Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Rectidentini | Rectidentina

genus Ensidens Frierson, 1911: cladomics

6 Recent valid species (6 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 6 (6 total)
analyzed in 47 cladograms in 33 publications
maximum number of terminals in an analysis = 6 (mean = 2.74, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 21
maximum number of included terminals = 6, (mean = 4.9 median = 5)
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Ensidens cladogram distances
PCoA Ordination

NJ Tree of Ensidens cladogram distances
NJ Tree

Cladograms

Konopleva et al., 2019 [Publ 4041], fig. 3

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
6 12 0 7 28 6 2 0 
  +-- Ensidens_ingallsianus **
  |                        
  +-- Ensidens_telus **       
  |                        
  +-- Ensidens_dugasti **     
  |                        
  +-- Hyriopsis_khoratensis
  |                        
  +-- Hyriopsis_bialata    
  |                        
  +-- Hyriopsis_myersiana  
  |                        
--+-- Hyriopsis_desowitzi  
  |                        
  +-- Rectidens            
  |                        
  +-- Hyriopsis_panhai     
  |                        
  +-- Ensidens_jaculus__1 **  
  |                        
  +-- Ensidens_jaculus__2 **  
  |                        
  +-- Ensidens_spiculus **    
  |                        
  +-- Ensidens_sagittarius **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
6 6 4 6 625 11 2 11 
                 +-- Ensidens_ingallsianus **  
           +-----+                          
           |     +-- Ensidens_telus **         
           |                                
  +--------+     +-- Ensidens_dugasti **       
  |        |  +--+                          
  |        |  |  +-- Ensidens_sagittarius **   
  |        +--+                             
  |           |  +-- Ensidens_jaculus **       
  |           +--+                          
--+              +-- Ensidens_spiculus **      
  |                                         
  |           +----- Rectidens              
  |  +--------+                             
  |  |        |  +-- Hyriopsis_delaportei   
  |  |        +--+                          
  |  |           +-- Hyriopsis_desowitzi    
  +--+                                      
     |  +----------- Hyriopsis_panhai       
     |  |                                   
     |  |        +-- Hyriopsis_bialata      
     +--+  +-----+                          
        |  |     +-- Hyriopsis_khoratensis  
        |  |                                
        +--+     +-- Hyriopsis_kratiensis   
           |  +--+                          
           |  |  +-- Hyriopsis_phuphaniensis
           +--+                             
              |  +-- Hyriopsis_myersiana    
              +--+                          
                 +-- Hyriopsis_sakhonensis

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
6 6 4 6 76 10 2 11 
                       +-- Ensidens_ingallsianus **  
                 +-----+                          
                 |     +-- Ensidens_telus **         
                 |                                
  +--------------+     +-- Ensidens_jaculus **       
  |              |  +--+                          
  |              |  |  +-- Ensidens_spiculus **      
  |              +--+                             
  |                 |  +-- Ensidens_sagittarius **   
--+                 +--+                          
  |                    +-- Ensidens_dugasti **       
  |                                               
  |                    +-- Hyriopsis_desowitzi    
  |  +-----------------+                          
  |  |                 +-- Hyriopsis_delaportei   
  +--+                                            
     |  +----------------- Rectidens              
     |  |                                         
     +--+  +-------------- Hyriopsis_panhai       
        |  |                                      
        +--+  +----------- Hyriopsis_myersiana    
           |  |                                   
           +--+        +-- Hyriopsis_khoratensis  
              |  +-----+                          
              |  |     +-- Hyriopsis_bialata      
              +--+                                
                 |  +----- Hyriopsis_sakhonensis  
                 +--+                             
                    |  +-- Hyriopsis_phuphaniensis
                    +--+                          
                       +-- Hyriopsis_kratiensis

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
6 6 4 6 111 6 2 11 
     +----------- Hyriopsis_desowitzi  
     |                                 
  +--+  +-------- Rectidens            
  |  |  |                              
  |  +--+     +-- Hyriopsis_khoratensis
  |     |  +--+                        
  |     |  |  +-- Hyriopsis_bialata    
  |     +--+                           
  |        |  +-- Hyriopsis_myersiana  
--+        +--+                        
  |           +-- Hyriopsis_panhai     
  |                                    
  |           +-- Ensidens_telus **       
  |     +-----+                        
  |     |     +-- Ensidens_ingallsianus **
  |     |                              
  +-----+     +-- Ensidens_jaculus **     
        |  +--+                        
        |  |  +-- Ensidens_spiculus **    
        +--+                           
           |  +-- Ensidens_dugasti **     
           +--+                        
              +-- Ensidens_sagittarius **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
6 6 4 6 105 6 2 11 
     +-------------- Hyriopsis_desowitzi  
  +--+                                    
  |  |  +----------- Rectidens            
  |  +--+                                 
  |     |  +-------- Hyriopsis_panhai     
  |     +--+                              
  |        |  +----- Hyriopsis_myersiana  
  |        +--+                           
--+           |  +-- Hyriopsis_khoratensis
  |           +--+                        
  |              +-- Hyriopsis_bialata    
  |                                       
  |              +-- Ensidens_telus **       
  |        +-----+                        
  |        |     +-- Ensidens_ingallsianus **
  |        |                              
  +--------+     +-- Ensidens_dugasti **     
           |  +--+                        
           |  |  +-- Ensidens_sagittarius ** 
           +--+                           
              |  +-- Ensidens_spiculus **    
              +--+                        
                 +-- Ensidens_jaculus **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
6 6 4 6 23 7 2 11 
              +-- Rectidens            
  +-----------+                        
  |           +-- Hyriopsis            
  |                                    
--+           +-- Ensidens_ingallsianus **
  |  +--------+                        
  |  |        +-- Ensidens_telus **       
  +--+                                 
     |  +-------- Ensidens_spiculus **    
     +--+                              
        |  +----- Ensidens_dugasti **     
        +--+                           
           |  +-- Ensidens_jaculus **     
           +--+                        
              +-- Ensidens_sagittarius **

Pfeiffer et al., 2021 [Publ 4126], fig. 30

iTtTbiriSoTroTroSiD
6 6 4 6 1 1 1 11 
  +-------------- Rectidens            
  |                                    
--+           +-- Ensidens_telus **       
  |  +--------+                        
  |  |        +-- Ensidens_ingallsianus **
  +--+                                 
     |  +-------- Ensidens_dugasti **     
     +--+                              
        |  +----- Ensidens_sagittarius ** 
        +--+                           
           |  +-- Ensidens_spiculus **    
           +--+                        
              +-- Ensidens_jaculus **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 119 6 2 11 
           +-- Ensidens_telus **       
     +-----+                        
     |     +-- Ensidens_ingallsianus **
  +--+                              
  |  |  +----- Ensidens_jaculus **     
  |  +--+                           
  |     |  +-- Ensidens_spiculus **    
  |     +--+                        
--+        +-- Ensidens_sagittarius ** 
  |                                 
  |  +-------- Hyriopsis_desowitzi  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai     
     |  |                           
     +--+  +-- Hyriopsis_myersiana  
        |  |                        
        +--+-- Hyriopsis_bialata    
           |                        
           +-- Hyriopsis_khoratensis

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
5 5 3 5 46 6 2 11 
              +-- Ensidens_ingallsianus **
        +-----+                        
        |     +-- Ensidens_telus **       
  +-----+                              
  |     |  +----- Ensidens_sagittarius ** 
  |     +--+                           
  |        |  +-- Ensidens_jaculus **     
--+        +--+                        
  |           +-- Ensidens_spiculus **    
  |                                    
  |  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+----------- Rectidens            
     |                                 
     |  +-------- Hyriopsis_panhai     
     +--+                              
        |  +----- Hyriopsis_myersiana  
        +--+                           
           |  +-- Hyriopsis_khoratensis
           +--+                        
              +-- Hyriopsis_bialata

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 3 5 195 6 2 11 
  +-------------- Hyriopsis_desowitzi  
  |                                    
  +-------------- Hyriopsis_myersiana  
  |                                    
  +-------------- Hyriopsis_panhai     
  |                                    
--+           +-- Hyriopsis_bialata    
  +-----------+                        
  |           +-- Hyriopsis_khoratensis
  |                                    
  |  +----------- Rectidens            
  |  |                                 
  +--+        +-- Ensidens_telus **       
     |  +-----+                        
     |  |     +-- Ensidens_ingallsianus **
     +--+                              
        |  +----- Ensidens_sagittarius ** 
        +--+                           
           |  +-- Ensidens_spiculus **    
           +--+                        
              +-- Ensidens_jaculus **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 5 193 6 2 11 
              +-- Ensidens_telus **       
        +-----+                        
        |     +-- Ensidens_ingallsianus **
  +-----+                              
  |     |  +----- Ensidens_sagittarius ** 
  |     +--+                           
  |        |  +-- Ensidens_spiculus **    
--+        +--+                        
  |           +-- Ensidens_jaculus **     
  |                                    
  |  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+  +-------- Rectidens            
     |  |                              
     +--+     +-- Hyriopsis_panhai     
        |  +--+                        
        |  |  +-- Hyriopsis_khoratensis
        +--+                           
           |  +-- Hyriopsis_bialata    
           +--+                        
              +-- Hyriopsis_myersiana

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 133 6 2 11 
              +-- Ensidens_telus **       
        +-----+                        
        |     +-- Ensidens_ingallsianus **
  +-----+                              
  |     |  +----- Ensidens_sagittarius ** 
  |     +--+                           
  |        |  +-- Ensidens_spiculus **    
  |        +--+                        
--+           +-- Ensidens_jaculus **     
  |                                    
  |           +-- Hyriopsis_bialata    
  |  +--------+                        
  |  |        +-- Hyriopsis_khoratensis
  +--+                                 
     |  +-------- Hyriopsis_panhai     
     +--+                              
        |  +----- Hyriopsis_myersiana  
        +--+                           
           |  +-- Rectidens            
           +--+                        
              +-- Hyriopsis_desowitzi

Muanta et al., 2019 [Publ 3921], fig. 2

iTtTbiriSoTroTroSiD
5 5 4 6 4 4 1 12 
  +-------------- Hyriopsis              
  |                                      
--+           +-- Ensidens_ingallsianus **  
  |  +--------+                          
  |  |        +-- Ensidens_telus **         
  +--+                                   
     |  +-------- Ensidens_sagittarius__2 **
     +--+                                
        |  +----- Ensidens_jaculus **       
        +--+                             
           |  +-- Ensidens_sagittarius__1 **
           +--+                          
              +-- Ensidens_dugasti **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
5 8 0 5 77 3 1 5 
  +-- Ensidens_telus **       
  |                        
  +-- Ensidens_ingallsianus **
  |                        
  +-- Ensidens_jaculus **     
  |                        
  +-- Hyriopsis_desowitzi  
--+                        
  +-- Hyriopsis_myersiana  
  |                        
  +-- Hyriopsis_khoratensis
  |                        
  +-- Ensidens_spiculus **    
  |                        
  +-- Ensidens_sagittarius **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 5 195 6 2 11 
  +----------- Hyriopsis_desowitzi  
  |                                 
  +----------- Rectidens            
  |                                 
  |        +-- Hyriopsis_bialata    
  |        |                        
  |        +-- Hyriopsis_myersiana  
--+--------+                        
  |        +-- Hyriopsis_panhai     
  |        |                        
  |        +-- Hyriopsis_khoratensis
  |                                 
  |        +-- Ensidens_ingallsianus **
  |  +-----+                        
  |  |     +-- Ensidens_telus **       
  +--+                              
     |  +----- Ensidens_sagittarius ** 
     +--+                           
        |  +-- Ensidens_spiculus **    
        +--+                        
           +-- Ensidens_jaculus **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
5 5 3 5 133 6 2 11 
           +----- Hyriopsis_panhai     
           |                           
  +--------+----- Hyriopsis_khoratensis
  |        |                           
  |        |  +-- Hyriopsis_myersiana  
  |        +--+                        
--+           +-- Hyriopsis_bialata    
  |                                    
  |  +----------- Hyriopsis_desowitzi  
  |  |                                 
  |  +----------- Rectidens            
  +--+                                 
     |        +-- Ensidens_telus **       
     |  +-----+                        
     |  |     +-- Ensidens_ingallsianus **
     +--+                              
        |  +----- Ensidens_sagittarius ** 
        +--+                           
           |  +-- Ensidens_spiculus **    
           +--+                        
              +-- Ensidens_jaculus **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
5 5 3 5 26 6 2 11 
           +-- Ensidens_telus **       
     +-----+                        
     |     +-- Ensidens_ingallsianus **
  +--+                              
  |  |  +----- Ensidens_sagittarius ** 
  |  +--+                           
  |     |  +-- Ensidens_spiculus **    
  |     +--+                        
--+        +-- Ensidens_jaculus **     
  |                                 
  |  +-------- Hyriopsis_desowitzi  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai     
     |  |                           
     +--+  +-- Hyriopsis_khoratensis
        |  |                        
        +--+-- Hyriopsis_bialata    
           |                        
           +-- Hyriopsis_myersiana

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3 3 1 3 88 1 1 11 
  +-------- Hyriopsis           
--+                             
  |  +----- Ensidens_telus **      
  +--+                          
     |  +-- Ensidens_spiculus **   
     +--+                       
        +-- Ensidens_sagittarius **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3 3 1 3 88 1 1 11 
  +-------- Hyriopsis           
--+                             
  |  +----- Ensidens_telus **      
  +--+                          
     |  +-- Ensidens_spiculus **   
     +--+                       
        +-- Ensidens_sagittarius **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
3 3 0 3 77 4 2 10 
  +----- Hyriopsis_desowitzi  
  |                           
  +----- Rectidens            
  |                           
  |  +-- Hyriopsis_khoratensis
--+--+                        
  |  +-- Hyriopsis_myersiana  
  |                           
  |  +-- Ensidens_ingallsianus **
  |  |                        
  +--+-- Ensidens_telus **       
     |                        
     +-- Ensidens_jaculus **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iTtTbiriSoTroTroSiD
2 2 0 2 19 9 2 -- 
                 +-- Ensidens_ingallsianus **  
  +--------------+                          
  |              +-- Ensidens_jaculus **       
  |                                         
--+              +-- Hyriopsis_desowitzi    
  |  +-----------+                          
  |  |           +-- Hyriopsis_delaportei   
  |  |                                      
  +--+  +----------- Rectidens              
     |  |                                   
     |  |     +----- Hyriopsis_myersiana    
     +--+  +--+                             
        |  |  |  +-- Hyriopsis_bialata      
        |  |  +--+                          
        +--+     +-- Hyriopsis_panhai       
           |                                
           |  +----- Hyriopsis_khoratensis  
           +--+                             
              |  +-- Hyriopsis_phuphaniensis
              +--+                          
                 +-- Hyriopsis_sakhonensis

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 34 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 30 3 2 -- 
  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Ensidens_ingallsianus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Rectidens            
  +--+                        
     +-- Hyriopsis

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 6 2 2 -- 
  +-- Ensidens **    
  |               
  +-- Hyriopsis__1
--+               
  +-- Hyriopsis__2
  |               
  +-- Rectidens

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Ensidens_ingallsianus **
--+                        
  +-- Hyriopsis

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 0 0 -- 
-- Ensidens_telus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +-- Ensidens_ingallsianus **
  |                        
--+-- Hyriopsis            
  |                        
  +-- Rectidens

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 45 1 1 -- 
  +-- Ensidens_ingallsianus **
--+                        
  +-- Hyriopsis

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 47 1 1 -- 
  +-- Ensidens_ingallsianus **
--+                        
  +-- Hyriopsis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 1 1 -- 
-- Ensidens_sagittarius **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
  +-------- Ensidens_sagittarius **
--+                             
  |  +----- Hyriopsis_bialata   
  +--+                          
     |  +-- Hyriopsis_desowitzi 
     +--+                       
        +-- Rectidens

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
1 1 0 1 44 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Ensidens_ingallsianus **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 44 4 2 -- 
  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 8 0 0 -- 
-- Ensidens_ingallsianus **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 0 0 -- 
-- Ensidens_sagittarius **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
-- Ensidens_sagittarius **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 58 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Zieritz et al., 2024 [Publ 4381], fig. 6A

iTtTbiriSoTroTroSiD
1 1 0 1 2 2 1 -- 
  +-- Ensidens_ingallsianus **
--+                        
  +-- Rectidens

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
  +----- Ensidens_sagittarius **
--+                          
  |  +-- Rectidens           
  +--+                       
     +-- Hyriopsis

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
-- Ensidens_sagittarius **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 2 -- 
  +----- Ensidens_ingallsianus **
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens