MUSSELp

MUSSELpdb

Introduction

Browse

d'basing

Search the MUSSELpdb

Taxonomy

all genera

Unionidae

Margaritiferidae

Hyriidae

Etheriidae

Mycetopodidae

Iridinidae


Species & Genus Richness by
Taxonomy

Geography

Summary

North America

Central America

South America

Afrotropics

North Eurasia

East Asia

Australasia


Species Richness & Endemism by
Geography

Data last updated
Thu 18 Dec 2025

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Rectidentini

genus Ensidens Frierson, 1911: cladomics

6 Recent valid species (6 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 6 (6 total)
analyzed in 45 cladograms in 31 publications
maximum number of terminals in an analysis = 6 (mean = 2.69, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 20
maximum number of included terminals = 6, (mean = 4.8 median = 5)
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Ensidens cladogram distances
PCoA Ordination

NJ Tree of Ensidens cladogram distances
NJ Tree

Cladograms

Konopleva et al., 2019 [Publ 4041], fig. 3

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
6 12 0 7 28 6 2 0 
  +-- Ensidens_ingallsianus **
  |                        
  +-- Ensidens_telus **       
  |                        
  +-- Ensidens_dugasti **     
  |                        
  +-- Hyriopsis_khoratensis
  |                        
  +-- Hyriopsis_bialata    
  |                        
  +-- Hyriopsis_myersiana  
  |                        
--+-- Hyriopsis_desowitzi  
  |                        
  +-- Rectidens            
  |                        
  +-- Hyriopsis_panhai     
  |                        
  +-- Ensidens_jaculus__1 **  
  |                        
  +-- Ensidens_jaculus__2 **  
  |                        
  +-- Ensidens_spiculus **    
  |                        
  +-- Ensidens_sagittarius **

Konopleva et al., 2023 [Publ 4262], fig. 8

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
5 5 4 6 77 10 2 12 
                       +-- Ensidens_ingallsianus **  
                 +-----+                          
                 |     +-- Ensidens_telus **         
                 |                                
  +--------------+     +-- Ensidens_jaculus **       
  |              |  +--+                          
  |              |  |  +-- Ensidens_spiculus__1 **   
  |              +--+                             
  |                 |  +-- Ensidens_spiculus__2 **   
--+                 +--+                          
  |                    +-- Ensidens_dugasti **       
  |                                               
  |                    +-- Hyriopsis_desowitzi    
  |  +-----------------+                          
  |  |                 +-- Hyriopsis_delaportei   
  +--+                                            
     |  +----------------- Rectidens              
     |  |                                         
     +--+  +-------------- Hyriopsis_panhai       
        |  |                                      
        +--+  +----------- Hyriopsis_myersiana    
           |  |                                   
           +--+        +-- Hyriopsis_khoratensis  
              |  +-----+                          
              |  |     +-- Hyriopsis_bialata      
              +--+                                
                 |  +----- Hyriopsis_sakhonensis  
                 +--+                             
                    |  +-- Hyriopsis_phuphaniensis
                    +--+                          
                       +-- Hyriopsis_kratiensis

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
6 6 4 6 112 6 2 11 
     +----------- Hyriopsis_desowitzi  
     |                                 
  +--+  +-------- Rectidens            
  |  |  |                              
  |  +--+     +-- Hyriopsis_khoratensis
  |     |  +--+                        
  |     |  |  +-- Hyriopsis_bialata    
  |     +--+                           
  |        |  +-- Hyriopsis_myersiana  
--+        +--+                        
  |           +-- Hyriopsis_panhai     
  |                                    
  |           +-- Ensidens_telus **       
  |     +-----+                        
  |     |     +-- Ensidens_ingallsianus **
  |     |                              
  +-----+     +-- Ensidens_jaculus **     
        |  +--+                        
        |  |  +-- Ensidens_spiculus **    
        +--+                           
           |  +-- Ensidens_dugasti **     
           +--+                        
              +-- Ensidens_sagittarius **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
6 6 4 6 23 7 2 11 
              +-- Rectidens            
  +-----------+                        
  |           +-- Hyriopsis            
  |                                    
--+           +-- Ensidens_ingallsianus **
  |  +--------+                        
  |  |        +-- Ensidens_telus **       
  +--+                                 
     |  +-------- Ensidens_spiculus **    
     +--+                              
        |  +----- Ensidens_dugasti **     
        +--+                           
           |  +-- Ensidens_jaculus **     
           +--+                        
              +-- Ensidens_sagittarius **

Pfeiffer et al., 2021 [Publ 4126], fig. 30

iTtTbiriSoTroTroSiD
6 6 4 6 1 1 1 11 
  +-------------- Rectidens            
  |                                    
--+           +-- Ensidens_telus **       
  |  +--------+                        
  |  |        +-- Ensidens_ingallsianus **
  +--+                                 
     |  +-------- Ensidens_dugasti **     
     +--+                              
        |  +----- Ensidens_sagittarius ** 
        +--+                           
           |  +-- Ensidens_spiculus **    
           +--+                        
              +-- Ensidens_jaculus **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
6 6 4 6 106 6 2 11 
     +-------------- Hyriopsis_desowitzi  
  +--+                                    
  |  |  +----------- Rectidens            
  |  +--+                                 
  |     |  +-------- Hyriopsis_panhai     
  |     +--+                              
  |        |  +----- Hyriopsis_myersiana  
  |        +--+                           
--+           |  +-- Hyriopsis_khoratensis
  |           +--+                        
  |              +-- Hyriopsis_bialata    
  |                                       
  |              +-- Ensidens_telus **       
  |        +-----+                        
  |        |     +-- Ensidens_ingallsianus **
  |        |                              
  +--------+     +-- Ensidens_dugasti **     
           |  +--+                        
           |  |  +-- Ensidens_sagittarius ** 
           +--+                           
              |  +-- Ensidens_spiculus **    
              +--+                        
                 +-- Ensidens_jaculus **

Muanta et al., 2019 [Publ 3921], fig. 2

iTtTbiriSoTroTroSiD
5 5 4 6 4 4 1 12 
  +-------------- Hyriopsis              
  |                                      
--+           +-- Ensidens_ingallsianus **  
  |  +--------+                          
  |  |        +-- Ensidens_telus **         
  +--+                                   
     |  +-------- Ensidens_sagittarius__2 **
     +--+                                
        |  +----- Ensidens_jaculus **       
        +--+                             
           |  +-- Ensidens_sagittarius__1 **
           +--+                          
              +-- Ensidens_dugasti **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
5 8 0 5 78 3 1 5 
  +-- Ensidens_telus **       
  |                        
  +-- Ensidens_ingallsianus **
  |                        
  +-- Ensidens_jaculus **     
  |                        
  +-- Hyriopsis_desowitzi  
--+                        
  +-- Hyriopsis_myersiana  
  |                        
  +-- Hyriopsis_khoratensis
  |                        
  +-- Ensidens_spiculus **    
  |                        
  +-- Ensidens_sagittarius **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 47 6 2 11 
              +-- Ensidens_ingallsianus **
        +-----+                        
        |     +-- Ensidens_telus **       
  +-----+                              
  |     |  +----- Ensidens_sagittarius ** 
  |     +--+                           
  |        |  +-- Ensidens_jaculus **     
--+        +--+                        
  |           +-- Ensidens_spiculus **    
  |                                    
  |  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+----------- Rectidens            
     |                                 
     |  +-------- Hyriopsis_panhai     
     +--+                              
        |  +----- Hyriopsis_myersiana  
        +--+                           
           |  +-- Hyriopsis_khoratensis
           +--+                        
              +-- Hyriopsis_bialata

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
5 5 3 5 134 6 2 11 
           +----- Hyriopsis_panhai     
           |                           
  +--------+----- Hyriopsis_khoratensis
  |        |                           
  |        |  +-- Hyriopsis_myersiana  
  |        +--+                        
--+           +-- Hyriopsis_bialata    
  |                                    
  |  +----------- Hyriopsis_desowitzi  
  |  |                                 
  |  +----------- Rectidens            
  +--+                                 
     |        +-- Ensidens_telus **       
     |  +-----+                        
     |  |     +-- Ensidens_ingallsianus **
     +--+                              
        |  +----- Ensidens_sagittarius ** 
        +--+                           
           |  +-- Ensidens_spiculus **    
           +--+                        
              +-- Ensidens_jaculus **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 5 194 6 2 11 
              +-- Ensidens_telus **       
        +-----+                        
        |     +-- Ensidens_ingallsianus **
  +-----+                              
  |     |  +----- Ensidens_sagittarius ** 
  |     +--+                           
  |        |  +-- Ensidens_spiculus **    
--+        +--+                        
  |           +-- Ensidens_jaculus **     
  |                                    
  |  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+  +-------- Rectidens            
     |  |                              
     +--+     +-- Hyriopsis_panhai     
        |  +--+                        
        |  |  +-- Hyriopsis_khoratensis
        +--+                           
           |  +-- Hyriopsis_bialata    
           +--+                        
              +-- Hyriopsis_myersiana

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 3 5 196 6 2 11 
  +-------------- Hyriopsis_desowitzi  
  |                                    
  +-------------- Hyriopsis_myersiana  
  |                                    
  +-------------- Hyriopsis_panhai     
  |                                    
--+           +-- Hyriopsis_bialata    
  +-----------+                        
  |           +-- Hyriopsis_khoratensis
  |                                    
  |  +----------- Rectidens            
  |  |                                 
  +--+        +-- Ensidens_telus **       
     |  +-----+                        
     |  |     +-- Ensidens_ingallsianus **
     +--+                              
        |  +----- Ensidens_sagittarius ** 
        +--+                           
           |  +-- Ensidens_spiculus **    
           +--+                        
              +-- Ensidens_jaculus **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 5 196 6 2 11 
  +----------- Hyriopsis_desowitzi  
  |                                 
  +----------- Rectidens            
  |                                 
  |        +-- Hyriopsis_bialata    
  |        |                        
  |        +-- Hyriopsis_myersiana  
--+--------+                        
  |        +-- Hyriopsis_panhai     
  |        |                        
  |        +-- Hyriopsis_khoratensis
  |                                 
  |        +-- Ensidens_ingallsianus **
  |  +-----+                        
  |  |     +-- Ensidens_telus **       
  +--+                              
     |  +----- Ensidens_sagittarius ** 
     +--+                           
        |  +-- Ensidens_spiculus **    
        +--+                        
           +-- Ensidens_jaculus **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 120 6 2 11 
           +-- Ensidens_telus **       
     +-----+                        
     |     +-- Ensidens_ingallsianus **
  +--+                              
  |  |  +----- Ensidens_jaculus **     
  |  +--+                           
  |     |  +-- Ensidens_spiculus **    
  |     +--+                        
--+        +-- Ensidens_sagittarius ** 
  |                                 
  |  +-------- Hyriopsis_desowitzi  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai     
     |  |                           
     +--+  +-- Hyriopsis_myersiana  
        |  |                        
        +--+-- Hyriopsis_bialata    
           |                        
           +-- Hyriopsis_khoratensis

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 26 6 2 11 
           +-- Ensidens_telus **       
     +-----+                        
     |     +-- Ensidens_ingallsianus **
  +--+                              
  |  |  +----- Ensidens_sagittarius ** 
  |  +--+                           
  |     |  +-- Ensidens_spiculus **    
  |     +--+                        
--+        +-- Ensidens_jaculus **     
  |                                 
  |  +-------- Hyriopsis_desowitzi  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai     
     |  |                           
     +--+  +-- Hyriopsis_khoratensis
        |  |                        
        +--+-- Hyriopsis_bialata    
           |                        
           +-- Hyriopsis_myersiana

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 134 6 2 11 
              +-- Ensidens_telus **       
        +-----+                        
        |     +-- Ensidens_ingallsianus **
  +-----+                              
  |     |  +----- Ensidens_sagittarius ** 
  |     +--+                           
  |        |  +-- Ensidens_spiculus **    
  |        +--+                        
--+           +-- Ensidens_jaculus **     
  |                                    
  |           +-- Hyriopsis_bialata    
  |  +--------+                        
  |  |        +-- Hyriopsis_khoratensis
  +--+                                 
     |  +-------- Hyriopsis_panhai     
     +--+                              
        |  +----- Hyriopsis_myersiana  
        +--+                           
           |  +-- Rectidens            
           +--+                        
              +-- Hyriopsis_desowitzi

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 3 88 1 1 11 
  +-------- Hyriopsis           
--+                             
  |  +----- Ensidens_telus **      
  +--+                          
     |  +-- Ensidens_spiculus **   
     +--+                       
        +-- Ensidens_sagittarius **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 88 1 1 11 
  +-------- Hyriopsis           
--+                             
  |  +----- Ensidens_telus **      
  +--+                          
     |  +-- Ensidens_spiculus **   
     +--+                       
        +-- Ensidens_sagittarius **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
3 3 0 3 77 4 2 10 
  +----- Hyriopsis_desowitzi  
  |                           
  +----- Rectidens            
  |                           
  |  +-- Hyriopsis_khoratensis
--+--+                        
  |  +-- Hyriopsis_myersiana  
  |                           
  |  +-- Ensidens_ingallsianus **
  |  |                        
  +--+-- Ensidens_telus **       
     |                        
     +-- Ensidens_jaculus **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 19 9 2 -- 
                 +-- Ensidens_ingallsianus **  
  +--------------+                          
  |              +-- Ensidens_jaculus **       
  |                                         
--+              +-- Hyriopsis_desowitzi    
  |  +-----------+                          
  |  |           +-- Hyriopsis_delaportei   
  |  |                                      
  +--+  +----------- Rectidens              
     |  |                                   
     |  |     +----- Hyriopsis_myersiana    
     +--+  +--+                             
        |  |  |  +-- Hyriopsis_bialata      
        |  |  +--+                          
        +--+     +-- Hyriopsis_panhai       
           |                                
           |  +----- Hyriopsis_khoratensis  
           +--+                             
              |  +-- Hyriopsis_phuphaniensis
              +--+                          
                 +-- Hyriopsis_sakhonensis

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 45 4 2 -- 
  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Ensidens_ingallsianus **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 6 2 2 -- 
  +-- Ensidens **    
  |               
  +-- Hyriopsis__1
--+               
  +-- Hyriopsis__2
  |               
  +-- Rectidens

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 47 1 1 -- 
  +-- Ensidens_ingallsianus **
--+                        
  +-- Hyriopsis

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 36 3 2 -- 
  +-------- Ensidens_sagittarius **
--+                             
  |  +----- Hyriopsis_bialata   
  +--+                          
     |  +-- Hyriopsis_desowitzi 
     +--+                       
        +-- Rectidens

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 36 3 2 -- 
-- Ensidens_sagittarius **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Ensidens_sagittarius **

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Ensidens_ingallsianus **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 1 1 -- 
-- Ensidens_sagittarius **

Zieritz et al., 2024 [Publ 4381], fig. 6A

iTtTbiriSoTroTroSiD
1 1 0 1 2 2 1 -- 
  +-- Ensidens_ingallsianus **
--+                        
  +-- Rectidens

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
1 1 0 1 36 3 2 -- 
  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 37 3 2 -- 
  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Ensidens_ingallsianus **
--+                        
  +-- Hyriopsis

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +-- Ensidens_ingallsianus **
  |                        
--+-- Hyriopsis            
  |                        
  +-- Rectidens

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
1 1 0 1 36 3 2 -- 
-- Ensidens_sagittarius **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 57 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Ensidens_ingallsianus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Rectidens            
  +--+                        
     +-- Hyriopsis

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 0 0 -- 
-- Ensidens_telus **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 31 3 2 -- 
  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 36 3 2 -- 
  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 35 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens
 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."