Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Limnoscaphini | Limnoscaphina

genus Ensidens Frierson, 1911: cladomics

6 Recent valid species (6 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 6 (6 total)
analyzed in 47 cladograms in 33 publications
maximum number of terminals in an analysis = 6 (mean = 2.74, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 21
maximum number of included terminals = 6, (mean = 4.9 median = 5)
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Ensidens cladogram distances
PCoA Ordination

NJ Tree of Ensidens cladogram distances
NJ Tree

Cladograms

Pfeiffer et al., 2021 [Publ 4126], fig. 9

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 23 7 2 **
Cladomics cladogram Rectidens Hyriopsis Ensidens ingallsianus Ensidens telus Ensidens spiculus Ensidens dugasti Ensidens jaculus Ensidens sagittarius

Konopleva et al., 2019 [Publ 4041], fig. 3

cst iT tT bi riS oT roT roS iD
0.500 6 12 0 6 28 6 2 11.0
Cladomics cladogram Yaukthwa Lens Trapezoideus Physunio Ensidens ingallsianus Ensidens telus Ensidens dugasti Hyriopsis khoratensis Hyriopsis bialata Hyriopsis myersiana Hyriopsis desowitzi Rectidens Hyriopsis panhai Ensidens jaculus 1 Ensidens jaculus 2 Ensidens spiculus Ensidens sagittarius

Bolotov et al., 2020 [Publ 4100], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 6 6 4 6 111 6 2 4.0
Cladomics cladogram Hyriopsis desowitzi Rectidens Hyriopsis khoratensis Hyriopsis bialata Hyriopsis myersiana Hyriopsis panhai Ensidens telus Ensidens ingallsianus Ensidens jaculus Ensidens spiculus Ensidens dugasti Ensidens sagittarius

Bolotov et al., 2020 [Publ 4100], fig. S1

cst iT tT bi riS oT roT roS iD
0.750 6 6 4 6 105 6 2 4.0
Cladomics cladogram Hyriopsis desowitzi Rectidens Hyriopsis panhai Hyriopsis myersiana Hyriopsis khoratensis Hyriopsis bialata Ensidens telus Ensidens ingallsianus Ensidens dugasti Ensidens sagittarius Ensidens spiculus Ensidens jaculus

Konopleva et al., 2023 [Publ 4262], fig. 8

cst iT tT bi riS oT roT roS iD
0.750 6 6 4 6 76 10 2 4.0
Cladomics cladogram Ensidens ingallsianus Ensidens telus Ensidens jaculus Ensidens spiculus Ensidens sagittarius Ensidens dugasti Hyriopsis desowitzi Hyriopsis delaportei Rectidens Hyriopsis panhai Hyriopsis myersiana Hyriopsis khoratensis Hyriopsis bialata Hyriopsis sakhonensis Hyriopsis phuphaniensis Hyriopsis kratiensis

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.750 6 6 4 6 627 11 2 4.0
Cladomics cladogram Ensidens ingallsianus Ensidens telus Ensidens dugasti Ensidens sagittarius Ensidens jaculus Ensidens spiculus Rectidens Hyriopsis delaportei Hyriopsis desowitzi Hyriopsis panhai Hyriopsis bialata Hyriopsis khoratensis Hyriopsis kratiensis Hyriopsis phuphaniensis Hyriopsis myersiana Hyriopsis sakhonensis

Pfeiffer et al., 2021 [Publ 4126], fig. 30

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 1 1 1 4.0
Cladomics cladogram Rectidens Ensidens telus Ensidens ingallsianus Ensidens dugasti Ensidens sagittarius Ensidens spiculus Ensidens jaculus

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 5 134 6 2 3.0
Cladomics cladogram Ensidens telus Ensidens ingallsianus Ensidens sagittarius Ensidens spiculus Ensidens jaculus Hyriopsis bialata Hyriopsis khoratensis Hyriopsis panhai Hyriopsis myersiana Rectidens Hyriopsis desowitzi

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 5 134 6 2 3.0
Cladomics cladogram Hyriopsis panhai Hyriopsis khoratensis Hyriopsis myersiana Hyriopsis bialata Hyriopsis desowitzi Rectidens Ensidens telus Ensidens ingallsianus Ensidens sagittarius Ensidens spiculus Ensidens jaculus

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 5 120 6 2 3.0
Cladomics cladogram Ensidens telus Ensidens ingallsianus Ensidens jaculus Ensidens spiculus Ensidens sagittarius Hyriopsis desowitzi Rectidens Hyriopsis panhai Hyriopsis myersiana Hyriopsis bialata Hyriopsis khoratensis

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 5 196 6 2 3.0
Cladomics cladogram Hyriopsis desowitzi Rectidens Hyriopsis bialata Hyriopsis myersiana Hyriopsis panhai Hyriopsis khoratensis Ensidens ingallsianus Ensidens telus Ensidens sagittarius Ensidens spiculus Ensidens jaculus

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 5 194 6 2 3.0
Cladomics cladogram Ensidens telus Ensidens ingallsianus Ensidens sagittarius Ensidens spiculus Ensidens jaculus Hyriopsis desowitzi Rectidens Hyriopsis panhai Hyriopsis khoratensis Hyriopsis bialata Hyriopsis myersiana

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 5 196 6 2 3.0
Cladomics cladogram Hyriopsis desowitzi Hyriopsis myersiana Hyriopsis panhai Hyriopsis bialata Hyriopsis khoratensis Rectidens Ensidens telus Ensidens ingallsianus Ensidens sagittarius Ensidens spiculus Ensidens jaculus

Konopleva et al., 2019 [Publ 3900], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 5 46 6 2 3.0
Cladomics cladogram Ensidens ingallsianus Ensidens telus Ensidens sagittarius Ensidens jaculus Ensidens spiculus Hyriopsis desowitzi Rectidens Hyriopsis panhai Hyriopsis myersiana Hyriopsis khoratensis Hyriopsis bialata

Zieritz et al., 2020 [Publ 4102], fig. 4

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 5 26 6 2 3.0
Cladomics cladogram Ensidens telus Ensidens ingallsianus Ensidens sagittarius Ensidens spiculus Ensidens jaculus Hyriopsis desowitzi Rectidens Hyriopsis panhai Hyriopsis khoratensis Hyriopsis bialata Hyriopsis myersiana

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
0.333 5 8 0 5 77 3 1 7.0
Cladomics cladogram Ensidens telus Ensidens ingallsianus Ensidens jaculus Hyriopsis desowitzi Hyriopsis myersiana Hyriopsis khoratensis Ensidens spiculus Ensidens sagittarius Lens eximius Physunio modelli 1 Yaukthwa nesemanni Yaukthwa panhai Lens comptus Lens novoselovi Physunio modelli 2 Physunio Yaukthwa inlenensis Lens contradens Lens micropterus

Muanta et al., 2019 [Publ 3921], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 4 5 4 4 1 4.0
Cladomics cladogram Hyriopsis Ensidens ingallsianus Ensidens telus Ensidens sagittarius 2 Ensidens jaculus Ensidens sagittarius 1 Ensidens dugasti

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
0.750 3 3 0 3 78 4 2 4.0
Cladomics cladogram Hyriopsis desowitzi Rectidens Hyriopsis khoratensis Hyriopsis myersiana Ensidens ingallsianus Ensidens telus Ensidens jaculus

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 88 1 1 3.0
Cladomics cladogram Hyriopsis Ensidens telus Ensidens spiculus Ensidens sagittarius

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 88 1 1 3.0
Cladomics cladogram Hyriopsis Ensidens telus Ensidens spiculus Ensidens sagittarius

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

cst iT tT bi riS oT roT roS iD
0.750 2 2 0 2 19 9 2 --
Cladomics cladogram Ensidens ingallsianus Ensidens jaculus Hyriopsis desowitzi Hyriopsis delaportei Rectidens Hyriopsis myersiana Hyriopsis bialata Hyriopsis panhai Hyriopsis khoratensis Hyriopsis phuphaniensis Hyriopsis sakhonensis

Pfeiffer et al., 2018 [Publ 3827], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 6 2 2 --
Cladomics cladogram Ensidens Hyriopsis 1 Hyriopsis 2 Rectidens

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 2 2 --
Cladomics cladogram Ensidens ingallsianus Hyriopsis Rectidens

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 2 2 --
Cladomics cladogram Ensidens ingallsianus Hyriopsis Rectidens

Zieritz et al., 2021 [Publ 4218], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 3 2 --
Cladomics cladogram Ensidens sagittarius Rectidens Hyriopsis

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 2 2 --
Cladomics cladogram Ensidens ingallsianus Hyriopsis Rectidens

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 34 2 2 --
Cladomics cladogram Ensidens ingallsianus Hyriopsis Rectidens

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 2 2 --
Cladomics cladogram Ensidens ingallsianus Hyriopsis Rectidens

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 2 2 --
Cladomics cladogram Ensidens ingallsianus Rectidens Hyriopsis

Wu et al., 2024 [Publ 4350], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 3 2 --
Cladomics cladogram Ensidens sagittarius Rectidens Hyriopsis

Wu et al., 2024 [Publ 4350], fig. 5b

cst iT tT bi riS oT roT roS iD
0.500 1 1 0 1 35 3 2 --
Cladomics cladogram Ensidens sagittarius Hyriopsis Yaukthwa elongatula Trapezoideus Physunio Yaukthwa inlenensis Yaukthwa paiensis Lens

Wu et al., 2024 [Publ 4350], fig. 5c

cst iT tT bi riS oT roT roS iD
0.500 1 1 0 1 35 3 2 --
Cladomics cladogram Ensidens sagittarius Hyriopsis desowitzi Rectidens Hyriopsis bialata Khairuloconcha Physunio Yaukthwa paiensis Yaukthwa inlenensis Trapezoideus Yaukthwa elongatula Lens

Wu et al., 2024 [Publ 4350], fig. 6-7

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 3 2 --
Cladomics cladogram Ensidens sagittarius Rectidens Hyriopsis

Liu et al., 2024 [Publ 4380], fig. 5

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 35 3 2 --
Cladomics cladogram Ensidens sagittarius Hyriopsis bialata Hyriopsis desowitzi Rectidens

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 4 2 --
Cladomics cladogram Ensidens sagittarius Rectidens Hyriopsis

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 58 2 2 --
Cladomics cladogram Ensidens ingallsianus Hyriopsis Rectidens

Wu et al., 2024 [Publ 4388], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 3 2 --
Cladomics cladogram Ensidens sagittarius Rectidens Hyriopsis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 121 1 1 --
Cladomics cladogram Lens Ensidens sagittarius Physunio Rectidens Yaukthwa

Dai et al., 2023 [Publ 4285], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 1 1 --
Cladomics cladogram Hyriopsis Ensidens ingallsianus

Dai et al., 2023 [Publ 4285], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 1 1 --
Cladomics cladogram Hyriopsis Ensidens ingallsianus

Zieritz et al., 2024 [Publ 4381], fig. 6A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 2 1 --
Cladomics cladogram Ensidens ingallsianus Rectidens

Dai et al., 2024 [Publ 4389], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 1 1 --
Cladomics cladogram Ensidens ingallsianus Hyriopsis

Dai et al., 2025 [Publ 4441], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 1 1 --
Cladomics cladogram Ensidens ingallsianus Hyriopsis

Chen et al., 2025 [Publ 4663], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 1 1 --
Cladomics cladogram Ensidens ingallsianus Hyriopsis

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 0 0 --
Cladomics cladogram Ensidens telus Lens Physunio

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Ensidens sagittarius Lens Physunio

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 8 0 0 --
Cladomics cladogram Ensidens ingallsianus Physunio Lens