Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Cristaria Schumacher, 1817: cladomics

8 Recent valid species (11 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 8 (11 total)
analyzed in 131 cladograms in 88 publications
maximum number of terminals in an analysis = 7 (mean = 1.49, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 24
maximum number of included terminals = 7, (mean = 3.67 median = 3)
proportion of cladograms that support basic monophyly = 0.96

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Cristaria cladogram distances
PCoA Ordination

NJ Tree of Cristaria cladogram distances
NJ Tree

Cladograms

Dai et al., 2026 [Publ 4757], fig. 3

** Top Monophyly Cladogram **

cstiTtTbiriSoTroTroSiD
1.000 7 7 6 7 27 23 6 ** 
                    +----- Amuranodonta         
  +-----------------+                           
  |                 |  +-- Anemina              
  |                 +--+                        
  |                    +-- Buldowskia           
--+                                             
  |                 +----- Pletholophus         
  |  +--------------+                           
  |  |              |  +-- Beringiana           
  |  |              +--+                        
  +--+                 +-- Sinanodonta          
     |                                          
     |  +----------------- Cristaria_wangyucheni **
     |  |                                       
     +--+  +-------------- Cristaria_bellua **     
        |  |                                    
        +--+  +----------- Cristaria_clessini **   
           |  |                                 
           +--+        +-- Cristaria_plicata **    
              |  +-----+                        
              |  |     +-- Cristaria_truncata **   
              +--+                              
                 |  +----- Cristaria_radiata__2 ** 
                 |  |                           
                 +--+----- Cristaria_radiata__3 ** 
                    |                           
                    |  +-- Cristaria_liboi **      
                    +--+                        
                       +-- Cristaria_radiata__1 **

Chen et al., 2026 [Publ 4805], fig. 4

** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
1.000 6 6 4 6 28 27 8 5.0 
        +----------- Cristaria_clessini **
        |                              
  +-----+  +-------- Cristaria_bellua **  
  |     |  |                           
  |     +--+     +-- Cristaria_plicata ** 
  |        |  +--+                     
  |        |  |  +-- Cristaria_truncata **
  |        +--+                        
--+           |  +-- Cristaria_radiata ** 
  |           +--+                     
  |              +-- Cristaria_liboi **   
  |                                    
  |  +-------------- Beringiana        
  |  |                                 
  |  |        +----- Mimo              
  +--+  +-----+                        
     |  |     |  +-- Acudonta          
     |  |     +--+                     
     +--+        +-- Sinanodonta       
        |                              
        |  +-------- Pletholophus      
        +--+                           
           |  +----- Anemina           
           +--+                        
              |  +-- Amuranodonta      
              +--+                     
                 +-- Buldowskia

Tumpeesuwan et al., 2026 [Publ 4845], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 7 7 5 7 3 3 3 8.0 
                 +-- Beringiana        
  +--------------+                     
  |              +-- Sinanodonta       
  |                                    
--+  +-------------- Buldowskia        
  |  |                                 
  |  |           +-- Cristaria_liboi **   
  +--+  +--------+                     
     |  |        +-- Cristaria_radiata ** 
     |  |                              
     +--+        +-- Cristaria_plicata ** 
        |  +-----+                     
        |  |     +-- Cristaria_truncata **
        +--+                           
           |  +----- Cristaria_clessini **
           +--+                        
              |  +-- Cristaria_hospes **  
              +--+                     
                 +-- Cristaria_bellua **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cstiTtTbiriSoTroTroSiD
0.900 6 6 4 6 627 27 8 7.0 
                                   +-- Cristaria_liboi **             
                             +-----+                               
                             |     +-- Cristaria_radiata **           
                             |                                     
  +--------------------------+     +-- Cristaria_bellua **            
  |                          |  +--+                               
  |                          |  |  +-- Cristaria_clessini **          
  |                          +--+                                  
  |                             |  +-- Cristaria_plicata **           
  |                             +--+                               
  |                                +-- Cristaria_truncata **          
  |                                                                
  |     +----------------------------- Lepidodesma                 
  |     |                                                          
  |     |                 +----------- Lanceolaria                 
  |     |  +--------------+                                        
  |  +--+  |              |  +-------- Pseudoacuticosta_retiaria__2
  |  |  |  |              +--+                                     
  |  |  |  |                 |  +----- Acuticosta_chinensis__2     
  |  |  |  |                 +--+                                  
--+  |  +--+                    |  +-- Acuticosta_chinensis__1     
  |  |     |                    +--+                               
  |  |     |                       +-- Pseudoacuticosta_retiaria__1
  |  |     |                                                       
  |  |     |  +----------------------- Globunio                    
  |  |     |  |                                                    
  |  |     +--+  +-------------------- Scriptolamprotula           
  |  |        |  |                                                 
  |  |        +--+  +----------------- Middendorffinaia            
  |  |           |  |                                              
  |  |           +--+  +-------------- Unio                        
  |  |              |  |                                           
  |  |              |  |           +-- Schistodesmus               
  |  |              +--+  +--------+                               
  |  |                 |  |        +-- Nodularia                   
  |  |                 |  |                                        
  +--+                 +--+        +-- Tchangsinaia                
     |                    |  +-----+                               
     |                    |  |     +-- Inversiunio                 
     |                    +--+                                     
     |                       |  +----- Pseudocuneopsis             
     |                       +--+                                  
     |                          |  +-- Diaurora                    
     |                          +--+                               
     |                             +-- Cuneopsis                   
     |                                                             
     |                       +-------- Pseudodontoideus            
     |  +--------------------+                                     
     |  |                    |  +----- Lasmigona                   
     |  |                    +--+                                  
     |  |                       |  +-- Pterosyna                   
     |  |                       +--+                               
     |  |                          +-- Arcidens                    
     |  |                                                          
     |  |                          +-- Utterbackia                 
     |  |           +--------------+                               
     +--+           |              +-- Pyganodon                   
        |  +--------+                                              
        |  |        |  +-------------- Utterbackiana               
        |  |        |  |                                           
        |  |        +--+  +----------- Simpsonaias                 
        |  |           |  |                                        
        |  |           |  |        +-- Alasminota                  
        |  |           +--+  +-----+                               
        |  |              |  |     +-- Alasmidonta                 
        |  |              |  |                                     
        +--+              +--+     +-- Prolasmidonta               
           |                 |  +--+                               
           |                 |  |  +-- Platynaias                  
           |                 +--+                                  
           |                    |  +-- Anodontoides                
           |                    +--+                               
           |                       +-- Strophitus                  
           |                                                       
           |  +----------------------- Anodonta_anatina            
           |  |                                                    
           +--+  +-------------------- Pseudanodonta               
              |  |                                                 
              +--+  +----------------- Brachyanodon_oregonensis__2 
                 |  |                                              
                 |  |           +----- Anodonta_cygnea             
                 +--+  +--------+                                  
                    |  |        |  +-- Anodonta_seddonae           
                    |  |        +--+                               
                    +--+           +-- Anodonta_exulcerata         
                       |                                           
                       |  +----------- Brachyanodon_impura         
                       |  |                                        
                       +--+  +-------- Brachyanodon                
                          |  |                                     
                          +--+  +----- Brachyanodon_nuttalliana__1 
                             |  |                                  
                             +--+----- Brachyanodon_nuttalliana__2 
                                |                                  
                                |  +-- Brachyanodon_californiensis 
                                +--+                               
                                   +-- Brachyanodon_oregonensis__1

Wu et al., 2025 [Publ 4442], fig. 2

cstiTtTbiriSoTroTroSiD
0.889 6 6 4 6 29 27 7 5.0 
                 +-- Cristaria_clessini **         
           +-----+                              
           |     +-- Cristaria_bellua **           
           |                                    
  +--------+     +-- Cristaria_plicata **          
  |        |  +--+                              
  |        |  |  +-- Cristaria_truncata **         
  |        +--+                                 
  |           |  +-- Cristaria_radiata **          
  |           +--+                              
--+              +-- Cristaria_liboi **            
  |                                             
  |           +----- Beringiana                 
  |  +--------+                                 
  |  |        |  +-- Acudonta                   
  |  |        +--+                              
  |  |           +-- Sinanodonta                
  +--+                                          
     |        +----- Pletholophus_guangzhouensis
     |  +-----+                                 
     |  |     |  +-- Pletholophus_tenuis        
     |  |     +--+                              
     +--+        +-- Pletholophus_reinianus     
        |                                       
        |  +-------- Pletholophus_honglinhensis 
        +--+                                    
           |  +----- Anemina                    
           +--+                                 
              |  +-- Amuranodonta               
              +--+                              
                 +-- Buldowskia

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cstiTtTbiriSoTroTroSiD
1.000 5 5 3 5 51 8 5 8.0 
           +----- Anemina           
  +--------+                        
  |        |  +-- Amuranodonta      
  |        +--+                     
  |           +-- Buldowskia        
--+                                 
  |           +-- Beringiana        
  |  +--------+                     
  |  |        +-- Sinanodonta       
  +--+                              
     |  +-------- Cristaria_radiata ** 
     |  |                           
     +--+     +-- Cristaria_truncata **
        |  +--+                     
        |  |  +-- Cristaria_plicata ** 
        +--+                        
           |  +-- Cristaria_bellua **  
           +--+                     
              +-- Cristaria_clessini **

Xiang et al., 2025 [Publ 4433], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 24 23 7 5.0 
  +----------------- Pletholophus      
  |                                    
  |           +----- Beringiana        
--+  +--------+                        
  |  |        |  +-- Mimo              
  |  |        +--+                     
  |  |           +-- Sinanodonta       
  +--+                                 
     |        +----- Anemina           
     |  +-----+                        
     |  |     |  +-- Amuranodonta      
     |  |     +--+                     
     +--+        +-- Buldowskia        
        |                              
        |  +-------- Cristaria_bellua **  
        +--+                           
           |  +----- Cristaria_clessini **
           +--+                        
              |  +-- Cristaria_plicata ** 
              +--+                     
                 +-- Cristaria_truncata **

Bogan et al., 2023 [Publ 4295], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 27 23 6 5.0 
     +-------- Cristaria_bellua **  
  +--+                           
  |  |  +----- Cristaria_clessini **
  |  +--+                        
  |     |  +-- Cristaria_plicata ** 
  |     +--+                     
--+        +-- Cristaria_truncata **
  |                              
  |     +----- Amuranodonta      
  |  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  +--+     +-- Buldowskia        
     |                           
     |  +----- Pletholophus      
     +--+                        
        |  +-- Beringiana        
        +--+                     
           +-- Sinanodonta

Dai et al., 2024 [Publ 4385], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 27 23 6 5.0 
           +----- Amuranodonta      
  +--------+                        
  |        |  +-- Anemina           
  |        +--+                     
  |           +-- Buldowskia        
--+                                 
  |        +----- Pletholophus      
  |  +-----+                        
  |  |     |  +-- Beringiana        
  |  |     +--+                     
  +--+        +-- Sinanodonta       
     |                              
     |  +-------- Cristaria_bellua **  
     +--+                           
        |  +----- Cristaria_clessini **
        +--+                        
           |  +-- Cristaria_plicata ** 
           +--+                     
              +-- Cristaria_truncata **

Dai et al., 2024 [Publ 4385], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 27 23 6 5.0 
           +----- Amuranodonta      
  +--------+                        
  |        |  +-- Anemina           
  |        +--+                     
  |           +-- Buldowskia        
--+                                 
  |        +----- Pletholophus      
  |  +-----+                        
  |  |     |  +-- Beringiana        
  |  |     +--+                     
  +--+        +-- Sinanodonta       
     |                              
     |  +-------- Cristaria_bellua **  
     +--+                           
        |  +----- Cristaria_clessini **
        +--+                        
           |  +-- Cristaria_plicata ** 
           +--+                     
              +-- Cristaria_truncata **

Bogan et al., 2023 [Publ 4295], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 4 4 2 4 3 2 2 7.0 
           +-- Sinanodonta       
  +--------+                     
  |        +-- Pletholophus      
--+                              
  |  +-------- Cristaria_radiata ** 
  +--+                           
     |  +----- Cristaria_clessini **
     +--+                        
        |  +-- Cristaria_plicata ** 
        +--+                     
           +-- Cristaria_truncata **

Wu et al., 2025 [Publ 4442], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 14 12 7 6.0 
        +----- Cristaria_liboi **   
  +-----+                        
  |     |  +-- Cristaria_plicata ** 
  |     +--+                     
  |        +-- Cristaria_truncata **
  |                              
--+     +----- Amuranodonta      
  |  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  |  |     +-- Buldowskia        
  +--+                           
     |     +-- Beringiana        
     |  +--+                     
     |  |  +-- Pletholophus      
     +--+                        
        |  +-- Acudonta          
        +--+                     
           +-- Sinanodonta

Wang et al., 2025 [Publ 4488], fig. 3B

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 61 12 7 6.0 
        +----- Cristaria_liboi **   
  +-----+                        
  |     |  +-- Cristaria_plicata ** 
  |     +--+                     
  |        +-- Cristaria_truncata **
  |                              
--+     +----- Amuranodonta      
  |  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  |  |     +-- Buldowskia        
  +--+                           
     |     +-- Beringiana        
     |  +--+                     
     |  |  +-- Pletholophus      
     +--+                        
        |  +-- Acudonta          
        +--+                     
           +-- Sinanodonta

Wu et al., 2026 [Publ 4811], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 92 12 7 6.0 
        +----- Cristaria_liboi **   
  +-----+                        
  |     |  +-- Cristaria_plicata ** 
  |     +--+                     
  |        +-- Cristaria_truncata **
  |                              
--+     +----- Amuranodonta      
  |  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  |  |     +-- Buldowskia        
  +--+                           
     |     +-- Beringiana        
     |  +--+                     
     |  |  +-- Pletholophus      
     +--+                        
        |  +-- Acudonta          
        +--+                     
           +-- Sinanodonta

Wu et al., 2026 [Publ 4825], fig. 6B

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 62 11 7 6.0 
        +----- Cristaria_liboi **   
  +-----+                        
  |     |  +-- Cristaria_plicata ** 
  |     +--+                     
  |        +-- Cristaria_truncata **
  |                              
--+     +----- Amuranodonta      
  |  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  |  |     +-- Buldowskia        
  +--+                           
     |     +-- Beringiana        
     |  +--+                     
     |  |  +-- Pletholophus      
     +--+                        
        |  +-- Acudonta          
        +--+                     
           +-- Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 4 2 1 6.0 
  +-------- Sinanodonta       
--+                           
  |  +----- Cristaria_clessini **
  +--+                        
     |  +-- Cristaria_plicata ** 
     +--+                     
        +-- Cristaria_radiata **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 120 22 6 -- 
        +-- Cristaria_plicata ** 
  +-----+                     
  |     +-- Cristaria_clessini **
  |                           
  |  +----- Anemina           
  +--+                        
--+  |  +-- Amuranodonta      
  |  +--+                     
  |     +-- Buldowskia        
  |                           
  |  +----- Pletholophus      
  +--+                        
     |  +-- Beringiana        
     +--+                     
        +-- Sinanodonta

Froufe et al., 2017 [Publ 3802], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 10 8 3 -- 
  +-------- Anemina          
  |                          
--+     +-- Cristaria_plicata **
  |  +--+                    
  |  |  +-- Cristaria_radiata **
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 12

cstiTtTbiriSoTroTroSiD
0.600 2 2 0 2 11 5 3 -- 
  +----- Amuranodonta     
--+                       
  |  +-- Cristaria_plicata **
  +--+                    
     +-- Cristaria **

Klishko et al., 2014 [Publ 3666], fig. 8

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 2 2 2 -- 
     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Beringiana       
  +--+                    
     +-- Sinanodonta

Klishko et al., 2016 [Publ 3733], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 2 2 2 -- 
     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Beringiana

Bolotov et al., 2017 [Publ 3761], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 137 4 2 -- 
     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta

Bolotov et al., 2017 [Publ 3761], fig. S2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 137 4 2 -- 
     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta

Bolotov et al., 2017 [Publ 3764], fig. S1

cstiTtTbiriSoTroTroSiD
0.250 2 18 0 2 123 4 2 22.0 
     +-- Unio_crassus            
     |                           
     +-- Unio_pictorum           
     |                           
     +-- Unio_tumidus            
     |                           
     +-- Scriptolamprotula       
     |                           
     +-- Cuneopsis_szechenyii    
  +--+                           
  |  +-- Tchangsinaia            
  |  |                           
  |  +-- Cuneopsis_rufescens     
  |  |                           
  |  +-- Nodularia_douglasiae    
  |  |                           
  |  +-- Nodularia_nuxpersicae   
  |  |                           
  |  +-- Schistodesmus           
  |                              
  |  +-- Alasmidonta             
  |  |                           
  |  +-- Anodonta_anatina        
--+  |                           
  |  +-- Anodonta_cygnea         
  |  |                           
  |  +-- Brachyanodon            
  |  |                           
  |  +-- Lasmigona               
  |  |                           
  |  +-- Pseudanodonta           
  |  |                           
  |  +-- Pyganodon               
  |  |                           
  |  +-- Simpsonaias             
  |  |                           
  |  +-- Strophitus              
  +--+                           
     +-- Sinanodonta_lauta       
     |                           
     +-- Cristaria_plicata **       
     |                           
     +-- Cristaria_radiata **       
     |                           
     +-- Pletholophus            
     |                           
     +-- Sinanodonta_lucida      
     |                           
     +-- Sinanodonta_jourdyi     
     |                           
     +-- Lanceolaria_gladiola    
     |                           
     +-- Lanceolaria_grayii      
     |                           
     +-- Lanceolaria_danzhouensis

Bolotov et al., 2020 [Publ 4084], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 8 5 -- 
     +----- Amuranodonta     
  +--+                       
  |  |  +-- Anemina          
  |  +--+                    
--+     +-- Buldowskia       
  |                          
  |  +----- Cristaria_plicata **
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 2

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 68 10 5 -- 
  +-------------------------- Cristaria_plicata **
  |                                            
  |              +----------- Anodonta         
  |              |                             
--+  +-----------+        +-- Pyganodon        
  |  |           |  +-----+                    
  |  |           |  |     +-- Utterbackia      
  |  |           +--+                          
  |  |              |  +----- Alasmidonta      
  +--+              +--+                       
     |                 |  +-- Prolasmidonta    
     |                 +--+                    
     |                    +-- Platynaias       
     |                                         
     |  +-------------------- Lanceolaria      
     +--+                                      
        |  +----------------- Lepidodesma      
        +--+                                   
           |  +-------------- Scriptolamprotula
           +--+                                
              |  +----------- Unio             
              +--+                             
                 |  +-------- Inversiunio      
                 +--+                          
                    |  +----- Nodularia        
                    +--+                       
                       |  +-- Tchangsinaia     
                       +--+                    
                          +-- Schistodesmus

Sano et al., 2020 [Publ 4138], fig. 3

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 68 10 5 -- 
  +-------------------------- Cristaria_plicata **
  |                                            
  |              +----------- Anodonta         
  |              |                             
--+  +-----------+        +-- Pyganodon        
  |  |           |  +-----+                    
  |  |           |  |     +-- Utterbackia      
  |  |           +--+                          
  |  |              |  +----- Alasmidonta      
  +--+              +--+                       
     |                 |  +-- Platynaias       
     |                 +--+                    
     |                    +-- Prolasmidonta    
     |                                         
     |  +-------------------- Lanceolaria      
     +--+                                      
        |  +----------------- Lepidodesma      
        +--+                                   
           |  +-------------- Scriptolamprotula
           +--+                                
              |  +----------- Unio             
              +--+                             
                 |  +-------- Inversiunio      
                 +--+                          
                    |  +----- Nodularia        
                    +--+                       
                       |  +-- Schistodesmus    
                       +--+                    
                          +-- Tchangsinaia

Sano et al., 2020 [Publ 4138], fig. 4

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 68 10 5 -- 
  +-------------------------- Cristaria_plicata **
  |                                            
  |              +----------- Anodonta         
  |              |                             
--+  +-----------+        +-- Pyganodon        
  |  |           |  +-----+                    
  |  |           |  |     +-- Utterbackia      
  |  |           +--+                          
  |  |              |  +----- Alasmidonta      
  +--+              +--+                       
     |                 |  +-- Prolasmidonta    
     |                 +--+                    
     |                    +-- Platynaias       
     |                                         
     |  +-------------------- Lepidodesma      
     +--+                                      
        |  +----------------- Lanceolaria      
        +--+                                   
           |  +-------------- Scriptolamprotula
           +--+                                
              |  +----------- Unio             
              +--+                             
                 |  +-------- Nodularia        
                 +--+                          
                    |  +----- Inversiunio      
                    +--+                       
                       |  +-- Schistodesmus    
                       +--+                    
                          +-- Tchangsinaia

Sano et al., 2020 [Publ 4138], fig. 1

cstiTtTbiriSoTroTroSiD
0.286 1 22 5 1 33 9 5 36.0 
           +----- Sinohyriopsis            
     +-----+                               
     |     |  +-- Lamprotula               
     |     +--+                            
  +--+        +-- Pronodularia             
  |  |                                     
  |  |  +-------- Sinosolenaia             
  |  |  |                                  
  |  +--+  +----- Inversidens              
  |     |  |                               
  |     +--+  +-- Obovalis_omiensis__1     
  |        |  |                            
  |        +--+-- Obovalis_omiensis__2     
  |           |                            
  |           +-- Ptychorhynchus           
  |                                        
  |        +----- Lanceolaria_oxyrhyncha__1
  |        |                               
  |        +----- Lanceolaria_oxyrhyncha__2
  |        |                               
  |        +----- Cristaria_plicata__1 **     
  |        |                               
--+        +----- Cristaria_plicata__2 **     
  |        |                               
  |        +----- Sinanodonta_lauta        
  |        |                               
  |        +----- Beringiana_japonica      
  |        |                               
  |        +----- Beringiana_fukuharai__1  
  |        |                               
  |        +----- Beringiana_beringiana    
  |        |                               
  |        +----- Beringiana_fukuharai__2  
  |        |                               
  |        +----- Beringiana_fukuharai__3  
  |        |                               
  |        +----- Buldowskia_iwakawai__1   
  |        |                               
  |        +----- Buldowskia_iwakawai__2   
  |        |                               
  |        +----- Sinanodonta_tumens__1    
  +--------+                               
           +----- Sinanodonta_tumens__2    
           |                               
           +----- Pletholophus             
           |                               
           +----- Anemina                  
           |                               
           +----- Lanceolaria_lanceolata   
           |                               
           +----- Cristaria_plicata__3 **     
           |                               
           +----- Lanceolaria_gladiola     
           |                               
           +----- Sinanodonta_angula       
           |                               
           +----- Lepidodesma              
           |                               
           |  +-- Scriptolamprotula        
           +--+                            
           |  +-- Acuticosta               
           |                               
           |  +-- Cuneopsis                
           +--+                            
           |  +-- Nodularia                
           |                               
           |  +-- Schistodesmus            
           +--+                            
              +-- Inversiunio

Chen et al., 2023 [Publ 4302], fig. 1

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 40 5 5 -- 
  +-------------------- Lanceolaria_gladiola  
  |                                           
  +-------------------- Acuticosta            
  |                                           
  +-------------------- Lepidodesma           
  |                                           
  +-------------------- Lanceolaria_oxyrhyncha
  |                                           
  +-------------------- Cristaria_plicata **     
  |                                           
--+                 +-- Alasmidonta           
  |              +--+                         
  |              |  +-- Lasmigona             
  +--------------+                            
  |              |  +-- Anodonta              
  |              +--+                         
  |                 +-- Pseudanodonta         
  |                                           
  |  +----------------- Scriptolamprotula     
  |  |                                        
  |  |              +-- Middendorffinaia      
  +--+  +-----------+                         
     |  |           +-- Nodularia             
     |  |                                     
     +--+  +----------- Cuneopsis             
        |  |                                  
        |  |        +-- Diaurora              
        +--+--------+                         
           |        +-- Schistodesmus         
           |                                  
           |  +-------- Pseudocuneopsis       
           +--+                               
              |  +----- Inversiunio           
              +--+                            
                 |  +-- Tchangsinaia          
                 +--+                         
                    +-- Unio

Chen et al., 2023 [Publ 4302], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 5 5 -- 
  +----------- Cristaria_plicata **
  |                             
--+        +-- Anemina          
  |  +-----+                    
  |  |     +-- Amuranodonta     
  +--+                          
     |  +----- Pletholophus     
     +--+                       
        |  +-- Sinanodonta      
        +--+                    
           +-- Beringiana

Dai et al., 2025 [Publ 4440], fig. 1

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 43 5 5 -- 
  +----------------------------------- Cristaria_plicata **
  |                                                     
  |                                +-- Alasmidonta      
  |                             +--+                    
  |                             |  +-- Lasmigona        
  +-----------------------------+                       
--+                             |  +-- Anodonta         
  |                             +--+                    
  |                                +-- Pseudanodonta    
  |                                                     
  |  +-------------------------------- Lanceolaria      
  |  |                                                  
  +--+  +----------------------------- Lepidodesma      
     |  |                                               
     +--+  +-------------------------- Acuticosta       
        |  |                                            
        +--+  +----------------------- Globunio         
           |  |                                         
           +--+  +-------------------- Scriptolamprotula
              |  |                                      
              +--+  +----------------- Middendorffinaia 
                 |  |                                   
                 +--+  +-------------- Nodularia        
                    |  |                                
                    +--+  +----------- Cuneopsis        
                       |  |                             
                       |  |        +-- Diaurora         
                       +--+--------+                    
                          |        +-- Schistodesmus    
                          |                             
                          |  +-------- Pseudocuneopsis  
                          +--+                          
                             |  +----- Unio             
                             +--+                       
                                |  +-- Tchangsinaia     
                                +--+                    
                                   +-- Inversiunio

Dai et al., 2025 [Publ 4440], fig. 2

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 43 5 5 -- 
  +-------------------------- Lanceolaria_gladiola  
  |                                                 
  +-------------------------- Lanceolaria_oxyrhyncha
  |                                                 
--+-------------------------- Cristaria_plicata **     
  |                                                 
  |  +----------------------- Acuticosta            
  |  |                                              
  |  +----------------------- Lepidodesma           
  +--+                                              
     |  +-------------------- Globunio              
     |  |                                           
     +--+  +----------------- Scriptolamprotula     
        |  |                                        
        |  |              +-- Middendorffinaia      
        +--+  +-----------+                         
           |  |           +-- Nodularia             
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              |  |        +-- Diaurora              
              +--+--------+                         
                 |        +-- Schistodesmus         
                 |                                  
                 |  +-------- Pseudocuneopsis       
                 +--+                               
                    |  +----- Inversiunio           
                    +--+                            
                       |  +-- Tchangsinaia          
                       +--+                         
                          +-- Unio

Chen et al., 2025 [Publ 4487], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 40 5 5 -- 
  +----------- Cristaria_plicata **
  |                             
--+        +-- Anemina          
  |  +-----+                    
  |  |     +-- Amuranodonta     
  +--+                          
     |  +----- Pletholophus     
     +--+                       
        |  +-- Sinanodonta      
        +--+                    
           +-- Beringiana

Chen et al., 2025 [Publ 4487], fig. 3

cstiTtTbiriSoTroTroSiD
0.857 1 1 0 1 40 5 5 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Lanceolaria

Sano et al., 2017 [Publ 3799], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 20 7 4 -- 
        +-- Cristaria_plicata **
  +-----+                    
  |     +-- Buldowskia       
--+                          
  |  +----- Pletholophus     
  +--+                       
     |  +-- Sinanodonta      
     +--+                    
        +-- Beringiana

Sano et al., 2017 [Publ 3799], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 32 8 4 -- 
        +-- Cristaria_plicata **
  +-----+                    
  |     +-- Buldowskia       
--+                          
  |  +----- Beringiana       
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta

Sano et al., 2022 [Publ 4311], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 11 11 4 -- 
        +-- Beringiana       
  +-----+                    
  |     +-- Sinanodonta      
--+                          
  |  +----- Pletholophus     
  +--+                       
     |  +-- Cristaria_plicata **
     +--+                    
        +-- Buldowskia

Sano et al., 2022 [Publ 4311], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 17 17 4 -- 
        +-- Beringiana       
  +-----+                    
  |     +-- Sinanodonta      
--+                          
  |  +----- Pletholophus     
  +--+                       
     |  +-- Cristaria_plicata **
     +--+                    
        +-- Buldowskia

Xiang et al., 2025 [Publ 4433], fig. 7

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 12 6 4 -- 
  +----------- Cristaria_plicata **
--+                             
  |  +-------- Anemina          
  +--+                          
     |  +----- Beringiana       
     +--+                       
        |  +-- Mimo             
        +--+                    
           +-- Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 10

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 11 5 4 -- 
  +----------- Anemina          
--+                             
  |  +-------- Amuranodonta     
  +--+                          
     |  +----- Cristaria_plicata **
     +--+                       
        |  +-- Beringiana       
        +--+                    
           +-- Sinanodonta

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 80 4 3 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 200 8 3 -- 
  +-------- Anemina          
--+                          
  |  +----- Cristaria_plicata **
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 198 8 3 -- 
  +-------- Anemina          
--+                          
  |  +----- Cristaria_plicata **
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 200 8 3 -- 
  +-------- Acuticosta           
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea      
  |                              
  +-------- Cristaria_plicata **    
  |                              
  +-------- Brachyanodon         
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesma          
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unio                 
  |                              
  +-------- Scriptolamprotula    
  |                              
  +-------- Lanceolaria          
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 5 3 -- 
     +-- Anemina          
  +--+                    
  |  +-- Cristaria_plicata **
--+                       
  |  +-- Beringiana       
  +--+                    
     +-- Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 5 3 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 5 3 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 5 3 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Wu et al., 2024 [Publ 4379], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 5 3 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Wu et al., 2024 [Publ 4388], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 3 3 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Hou et al., 2024 [Publ 4400], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 74 5 3 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Hou et al., 2025 [Publ 4445], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 51 5 3 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 11

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 13 3 3 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Amuranodonta     
  +--+                       
     |  +-- Sinanodonta      
     +--+                    
        +-- Beringiana

Xue et al., 2016 [Publ 3659], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 14 3 2 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Platynaias

Wu et al., 2016 [Publ 3711], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 20 2 2 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anodonta         
  +--+                       
     |  +-- Pyganodon        
     +--+                    
        +-- Utterbackia

Fonseca et al., 2016 [Publ 3775], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 31 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Bolotov et al., 2018 [Publ 3826], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 81 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta

Huang et al., 2018 [Publ 3828], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 41 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2018 [Publ 3836], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 5 2 -- 
  +-- Anemina          
  |                    
--+-- Cristaria_plicata **
  |                    
  +-- Sinanodonta

Wang et al., 2018 [Publ 3846], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Aunins et al., 2018 [Publ 3863], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 39 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wang et al., 2018 [Publ 3865], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Cha et al., 2018 [Publ 3916], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 37 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2018 [Publ 4143], fig. 2

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 21 2 2 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anodonta         
  +--+                       
     |  +-- Pyganodon        
     +--+                    
        +-- Utterbackia

Huang et al., 2019 [Publ 3843], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 45 7 2 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Lanceolaria

Huang et al., 2019 [Publ 3843], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 55 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 46 7 2 -- 
  +-- Lepidodesma      
--+                    
  +-- Cristaria_plicata **

Wu et al., 2019 [Publ 3864], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 45 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Froufe et al., 2019 [Publ 3936], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 43 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Froufe et al., 2019 [Publ 3936], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 43 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 2 2 -- 
  +----- Cristaria_radiata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Pletholophus

Riccardi et al., 2020 [Publ 4089], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 18 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 53 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

cstiTtTbiriSoTroTroSiD
0.500 1 1 0 1 11 4 2 -- 
  +-------------- Cristaria_plicata **       
  |                                       
  |  +----------- Sinanodonta_tumens      
  |  |                                    
  |  |        +-- Sinanodonta_lauta__5    
  |  |        |                           
--+  |        +-- Sinanodonta_lauta__6    
  |  |        |                           
  |  |        +-- Beringiana_japonica__15 
  |  |        |                           
  |  |        +-- Beringiana_japonica__16 
  |  |  +-----+                           
  |  |  |     +-- Beringiana_japonica__17 
  +--+  |     |                           
     |  |     +-- Beringiana_japonica__18 
     |  |     |                           
     |  |     +-- Beringiana_japonica__19 
     |  |     |                           
     |  |     +-- Beringiana_japonica__20 
     |  |                                 
     |  |  +----- Beringiana_japonica__1  
     |  |  |                              
     |  |  +----- Beringiana_japonica__2  
     |  |  |                              
     +--+  +----- Beringiana_japonica__3  
        |  |                              
        |  +----- Beringiana_japonica__4  
        |  |                              
        |  +----- Beringiana_japonica__5  
        |  |                              
        |  +----- Sinanodonta_lauta__1    
        |  |                              
        |  +----- Beringiana_japonica__6  
        |  |                              
        |  +----- Sinanodonta_lauta__2    
        |  |                              
        |  +----- Sinanodonta_calipygos__1
        |  |                              
        +--+----- Beringiana_japonica__7  
           |                              
           +----- Sinanodonta_lauta__4    
           |                              
           +----- Beringiana_japonica__8  
           |                              
           +----- Beringiana_japonica__9  
           |                              
           +----- Beringiana_japonica__10 
           |                              
           +----- Beringiana_japonica__11 
           |                              
           +----- Beringiana_japonica__12 
           |                              
           +----- Beringiana_japonica__13 
           |                              
           +----- Beringiana_japonica__14 
           |                              
           |  +-- Sinanodonta_lauta__3    
           +--+                           
              +-- Sinanodonta_calipygos__2

Wu et al., 2020 [Publ 4121], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2020 [Publ 4147], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 29 2 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Anemina

Chen et al., 2020 [Publ 4170], fig. 1

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 33 2 2 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Anodonta

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 2_I

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 3 2 -- 
  +----- Anemina          
--+                       
  |  +-- Cristaria_plicata **
  +--+                    
     +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 67 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 3 2 -- 
  +----- Anemina          
--+                       
  |  +-- Cristaria_plicata **
  +--+                    
     +-- Sinanodonta

Wu et al., 2022 [Publ 4238], fig. 2d

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 37 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Wu et al., 2022 [Publ 4238], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 72 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 75 4 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 75 4 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 36 2 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Anemina

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 38 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Dai et al., 2023 [Publ 4285], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 2 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Bogan et al., 2023 [Publ 4295], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 4 2 -- 
  +----- Cristaria_truncata **
--+                        
  |  +-- Sinanodonta       
  +--+                     
     +-- Pletholophus

Bogan et al., 2023 [Publ 4295], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 12 11 2 -- 
  +----- Pletholophus      
--+                        
  |  +-- Cristaria_truncata **
  +--+                     
     +-- Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 18 3 2 -- 
  +----- Anemina          
--+                       
  |  +-- Cristaria_plicata **
  +--+                    
     +-- Sinanodonta

Dai et al., 2024 [Publ 4358], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 2 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Dai et al., 2024 [Publ 4389], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 45 2 2 -- 
  +----- Sinanodonta      
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Cristaria_plicata **

Dai et al., 2025 [Publ 4440], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 86 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Dai et al., 2025 [Publ 4441], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 46 2 2 -- 
  +----- Sinanodonta      
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Cristaria_plicata **

Chen et al., 2025 [Publ 4448], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 84 4 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Chen et al., 2025 [Publ 4663], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 2 2 -- 
  +----- Sinanodonta      
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Cristaria_plicata **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 139 3 2 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta

Huang et al., 2002 [Publ 2657], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 1 1 -- 
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Huang et al., 2002 [Publ 2657], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 1 1 -- 
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Zhang et al., 2004 [Publ 2735], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 13 1 1 -- 
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Zhou et al., 2007 [Publ 2658], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 20 1 1 -- 
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Zhou et al., 2007 [Publ 2658], fig. 2

cstiTtTbiriSoTroTroSiD
0.667 1 1 0 1 17 1 1 -- 
  +----------------------- Cristaria_plicata **   
  |                                            
  |              +-------- Scriptolamprotula   
--+  +-----------+                             
  |  |           |  +----- Acuticosta          
  |  |           +--+                          
  |  |              |  +-- Nodularia           
  +--+              +--+                       
     |                 +-- Cuneopsis           
     |                                         
     |  +----------------- Lamprotula          
     +--+                                      
        |  +-------------- Theliderma          
        +--+                                   
           |  +----------- Quadrula_quadrula   
           +--+                                
              |  +-------- Pustulosa_infucata  
              +--+                             
                 |  +----- Quadrula_verrucosa  
                 +--+                          
                    |  +-- Pustulosa_kieneriana
                    +--+                       
                       +-- Pustulosa_succissa

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 1 1 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Huang et al., 2013 [Publ 3629], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 15 1 1 -- 
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Zhang et al., 2013 [Publ 4214], fig. 5

cstiTtTbiriSoTroTroSiD
0.667 1 1 0 1 3 1 1 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Sinanodonta      
  +--+                       
     |  +-- Margaritifera    
     +--+                    
        +-- Lamprotula

Zhou et al., 2016 [Publ 3710], fig. 1

cstiTtTbiriSoTroTroSiD
0.667 1 1 0 1 19 1 1 -- 
  +-------- Cristaria_plicata **
--+                          
  |  +----- Platynaias       
  +--+                       
     |  +-- Pyganodon        
     +--+                    
        +-- Utterbackia

Xia et al., 2016 [Publ 3712], fig. 7B

cstiTtTbiriSoTroTroSiD
0.667 1 1 0 1 2 1 1 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Sinohyriopsis

Guerra et al., 2017 [Publ 3772], fig. 2a

cstiTtTbiriSoTroTroSiD
0.667 1 1 0 1 24 1 1 -- 
  +----------- Cristaria_plicata **
--+                             
  |  +-------- Anodonta         
  +--+                          
     |  +----- Platynaias       
     +--+                       
        |  +-- Pyganodon        
        +--+                    
           +-- Utterbackia

Guerra et al., 2017 [Publ 3772], fig. 2b

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 24 1 1 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Wen et al., 2017 [Publ 3795], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 15 1 1 -- 
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 1 1 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 1 1 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 11 1 1 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Wu et al., 2024 [Publ 4379], fig. 4_6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 2 1 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta

Jiang et al., 2010 [Publ 3438], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 3 0 0 -- 
  +----- Lampsilis        
  |                       
--+----- Cristaria_plicata **
  |                       
  |  +-- Pronodularia     
  +--+                    
     +-- Sinohyriopsis

Breton et al., 2011 [Publ 3376], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 37 0 0 -- 
  +-------------- Cristaria_plicata **
  |                                
--+           +-- Pterosyna        
  |  +--------+                    
  |  |        +-- Lasmigona        
  +--+                             
     |  +-------- Strophitus       
     |  |                          
     +--+     +-- Platynaias       
        |  +--+                    
        |  |  +-- Alasmidonta      
        +--+                       
           |  +-- Pyganodon        
           +--+                    
              +-- Utterbackia

Plazzi et al., 2011 [Publ 3676], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 17 0 0 -- 
  +----------------- Cristaria_plicata **
  |                                   
  |  +-------------- Lanceolaria      
  |  |                                
--+  +-------------- Anodonta         
  |  |                                
  |  +-------------- Unio             
  |  |                                
  |  |           +-- Pyganodon        
  +--+-----------+                    
     |           +-- Pseudanodonta    
     |                                
     |           +-- Alathyria        
     |           |                    
     |  +--------+-- Pronodularia     
     |  |        |                    
     |  |        +-- Sinohyriopsis    
     +--+                             
        |  +-------- Quadrula         
        |  |                          
        +--+-------- Parvaspina       
           |                          
           |  +----- Lampsilis        
           +--+                       
              |  +-- Epioblasma       
              +--+                    
                 +-- Venustaconcha

Chen et al., 2012 [Publ 3624], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 6 0 0 -- 
  +-- Pyganodon        
--+                    
  +-- Cristaria_plicata **

Peng et al., 2012 [Publ 3689], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 2 0 0 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Sinohyriopsis

Sheng et al., 2014 [Publ 3704], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 7 0 0 -- 
  +-------------- Cristaria_plicata **
  |                                
--+  +----------- Pyganodon        
  |  |                             
  +--+  +-------- Lampsilis        
     |  |                          
     +--+     +-- Venustaconcha    
        |  +--+                    
        |  |  +-- Quadrula         
        +--+                       
           |  +-- Lamprotula       
           +--+                    
              +-- Sinohyriopsis

Wei et al., 2016 [Publ 3665], fig. 0

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 5 0 0 -- 
  +-- Cristaria_plicata **
  |                    
  +-- Lampsilis        
--+                    
  +-- Pyganodon        
  |                    
  +-- Quadrula

Froufe et al., 2017 [Publ 3802], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 3 0 0 -- 
  +-- Cristaria_plicata **
--+                    
  +-- Anodonta

Smith et al., 2018 [Publ 3790], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 17 0 0 -- 
  +-------------- Cristaria_plicata **
  |                                
  |           +-- Utterbackia      
--+  +--------+                    
  |  |        +-- Pyganodon        
  |  |                             
  +--+        +-- Anodontoides     
     |  +-----+                    
     |  |     +-- Strophitus       
     |  |                          
     +--+     +-- Alasmidonta      
        |  +--+                    
        |  |  +-- Alasminota       
        +--+                       
           |  +-- Pseudodontoideus 
           +--+                    
              +-- Utterbackiana

Smith, 2021 [Publ 4331], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 0 0 -- 
  +----- Cristaria_plicata **
--+                       
  |  +-- Pyganodon        
  +--+                    
     +-- Utterbackia