906.
(intro'd as intra-sp. of Unio graniferus Lea, 1838)
Type(s): LECTOTYPE MCZ_20176 (by R.I. Johnson, 1956: 131), Green River, Kentucky.

Quadrula (Rotundaria) granifera var. pusilla — Simpson (1900), Simpson (1914)
Quadrula (Cyclonaias) tuberculata-pusilla — Frierson (1927)
Cyclonaias tuberculata pusilla — Haas (1969)
Cyclonaias tuberculata — R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), Lopes-Lima et al. (2019)

Quadrula (Cyclonaias) tuberculata utterbackiana Frierson, 1927

‘Lea’ Utterback, 1915. Amer. Midl. Nat. 4(5): 183, pl. 20, fig. 57a-b [as ‘Plethobasus cooperianus’].
Frierson, 1927. Check List N. Amer. Naiades: 52, sp. 224c.
(intro'd as intra-sp. of Obliquaria (Rotundaria) tuberculata Rafinesque, 1820)

Quadrula (Cyclonaias) tuberculata-utterbackiana — Frierson (1927)
Cyclonaias tuberculata utterbackiana — Haas (1969)
Cyclonaias tuberculata — R.I. Johnson (1980), Williams et al. (2008), Watters et al. (2009), Lopes-Lima et al. (2019)

Cyclonaias tuberculata compressa F.C. Baker, 1928

F.C. Baker, 1928. Bull. Wisconsin Geol. Nat. Hist. Survey 70(2): 107, pl. 75, figs. 1-2.
(intro'd as intra-sp. of Obliquaria (Rotundaria) tuberculata Rafinesque, 1820)
Type(s): HOLOTYPE (ORIG. DESIG.) INHS_82236, Big Vermilion River, 0.5 mi below junction with Middle Fork, [Vermilion Co.], Illinois.

Cyclonaias tuberculata compressa — F.C. Baker (1928)

Potamilus tubercularis Rafinesque, 1818

Rafinesque, 1818. Amer. Mon. Mag. 3: 355. [nomen nudum]
Rafinesque, 1864. [in] Binney & Tryon, reprint: 23.
(incorrect original spelling of Obliquaria (Rotundaria) tuberculata Rafinesque, 1820)

Plethobasus cooperianus Utterback, 1915

‘Lea’ Utterback, 1915. Amer. Midl. Nat. 4(5): 183, pl. 20, fig. 57a-b.
(misidentified reference to Unio cooperianus Lea, 1834)

Quadrula (Cyclonaias) tuberculata-utterbackiana — Frierson (1927)
Cyclonaias tuberculata utterbackiana — Haas (1969)
Cyclonaias tuberculata — Parmalee & Bogan (1998), Watters et al. (2009)

Cylonaias tuberculate Gasparini et al., 2020

Gasparini et al., 2020. Environmental DNA 2(3): Table S1.
(error for Obliquaria (Rotundaria) tuberculata Rafinesque, 1820)