Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Anemina Haas, 1969: cladomics

1 Recent valid species (1 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 1 (1 total)
analyzed in 99 cladograms in 69 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)

Cladograms

Chen et al., 2026 [Publ 4805], fig. 4

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 32 8 **
Cristaria Beringiana Mimo Acudonta Sinanodonta Pletholophus Anemina harlandi Amuranodonta Buldowskia

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cstiTtTbiriSoTroTroSiD
0.900 1 1 0 1 632 32 8 0.0
Amuranodonta Anemina harlandi Buldowskia

Xiang et al., 2025 [Publ 4433], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 27 26 7 0.0
Pletholophus Beringiana Mimo Sinanodonta Cristaria Anemina harlandi Amuranodonta Buldowskia

Wu et al., 2025 [Publ 4442], fig. 2

cstiTtTbiriSoTroTroSiD
0.889 1 1 0 1 34 32 7 0.0
Cristaria Beringiana Acudonta Sinanodonta Pletholophus guangzhouensis Pletholophus tenuis Pletholophus reinianus Pletholophus honglinhensis Anemina harlandi Amuranodonta Buldowskia

Wu et al., 2025 [Publ 4442], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 16 14 7 0.0
Cristaria Amuranodonta Anemina harlandi Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Wang et al., 2025 [Publ 4488], fig. 3B

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 63 14 7 0.0
Cristaria Amuranodonta Anemina harlandi Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Wu et al., 2026 [Publ 4811], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 14 7 0.0
Cristaria Amuranodonta Anemina harlandi Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Wu et al., 2026 [Publ 4825], fig. 6B

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 64 13 7 0.0
Cristaria Amuranodonta Anemina harlandi Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 121 23 6 0.0
Cristaria Anemina harlandi Amuranodonta Buldowskia Pletholophus Beringiana Sinanodonta

Bogan et al., 2023 [Publ 4295], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 26 6 0.0
Cristaria Amuranodonta Anemina harlandi Buldowskia Pletholophus Beringiana Sinanodonta

Dai et al., 2024 [Publ 4385], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 26 6 0.0
Amuranodonta Anemina harlandi Buldowskia Cristaria Pletholophus Beringiana Sinanodonta

Dai et al., 2024 [Publ 4385], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 26 6 0.0
Amuranodonta Anemina harlandi Buldowskia Cristaria Pletholophus Beringiana Sinanodonta

Dai et al., 2026 [Publ 4757], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 29 6 0.0
Amuranodonta Anemina harlandi Buldowskia Cristaria Pletholophus Beringiana Sinanodonta

Bolotov et al., 2020 [Publ 4084], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 8 5 0.0
Amuranodonta Anemina harlandi Buldowskia Cristaria Beringiana Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 1

cstiTtTbiriSoTroTroSiD
0.286 1 1 0 1 33 9 5 0.0
Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina harlandi Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula Lepidodesma Scriptolamprotula Acuticosta Cuneopsis Nodularia Schistodesmus Inversiunio

Sano et al., 2020 [Publ 4138], fig. 2

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 68 10 5 0.0
Anemina harlandi Buldowskia

Sano et al., 2020 [Publ 4138], fig. 3

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 68 10 5 0.0
Anemina harlandi Buldowskia

Sano et al., 2020 [Publ 4138], fig. 4

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 68 10 5 0.0
Anemina harlandi Buldowskia

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 55 12 5 0.0
Anemina harlandi Amuranodonta Buldowskia Cristaria Beringiana Sinanodonta

Chen et al., 2023 [Publ 4302], fig. 1

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 40 5 5 0.0
Anemina harlandi Amuranodonta

Chen et al., 2023 [Publ 4302], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 5 5 0.0
Cristaria Anemina harlandi Amuranodonta Pletholophus Sinanodonta Beringiana

Dai et al., 2025 [Publ 4440], fig. 1

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 43 5 5 0.0
Anemina harlandi Amuranodonta

Dai et al., 2025 [Publ 4440], fig. 2

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 43 5 5 0.0
Anemina harlandi Amuranodonta

Chen et al., 2025 [Publ 4487], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 5 5 0.0
Cristaria Anemina harlandi Amuranodonta Pletholophus Sinanodonta Beringiana

Chen et al., 2025 [Publ 4487], fig. 3

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 40 5 5 0.0
Anemina harlandi Amuranodonta Pletholophus Sinanodonta Beringiana

Xiang et al., 2025 [Publ 4433], fig. 7

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 12 6 4 0.0
Cristaria Anemina harlandi Beringiana Mimo Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 10

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 11 5 4 0.0
Anemina harlandi Amuranodonta Cristaria Beringiana Sinanodonta

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 80 4 3 0.0
Cristaria Anemina harlandi Pletholophus Sinanodonta

Froufe et al., 2017 [Publ 3802], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 11 9 3 0.0
Anemina harlandi Cristaria Beringiana Sinanodonta

Huang et al., 2019 [Publ 3843], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 200 8 3 0.0
Anemina harlandi Cristaria Pletholophus Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 198 8 3 0.0
Anemina harlandi Cristaria Pletholophus Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 200 8 3 0.0
Acuticosta Alasmidonta Anodonta anatina Anemina harlandi Anodonta cygnea Cristaria Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus Unio Scriptolamprotula Lanceolaria Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 10 8 3 0.0
Sinanodonta Amuranodonta Anemina harlandi Buldowskia

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 5 3 0.0
Anemina harlandi Cristaria Beringiana Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 5 3 0.0
Cristaria Anemina harlandi Beringiana Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 5 3 0.0
Cristaria Anemina harlandi Beringiana Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 5 3 0.0
Cristaria Anemina harlandi Beringiana Sinanodonta

Wu et al., 2024 [Publ 4379], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 5 3 0.0
Cristaria Anemina harlandi Beringiana Sinanodonta

Wu et al., 2024 [Publ 4388], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 3 3 0.0
Cristaria Anemina harlandi Beringiana Sinanodonta

Hou et al., 2024 [Publ 4400], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 74 5 3 0.0
Cristaria Anemina harlandi Beringiana Sinanodonta

Hou et al., 2025 [Publ 4445], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 51 5 3 0.0
Cristaria Anemina harlandi Beringiana Sinanodonta

Xue et al., 2016 [Publ 3659], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 14 3 2 0.0
Anemina harlandi Sinanodonta

Wu et al., 2016 [Publ 3711], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 20 2 2 0.0
Anemina harlandi Cristaria Anodonta Pyganodon Utterbackia

Fonseca et al., 2016 [Publ 3775], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 31 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Huang et al., 2018 [Publ 3828], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 41 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wu et al., 2018 [Publ 3836], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 5 2 0.0
Anemina harlandi Cristaria Sinanodonta

Wang et al., 2018 [Publ 3846], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Aunins et al., 2018 [Publ 3863], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 39 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wang et al., 2018 [Publ 3865], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Cha et al., 2018 [Publ 3916], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 37 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wu et al., 2018 [Publ 4143], fig. 2

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 21 2 2 0.0
Anemina harlandi Cristaria Anodonta Pyganodon Utterbackia

Huang et al., 2019 [Publ 3843], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 45 7 2 0.0
Anemina harlandi Acuticosta Sinanodonta Scriptolamprotula Cristaria Lanceolaria Lepidodesma Schistodesmus Nodularia Cuneopsis rufescens Tchangsinaia Cuneopsis celtiformis Cuneopsis szechenyii Cuneopsis heudei

Huang et al., 2019 [Publ 3843], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 55 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 46 7 2 0.0
Anemina harlandi Sinanodonta Acuticosta Lepidodesma Cristaria Lanceolaria Scriptolamprotula Nodularia Tchangsinaia Cuneopsis Schistodesmus

Wu et al., 2019 [Publ 3864], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 45 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Froufe et al., 2019 [Publ 3936], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 43 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Froufe et al., 2019 [Publ 3936], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 43 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 2 2 0.0
Cristaria Anemina harlandi Sinanodonta

Riccardi et al., 2020 [Publ 4089], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 18 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 53 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wu et al., 2020 [Publ 4121], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wu et al., 2020 [Publ 4147], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 29 2 2 0.0
Cristaria Sinanodonta Anemina harlandi

Chen et al., 2020 [Publ 4170], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 33 2 2 0.0
Alasmidonta Anemina harlandi

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 2_I

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 3 2 0.0
Anemina harlandi Cristaria Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 67 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 3 2 0.0
Anemina harlandi Cristaria Sinanodonta

Wu et al., 2022 [Publ 4238], fig. 2d

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 37 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wu et al., 2022 [Publ 4238], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 36 2 2 0.0
Cristaria Sinanodonta Anemina harlandi

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 38 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Dai et al., 2023 [Publ 4285], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 2 2 0.0
Cristaria Anemina harlandi Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 18 3 2 0.0
Anemina harlandi Cristaria Sinanodonta

Dai et al., 2024 [Publ 4358], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 2 2 0.0
Cristaria Anemina harlandi Sinanodonta

Dai et al., 2024 [Publ 4389], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 45 2 2 0.0
Sinanodonta Anemina harlandi Cristaria

Dai et al., 2025 [Publ 4440], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 86 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Dai et al., 2025 [Publ 4441], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 46 2 2 0.0
Sinanodonta Anemina harlandi Cristaria

Chen et al., 2025 [Publ 4448], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 84 4 2 0.0
Cristaria Anemina harlandi Sinanodonta

Chen et al., 2025 [Publ 4663], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 2 2 0.0
Sinanodonta Anemina harlandi Cristaria

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 139 3 2 0.0
Cristaria Anemina harlandi Sinanodonta

Ouyang et al., 2011 [Publ 3408], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 12 2 1 0.0
Anemina harlandi Sinanodonta

Ouyang et al., 2015 [Publ 4493], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 3 1 0.0
Anemina harlandi Sinanodonta

Ouyang et al., 2015 [Publ 4493], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 3 1 0.0
Anemina harlandi Sinanodonta

Feng et al., 2016 [Publ 3800], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 13 2 1 0.0
Anemina harlandi Sinanodonta

Vikhrev et al., 2017 [Publ 3810], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 9 7 1 0.0
Anemina harlandi Sinanodonta

Bespalaya et al., 2018 [Publ 3811], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 9 7 1 0.0
Anemina harlandi Sinanodonta

Kondakov et al., 2018 [Publ 3914], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 7 7 1 0.0
Anemina harlandi Sinanodonta

Riccardi et al., 2020 [Publ 4089], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 1 1 0.0
Sinanodonta Anemina harlandi

Bauer et al., 2021 [Publ 4365], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 7 7 1 0.0
Anemina harlandi Sinanodonta

Whelan et al., 2023 [Publ 4287], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 3 1 0.0
Anemina harlandi Sinanodonta

Kashiri et al., 2024 [Publ 4822], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 9 7 1 0.0
Anemina harlandi Sinanodonta

Zhou et al., 2017 [Publ 3788], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 15 0 0 0.0
Anemina harlandi Lanceolaria Scriptolamprotula Nodularia Tchangsinaia Quadrula Potamilus Lampsilis Sinohyriopsis Lamprotula Ptychorhynchus Sinosolenaia

Liu et al., 2017 [Publ 3939], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 14 0 0 0.0
Anemina harlandi Sinosolenaia

Dai et al., 2024 [Publ 4386], fig. 5-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 0 0 0.0
Acuticosta Anemina harlandi Anodonta Lanceolaria Scriptolamprotula Lepidodesma Unio Middendorffinaia Cuneopsis Nodularia Inversiunio Schistodesmus

Dai et al., 2024 [Publ 4386], fig. 5-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 21 0 0 0.0
Acuticosta Anemina harlandi