The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina genus Pilsbryoconcha Simpson, 1900: cladomics
9 Recent valid species (9 total species) Cladomics pages are experimental — even more so than the rest of this site!
The purpose of the cladomics pages for each genus and family-group level taxon is to
provide statistics and references indicating the extent to which published phylogenetic
work supports the classification used here. The Cladomics Summary provides an
overview of the number of studies that have included terminal species representing this
taxon as well as an indication of how frequently it has been recovered as monophyletic.
Following the summary, the available published clades representing the taxon are listed.
For more information, visit the Cladomics FARQ (frequently
asked rhetorical questions) page. Abbreviations for the various statistics are explained on
the FARQ! Cladomics Summarynumber of species = 9 (9 total)
analyzed in 79 cladograms in 50 publications
maximum number of terminals in an analysis =
9 (mean = 1.99, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 27
maximum number of included terminals = 9, (mean =
3.89 median = 2)
proportion of cladograms that support basic monophyly
= 0.85
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup
distances within the cladome. See the Cladmoics
FARQ for more information.
Bolotov et al., 2023 [Publ 4312], fig. S2 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 8 | 10 | 32 | 27 | 7 | 0 | +-------------------------------- Songkhlanaia
|
| +-- Nyeinchanconcha
--+ +--------------------------+
| | +-- Sundadontina
| |
+--+ +----- Namkongnaia
| +--------------------+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +----------------------- Pseudodon
| |
+--+ +-------------------- Pilsbryoconcha_carinifera **
| |
+--+ +-- Pilsbryoconcha_linguaeformis__2 **
| +--------------+
| | +-- Pilsbryoconcha **
+--+
| +-------------- Pilsbryoconcha_linguaeformis__1 **
| |
+--+ +-- Pilsbryoconcha_schomburgki **
| +--------+
| | +-- Pilsbryoconcha_exilis **
+--+
| +-------- Pilsbryoconcha_hoikaab **
+--+
| +----- Pilsbryoconcha_mekongiana **
+--+
| +-- Pilsbryoconcha_acuta **
+--+
+-- Pilsbryoconcha_kittitati **
Konopleva et al., 2025 [Publ 4446], fig. 4 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 8 | 37 | 32 | 9 | 12 | +-------------------- Lannanaia
|
| +-------------- Songkhlanaia
| |
| | +----- Nyeinchanconcha
| +--+ +-----+
--+ | | | | +-- Thaiconcha
| | | | +--+
| | +--+ +-- Bineurus
| | |
| | | +-------- Sundadontina
| | +--+
| | | +----- Pseudodon
+--+ +--+
| | +-- Namkongnaia
| +--+
| +-- Isannaia
|
| +-- Pilsbryoconcha_exilis__1 **
| +-----------+
| | +-- Pilsbryoconcha_schomburgki **
| |
+--+ +-- Pilsbryoconcha_carinifera **
| +--------+
| | +-- Pilsbryoconcha_exilis__2 **
+--+
| +-------- Pilsbryoconcha_kittitati **
+--+
| +----- Pilsbryoconcha_acuta **
+--+
| +-- Pilsbryoconcha_hoikaab **
+--+
+-- Pilsbryoconcha_mekongiana **
Bolotov et al., 2023 [Publ 4312], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 8 | 31 | 26 | 7 | 10 | +-------------- Songkhlanaia
|
+--+ +-- Namkongnaia
| | +--------+
| | | +-- Pseudodon
| +--+
| | +-------- Sundadontina
| +--+
| | +----- Nyeinchanconcha
| +--+
--+ | +-- Thaiconcha
| +--+
| +-- Bineurus
|
| +-- Pilsbryoconcha_exilis **
| +-----------+
| | +-- Pilsbryoconcha_schomburgki **
| |
+--+ +-- Pilsbryoconcha_carinifera **
| +--------+
| | +-- Pilsbryoconcha_linguaeformis **
+--+
| +-------- Pilsbryoconcha_acuta **
+--+
| +----- Pilsbryoconcha_kittitati **
+--+
| +-- Pilsbryoconcha_mekongiana **
+--+
+-- Pilsbryoconcha_hoikaab **
Bolotov et al., 2023 [Publ 4312], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 8 | 31 | 26 | 7 | 10 | +----- Nyeinchanconcha
+-----+
| | +-- Thaiconcha
| +--+
+-----+ +-- Bineurus
| |
| | +-------- Songkhlanaia
| +--+
| | +----- Sundadontina
| +--+
| | +-- Namkongnaia
--+ +--+
| +-- Pseudodon
|
| +-- Pilsbryoconcha_exilis **
| +-----------+
| | +-- Pilsbryoconcha_schomburgki **
| |
+--+ +-- Pilsbryoconcha_carinifera **
| +--------+
| | +-- Pilsbryoconcha_linguaeformis **
+--+
| +-------- Pilsbryoconcha_kittitati **
+--+
| +----- Pilsbryoconcha_acuta **
+--+
| +-- Pilsbryoconcha_hoikaab **
+--+
+-- Pilsbryoconcha_mekongiana **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 8 | 68 | 7 | 6 | 6 | +-- Nyeinchanconcha
+--+
| +-- Sundadontina
+--------------+
| | +-- Thaiconcha
| +--+
| +-- Bineurus
--+
| +-- Namkongnaia
| +--------------+
| | +-- Pseudodon
| |
+--+ +-- Pilsbryoconcha_exilis **
| +-----------+
| | +-- Pilsbryoconcha_schomburgki **
| |
+--+ +-- Pilsbryoconcha_carinifera **
| +--------+
| | +-- Pilsbryoconcha_linguaeformis **
+--+
| +-------- Pilsbryoconcha_hoikaab **
+--+
| +----- Pilsbryoconcha_mekongiana **
+--+
| +-- Pilsbryoconcha_acuta **
+--+
+-- Pilsbryoconcha_kittitati **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 8 | 68 | 7 | 6 | 6 | +-- Nyeinchanconcha
+--+
| +-- Sundadontina
+--------------+
| | +-- Thaiconcha
| +--+
| +-- Bineurus
--+
| +-- Namkongnaia
| +--------------+
| | +-- Pseudodon
| |
+--+ +-- Pilsbryoconcha_exilis **
| +-----------+
| | +-- Pilsbryoconcha_schomburgki **
| |
+--+ +-- Pilsbryoconcha_carinifera **
| +--------+
| | +-- Pilsbryoconcha_linguaeformis **
+--+
| +-------- Pilsbryoconcha_hoikaab **
+--+
| +----- Pilsbryoconcha_mekongiana **
+--+
| +-- Pilsbryoconcha_acuta **
+--+
+-- Pilsbryoconcha_kittitati **
Jeratthitikul et al., 2024 [Publ 4387], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 8 | 51 | 28 | 9 | 10 | +----------------------- Pseudodon
|
| +-------- Sundadontina
| +-----------+
--+ | | +----- Nyeinchanconcha
| | +--+
| | | +-- Thaiconcha
| | +--+
| | +-- Bineurus
+--+
| +----- Namkongnaia
| +-----------+
| | | +-- Lannanaia
| | +--+
| | +-- Songkhlanaia
+--+
| +-------------- Isannaia
| |
| | +-- Pilsbryoconcha_exilis__2 **
+--+ +--------+
| | +-- Pilsbryoconcha_schomburgki **
| |
+--+ +-- Pilsbryoconcha_exilis__1 **
| +-----+
| | +-- Pilsbryoconcha_carinifera **
| |
+--+ +-- Pilsbryoconcha_acuta **
| +--+
| | +-- Pilsbryoconcha_kittitati **
+--+
| +-- Pilsbryoconcha_hoikaab **
+--+
+-- Pilsbryoconcha_mekongiana **
Jeratthitikul et al., 2025 [Publ 4423], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 8 | 43 | 31 | 9 | 8 | +-------------- Songkhlanaia
|
| +----- Nyeinchanconcha
+-----+ +-----+
| | | | +-- Thaiconcha
| | | +--+
| +--+ +-- Bineurus
| |
| | +-------- Sundadontina
| +--+
| | +----- Isannaia
--+ +--+
| | +-- Namkongnaia
| +--+
| +-- Pseudodon
|
| +----------------- Lannanaia
| |
| | +-- Pilsbryoconcha_exilis__2 **
+--+ +-----------+
| | +-- Pilsbryoconcha_schomburgki **
| |
+--+ +-- Pilsbryoconcha_carinifera **
| +--------+
| | +-- Pilsbryoconcha_exilis__1 **
+--+
| +-------- Pilsbryoconcha_hoikaab **
+--+
| +----- Pilsbryoconcha_mekongiana **
+--+
| +-- Pilsbryoconcha_acuta **
+--+
+-- Pilsbryoconcha_kittitati **
Konopleva et al., 2023 [Publ 4262], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 6 | 77 | 21 | 7 | 12 | +-- Pilsbryoconcha_schomburgki **
+--------+
| +-- Pilsbryoconcha_exilis__1 **
+--------+
| | +-------- Pilsbryoconcha **
| +--+
| | +----- Pilsbryoconcha_linguaeformis **
| +--+
--+ | +-- Pilsbryoconcha_exilis__2 **
| +--+
| +-- Pilsbryoconcha_carinifera **
|
| +----------------- Songkhlanaia
+--+
| +-------------- Namkongnaia
+--+
| +----------- Pseudodon
+--+
| +-------- Sundadontina
+--+
| +----- Nyeinchanconcha
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Jeratthitikul et al., 2021 [Publ 4219], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 33 | 15 | 6 | 9 | +----- Pilsbryoconcha_exilis **
+-----+
| | +-- Pilsbryoconcha_linguaeformis **
| +--+
| +-- Pilsbryoconcha **
--+
| +----- Nyeinchanconcha
| +--+
| | | +-- Thaiconcha
| | +--+
+--+ +-- Bineurus
|
| +----- Sundadontina
+--+
| +-- Namkongnaia
+--+
+-- Pseudodon
Jeratthitikul et al., 2021 [Publ 4219], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 33 | 15 | 6 | 9 | +----- Pilsbryoconcha_exilis **
+--------+
| | +-- Pilsbryoconcha_linguaeformis **
| +--+
| +-- Pilsbryoconcha **
|
| +-- Nyeinchanconcha
--+ +--+
| | +-- Sundadontina_cumingii
| +-----+
| | | +-- Thaiconcha
| | +--+
| | +-- Bineurus
+--+
| +-- Namkongnaia
| +-----+
| | +-- Pseudodon
+--+
| +----- Sundadontina_tumida
+--+
| +-- Sundadontina_brandti
+--+
+-- Sundadontina_taskaevi
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 4 | 123 | 10 | 5 | -- | +-------------- Bineurus
|
| +-- Nyeinchanconcha
+-----------+
| +-- Sundadontina_cumingii
--+
| +-- Sundadontina_tumida
+-----------+
| +-- Pseudodon
|
| +----------- Thaiconcha
+--+
| +-------- Pilsbryoconcha_exilis__3 **
+--+
| +----- Pilsbryoconcha_exilis__2 **
+--+
| +-- Pilsbryoconcha_exilis__1 **
+--+
+-- Pilsbryoconcha_carinifera **
Dai et al., 2023 [Publ 4285], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 12 | 5 | -- | +-- Nyeinchanconcha
+--------+
| +-- Bineurus
|
--+ +-- Thaiconcha
| +-----+
| | +-- Sundadontina
+--+
| +----- Pseudodon
+--+
| +-- Pilsbryoconcha_carinifera **
+--+
+-- Pilsbryoconcha_exilis **
Bolotov et al., 2020 [Publ 4100], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 3 | 110 | 15 | 5 | -- | +----------------- Bineurus
|
--+ +-------------- Nyeinchanconcha
| |
+--+ +-- Pseudodon
| +--------+
| | +-- Sundadontina
+--+
| +-------- Thaiconcha
+--+
| +----- Pilsbryoconcha_exilis__2 **
+--+
| +-- Pilsbryoconcha_exilis__1 **
+--+
+-- Pilsbryoconcha **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 3 | 197 | 10 | 5 | -- | +-------- Nyeinchanconcha
|
+-------- Thaiconcha
|
+-------- Sundadontina_cumingii
|
+-------- Sundadontina_tumida__1
--+
+-------- Sundadontina_tumida__2
|
+-------- Pseudodon
|
+-------- Bineurus
|
| +----- Pilsbryoconcha_exilis__2 **
+--+
| +-- Pilsbryoconcha_carinifera **
+--+
+-- Pilsbryoconcha_exilis__1 **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 4 | 137 | 11 | 5 | -- | +-------- Pilsbryoconcha_exilis__3 **
--+
| +----- Pilsbryoconcha_exilis__2 **
+--+
| +-- Pilsbryoconcha_exilis__1 **
+--+
+-- Pilsbryoconcha_carinifera **
Dai et al., 2023 [Publ 4285], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 12 | 5 | -- | +-- Nyeinchanconcha
+-----+
| +-- Bineurus
|
| +-- Thaiconcha
--+-----+
| +-- Sundadontina
|
| +----- Pseudodon
+--+
| +-- Pilsbryoconcha_carinifera **
+--+
+-- Pilsbryoconcha_exilis **
Konopleva et al., 2019 [Publ 3900], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 8 | 0 | 2 | 50 | 6 | 2 | 14 | +-- Bineurus_mouhotii
|
+-- Pseudodon_vagulus
|
+-- Pseudodon_mekongi
|
+-- Pilsbryoconcha_exilis **
--+
+-- Pilsbryoconcha_carinifera **
|
+-- Bineurus_loeiensis
|
+-- Bineurus_exilis
|
+-- Pseudodon_vondembuschianus
Bolotov et al., 2020 [Publ 4100], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 3 | 116 | 15 | 5 | -- | +----------------- Bineurus
|
--+ +-------------- Nyeinchanconcha
| |
+--+ +-- Pseudodon
| +--------+
| | +-- Sundadontina
+--+
| +-------- Thaiconcha
+--+
| +----- Pilsbryoconcha_exilis__2 **
+--+
| +-- Pilsbryoconcha_exilis__1 **
+--+
+-- Pilsbryoconcha **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 4 | 137 | 10 | 5 | -- | +-------- Pilsbryoconcha_exilis__1 **
+--+
| | +----- Pilsbryoconcha_exilis__3 **
| +--+
| | +-- Pilsbryoconcha_exilis__2 **
| +--+
--+ +-- Pilsbryoconcha_carinifera **
|
| +-- Sundadontina_tumida
| +-----+
| | +-- Pseudodon
| |
+--+ +-- Nyeinchanconcha
| +--+
| | +-- Sundadontina_cumingii
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 2 | 89 | 1 | 1 | 9 | +----- Pilsbryoconcha_exilis **
--+
| +-- Pseudodon
+--+
+-- Pilsbryoconcha_carinifera **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 119 | 4 | 4 | -- | +-- Pilsbryoconcha_exilis **
+-----+
| +-- Pilsbryoconcha_carinifera **
--+
| +----- Pseudodon
| |
+--+ +-- Nyeinchanconcha
| |
+--+-- Bineurus
|
+-- Sundadontina
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 8 | 0 | 2 | 81 | 6 | 2 | 14 | +-- Bineurus_mouhotii
|
+-- Pseudodon_vagulus
|
+-- Pseudodon_mekongi
|
+-- Pilsbryoconcha_exilis **
--+
+-- Pilsbryoconcha_carinifera **
|
+-- Bineurus_loeiensis
|
+-- Bineurus_exilis
|
+-- Pseudodon_vondembuschianus
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 4 | 195 | 10 | 5 | -- | +-------- Pilsbryoconcha_exilis__3 **
+--+
| | +----- Pilsbryoconcha_exilis__2 **
| +--+
| | +-- Pilsbryoconcha_carinifera **
| +--+
--+ +-- Pilsbryoconcha_exilis__1 **
|
| +-- Sundadontina_tumida
| +-----+
| | +-- Pseudodon
| |
+--+ +-- Nyeinchanconcha
| +--+
| | +-- Sundadontina_cumingii
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 4 | 197 | 10 | 5 | -- | +-------- Pilsbryoconcha_exilis__3 **
+--+
| | +----- Pilsbryoconcha_exilis__2 **
| +--+
| | +-- Pilsbryoconcha_exilis__1 **
| +--+
--+ +-- Pilsbryoconcha_carinifera **
|
| +-- Pseudodon
| +-----+
| | +-- Sundadontina
+--+
| +----- Nyeinchanconcha
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 2 | 89 | 1 | 1 | 9 | +----- Pilsbryoconcha_exilis **
--+
| +-- Pseudodon
+--+
+-- Pilsbryoconcha_carinifera **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 78 | 3 | 3 | -- | +-- Pilsbryoconcha_exilis **
+-----+
| +-- Pilsbryoconcha_acuta **
--+
| +----- Pseudodon
+--+
| +-- Bineurus
+--+
+-- Nyeinchanconcha
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 7 | 4 | -- | +-- Pilsbryoconcha_exilis **
+-----+
| +-- Pseudodon
--+
| +----- Sundadontina
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 0 | 0 | -- | -- Pilsbryoconcha_exilis **
Konopleva et al., 2017 [Publ 3807], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 4 | -- | +-------- Sundadontina
|
--+ +-- Nyeinchanconcha
| +--+
| | +-- Bineurus
+--+
| +-- Pilsbryoconcha_carinifera **
+--+
+-- Pseudodon
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Froufe et al., 2019 [Publ 3936], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Froufe et al., 2019 [Publ 3936], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 5 | 4 | -- | +----------- Pilsbryoconcha_carinifera **
--+
| +-------- Pseudodon
+--+
| +----- Sundadontina
+--+
| +-- Nyeinchanconcha
+--+
+-- Bineurus
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Wu et al., 2024 [Publ 4350], fig. 6-7 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 5 | 3 | -- | +-------- Pilsbryoconcha_exilis **
--+
| +----- Pseudodon
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Wu et al., 2024 [Publ 4388], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 5 | 3 | -- | +-------- Pilsbryoconcha_exilis **
--+
| +----- Pseudodon
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 3 | 3 | -- | +-------- Pilsbryoconcha_exilis **
--+
| +----- Bineurus
+--+
| +-- Pseudodon
+--+
+-- Sundadontina
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 87 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 51 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Jeratthitikul et al., 2019 [Publ 3911], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | -- Pilsbryoconcha_exilis **
Wu et al., 2024 [Publ 4350], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 5 | 3 | -- | +-------- Pilsbryoconcha_exilis **
--+
| +----- Bineurus
+--+
| +-- Thaiconcha
+--+
+-- Pseudodon
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Wu et al., 2024 [Publ 4350], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 5 | 3 | -- | +-------- Pilsbryoconcha_exilis **
--+
| +----- Pseudodon
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 84 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 10 | 6 | -- | +----------- Bineurus
|
| +----- Namkongnaia
--+ +--+
| | | +-- Sundadontina
| | +--+
+--+ +-- Nyeinchanconcha
|
| +----- Pseudodon
+--+
| +-- Pilsbryoconcha_exilis **
+--+
+-- Thaiconcha
Matos et al., 2024 [Publ 4384], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Kongim et al., 2023 [Publ 4290], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | +-- Namkongnaia
--+
+-- Pilsbryoconcha_exilis **
Zieritz et al., 2021 [Publ 4218], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 31 | 6 | 4 | -- | +----------- Pilsbryoconcha_exilis **
--+
| +-------- Pseudodon
+--+
| +----- Sundadontina
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Konopleva et al., 2021 [Publ 4213], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 2 | 0 | 2 | 26 | 18 | 6 | -- | +----- Pilsbryoconcha_exilis__2 **
+-----+
| | +-- Pilsbryoconcha_exilis__1 **
| +--+
| +-- Namkongnaia
--+
| +-- Pseudodon
| +-----+
| | +-- Sundadontina
+--+
| +----- Nyeinchanconcha
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Liu et al., 2024 [Publ 4380], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 5 | 3 | -- | +-------- Pilsbryoconcha_exilis **
--+
| +----- Pseudodon
+--+
| +-- Thaiconcha
+--+
+-- Bineurus
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 3 | 3 | -- | +-------- Bineurus
--+
| +----- Pilsbryoconcha_exilis **
+--+
| +-- Pseudodon
+--+
+-- Sundadontina
Wu et al., 2021 [Publ 4215], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 2 | 2 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_carinifera **
Dai et al., 2025 [Publ 4441], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 10 | 5 | -- | +-------------- Pilsbryoconcha_exilis **
--+
| +----------- Pseudodon
+--+
| +-------- Sundadontina
+--+
| +----- Nyeinchanconcha
+--+
| +-- Bineurus
+--+
+-- Thaiconcha
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Zhang et al., 2024 [Publ 4397], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Wu et al., 2024 [Publ 4350], fig. 5c iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 5 | 3 | -- | +-------- Pilsbryoconcha_exilis **
|
--+ +----- Bineurus_anodontinum
| |
+--+----- Bineurus_loeiensis
|
| +-- Thaiconcha
+--+
+-- Pseudodon
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Matos et al., 2024 [Publ 4383], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Dai et al., 2024 [Publ 4389], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 10 | 5 | -- | +-------------- Pilsbryoconcha_exilis **
--+
| +----------- Pseudodon
+--+
| +-------- Sundadontina
+--+
| +----- Nyeinchanconcha
+--+
| +-- Bineurus
+--+
+-- Thaiconcha
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- | +-- Pilsbryoconcha_exilis **
--+
+-- Pseudodon
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 1 | 1 | -- | +-- Pseudodon
--+
+-- Pilsbryoconcha_exilis **
Zieritz et al., 2020 [Publ 4102], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- | -- Pilsbryoconcha_carinifera **
|
