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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Cristaria Schumacher, 1817: cladomics
     6 Recent valid species (9 total species)

Cladomics Summary

number of species = 6 (9 total)
analyzed in 115 cladograms in 76 publications
maximum number of terminals in an analysis = 6 (mean = 1.3, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 16
maximum number of included terminals = 6, (mean = 3.19 median = 3)
proportion of cladograms that support basic monophyly = 0.94


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Wu et al., 2025 [Publ 4442], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
6646292770
                 +-- Cristaria_clessini **         
           +-----+                              
           |     +-- Cristaria_bellua **           
           |                                    
  +--------+     +-- Cristaria_plicata **          
  |        |  +--+                              
  |        |  |  +-- Cristaria_truncata **         
  |        +--+                                 
  |           |  +-- Cristaria_radiata **          
  |           +--+                              
--+              +-- Cristaria_beirensis **        
  |                                             
  |           +----- Beringiana                 
  |  +--------+                                 
  |  |        |  +-- Acudonta                   
  |  |        +--+                              
  |  |           +-- Sinanodonta                
  +--+                                          
     |        +----- Pletholophus_guangzhouensis
     |  +-----+                                 
     |  |     |  +-- Pletholophus_tenuis        
     |  |     +--+                              
     +--+        +-- Pletholophus_reinianus     
        |                                       
        |  +-------- Pletholophus_honglinhensis 
        +--+                                    
           |  +----- Anemina                    
           +--+                                 
              |  +-- Amuranodonta               
              +--+                              
                 +-- Buldowskia                 

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
553551853
           +----- Anemina           
  +--------+                        
  |        |  +-- Amuranodonta      
  |        +--+                     
  |           +-- Buldowskia        
--+                                 
  |           +-- Beringiana        
  |  +--------+                     
  |  |        +-- Sinanodonta       
  +--+                              
     |  +-------- Cristaria_radiata ** 
     |  |                           
     +--+     +-- Cristaria_truncata **
        |  +--+                     
        |  |  +-- Cristaria_plicata ** 
        +--+                        
           |  +-- Cristaria_bellua **  
           +--+                     
              +-- Cristaria_clessini **

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
4424272364
           +----- Amuranodonta      
  +--------+                        
  |        |  +-- Anemina           
  |        +--+                     
  |           +-- Buldowskia        
--+                                 
  |        +----- Pletholophus      
  |  +-----+                        
  |  |     |  +-- Beringiana        
  |  |     +--+                     
  +--+        +-- Sinanodonta       
     |                              
     |  +-------- Cristaria_bellua **  
     +--+                           
        |  +----- Cristaria_clessini **
        +--+                        
           |  +-- Cristaria_plicata ** 
           +--+                     
              +-- Cristaria_truncata **

Xiang et al., 2025 [Publ 4433], fig. 6

iTtTbiriSoTroTroSiD
4424242374
  +----------------- Pletholophus      
  |                                    
  |           +----- Beringiana        
--+  +--------+                        
  |  |        |  +-- Mimo              
  |  |        +--+                     
  |  |           +-- Sinanodonta       
  +--+                                 
     |        +----- Anemina           
     |  +-----+                        
     |  |     |  +-- Amuranodonta      
     |  |     +--+                     
     +--+        +-- Buldowskia        
        |                              
        |  +-------- Cristaria_bellua **  
        +--+                           
           |  +----- Cristaria_clessini **
           +--+                        
              |  +-- Cristaria_plicata ** 
              +--+                     
                 +-- Cristaria_truncata **

Bogan et al., 2023 [Publ 4295], fig. 4

iTtTbiriSoTroTroSiD
44243224
           +-- Sinanodonta       
  +--------+                     
  |        +-- Pletholophus      
--+                              
  |  +-------- Cristaria_radiata ** 
  +--+                           
     |  +----- Cristaria_clessini **
     +--+                        
        |  +-- Cristaria_plicata ** 
        +--+                     
           +-- Cristaria_truncata **

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
4424272364
           +----- Amuranodonta      
  +--------+                        
  |        |  +-- Anemina           
  |        +--+                     
  |           +-- Buldowskia        
--+                                 
  |        +----- Pletholophus      
  |  +-----+                        
  |  |     |  +-- Beringiana        
  |  |     +--+                     
  +--+        +-- Sinanodonta       
     |                              
     |  +-------- Cristaria_bellua **  
     +--+                           
        |  +----- Cristaria_clessini **
        +--+                        
           |  +-- Cristaria_plicata ** 
           +--+                     
              +-- Cristaria_truncata **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
4424272364
     +-------- Cristaria_bellua **  
  +--+                           
  |  |  +----- Cristaria_clessini **
  |  +--+                        
  |     |  +-- Cristaria_plicata ** 
  |     +--+                     
--+        +-- Cristaria_truncata **
  |                              
  |     +----- Amuranodonta      
  |  +--+                        
  |  |  |  +-- Anemina           
  |  |  +--+                     
  +--+     +-- Buldowskia        
     |                           
     |  +----- Pletholophus      
     +--+                        
        |  +-- Beringiana        
        +--+                     
           +-- Sinanodonta       

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

iTtTbiriSoTroTroSiD
33134213
  +-------- Sinanodonta       
--+                           
  |  +----- Cristaria_clessini **
  +--+                        
     |  +-- Cristaria_plicata ** 
     +--+                     
        +-- Cristaria_radiata ** 

Wu et al., 2025 [Publ 4442], fig. 6

iTtTbiriSoTroTroSiD
3313141273
        +----- Cristaria_beirensis **
  +-----+                         
  |     |  +-- Cristaria_plicata **  
  |     +--+                      
  |        +-- Cristaria_truncata ** 
  |                               
--+     +----- Amuranodonta       
  |  +--+                         
  |  |  |  +-- Anemina            
  |  |  +--+                      
  |  |     +-- Buldowskia         
  +--+                            
     |     +-- Beringiana         
     |  +--+                      
     |  |  +-- Pletholophus       
     +--+                         
        |  +-- Acudonta           
        +--+                      
           +-- Sinanodonta        

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
220213742--
     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta      

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
2202119226--
        +-- Cristaria_plicata ** 
  +-----+                     
  |     +-- Cristaria_clessini **
  |                           
  |  +----- Anemina           
  +--+                        
--+  |  +-- Amuranodonta      
  |  +--+                     
  |     +-- Buldowskia        
  |                           
  |  +----- Pletholophus      
  +--+                        
     |  +-- Beringiana        
     +--+                     
        +-- Sinanodonta       

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
220213742--
     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta      

Klishko et al., 2014 [Publ 3666], fig. 8

iTtTbiriSoTroTroSiD
2202222--
     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Beringiana       
  +--+                    
     +-- Sinanodonta      

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
22021083--
  +-------- Anemina          
  |                          
--+     +-- Cristaria_plicata **
  |  +--+                    
  |  |  +-- Cristaria_radiata **
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta      

Klishko et al., 2016 [Publ 3733], fig. 1

iTtTbiriSoTroTroSiD
2202222--
     +-- Cristaria_plicata **
  +--+                    
  |  +-- Cristaria_radiata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Beringiana       

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
217021234220
  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina     
  |                        
  +-- Anodonta_cygnea      
  |                        
  +-- Anodonta_nuttalliana 
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta    
  |                        
  +-- Cristaria_plicata **    
  |                        
  +-- Cristaria_radiata **    
  |                        
  +-- Pletholophus         
  |                        
  +-- Sinanodonta_lucida   
  |                        
  +-- Sinanodonta_jourdyi  
  |                        
  +-- Lanceolaria_gladiola 
  |                        
  +-- Lanceolaria_grayii__1
  |                        
  +-- Lanceolaria_grayii__2

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

iTtTbiriSoTroTroSiD
1101532--
  +----- Cristaria_plicata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Pletholophus     

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015222--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119883--
-- Cristaria_plicata **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014832--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
11014332--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
11014472--
-- Cristaria_plicata **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
11011511--
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119883--
  +-------- Anemina          
--+                          
  |  +----- Cristaria_plicata **
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta      

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
110167105--
-- Cristaria_plicata **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11013955--
-- Cristaria_plicata **

Wei et al., 2016 [Publ 3665], fig. 0

iTtTbiriSoTroTroSiD
1101500--
-- Cristaria_plicata **

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1101600--
-- Cristaria_plicata **

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
11011832--
  +----- Anemina          
--+                       
  |  +-- Cristaria_plicata **
  +--+                    
     +-- Sinanodonta      

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
11013322--
-- Cristaria_plicata **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
11013132--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019453--
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta      

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014532--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012411--
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta      

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
11013732--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
11014272--
-- Cristaria_plicata **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017232--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iTtTbiriSoTroTroSiD
1101522--
  +----- Cristaria_radiata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
11012121--
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta      

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
11012022--
-- Cristaria_plicata **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
12203339537
  +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1 **   
  |                             
  +----- Cristaria_plicata__2 **   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3 **   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio            

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
11012122--
-- Cristaria_plicata **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
11013011--
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta      

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119683--
  +-------- Anemina          
--+                          
  |  +----- Cristaria_plicata **
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta      

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015611--
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta      

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013311--
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
11015532--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
11012822--
  +----- Cristaria_plicata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Anemina          

Bogan et al., 2023 [Publ 4295], fig. 5

iTtTbiriSoTroTroSiD
1101542--
  +----- Cristaria_truncata **
--+                        
  |  +-- Sinanodonta       
  +--+                     
     +-- Pletholophus      

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
11014622--
  +----- Sinanodonta      
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Cristaria_plicata **

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
11013284--
        +-- Cristaria_plicata **
  +-----+                    
  |     +-- Buldowskia       
--+                          
  |  +----- Beringiana       
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta      

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
11012011--
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
11013032--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015333--
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta      

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017232--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017943--
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Pletholophus     
     +--+                    
        +-- Sinanodonta      

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
11011111--
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta      

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
11011700--
-- Cristaria_plicata **

Jiang et al., 2010 [Publ 3438], fig. 5

iTtTbiriSoTroTroSiD
1101300--
-- Cristaria_plicata **

Sano et al., 2022 [Publ 4311], fig. 3

iTtTbiriSoTroTroSiD
110117174--
        +-- Beringiana       
  +-----+                    
  |     +-- Sinanodonta      
--+                          
  |  +----- Pletholophus     
  +--+                       
     |  +-- Cristaria_plicata **
     +--+                    
        +-- Buldowskia       

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
11013032--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015611--
  +-- Cristaria_plicata **
--+                    
  +-- Sinanodonta      

Smith et al., 2018 [Publ 3790], fig. 1

iTtTbiriSoTroTroSiD
11011700--
-- Cristaria_plicata **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
11013032--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
11013732--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
11014455--
-- Cristaria_plicata **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
11013522--
  +----- Cristaria_plicata **
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Anemina          

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
1101311--
-- Cristaria_plicata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017542--
  +----- Cristaria_plicata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta      

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
11014722--
  +----- Sinanodonta      
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Cristaria_plicata **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
11014732--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
11011832--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Bogan et al., 2023 [Publ 4295], fig. 6

iTtTbiriSoTroTroSiD
110112112--
  +----- Pletholophus      
--+                        
  |  +-- Cristaria_truncata **
  +--+                     
     +-- Sinanodonta       

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
11011311--
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012411--
-- Cristaria_plicata **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
11011432--
-- Cristaria_plicata **

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1101700--
-- Cristaria_plicata **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
11015252--
  +-- Anemina          
  |                    
--+-- Cristaria_plicata **
  |                    
  +-- Sinanodonta      

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
11014332--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11013955--
  +----------- Cristaria_plicata **
  |                             
--+        +-- Anemina          
  |  +-----+                    
  |  |     +-- Amuranodonta     
  +--+                          
     |  +----- Pletholophus     
     +--+                       
        |  +-- Sinanodonta      
        +--+                    
           +-- Beringiana       

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
11012232--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
11013732--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019453--
     +-- Anemina          
  +--+                    
  |  +-- Cristaria_plicata **
--+                       
  |  +-- Beringiana       
  +--+                    
     +-- Sinanodonta      

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013311--
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
11017553--
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta      

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
110167105--
-- Cristaria_plicata **

Froufe et al., 2017 [Publ 3802], fig. 2

iTtTbiriSoTroTroSiD
1101300--
-- Cristaria_plicata **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019032--
  +----- Anemina          
--+                       
  |  +-- Cristaria_plicata **
  +--+                    
     +-- Sinanodonta      

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
11012074--
        +-- Cristaria_plicata **
  +-----+                    
  |     +-- Buldowskia       
--+                          
  |  +----- Pletholophus     
  +--+                       
     |  +-- Sinanodonta      
     +--+                    
        +-- Beringiana       

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018732--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
11011264--
  +----------- Cristaria_plicata **
--+                             
  |  +-------- Anemina          
  +--+                          
     |  +----- Beringiana       
     +--+                       
        |  +-- Mimo             
        +--+                    
           +-- Sinanodonta      

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
11011142--
  +-------------- Cristaria_plicata **       
  |                                       
  |  +----------- Sinanodonta_tumens      
  |  |                                    
  |  |        +-- Sinanodonta_lauta__5    
  |  |        |                           
--+  |        +-- Sinanodonta_lauta__6    
  |  |        |                           
  |  |        +-- Beringiana_japonica__15 
  |  |        |                           
  |  |        +-- Beringiana_japonica__16 
  |  |  +-----+                           
  |  |  |     +-- Beringiana_japonica__17 
  +--+  |     |                           
     |  |     +-- Beringiana_japonica__18 
     |  |     |                           
     |  |     +-- Beringiana_japonica__19 
     |  |     |                           
     |  |     +-- Beringiana_japonica__20 
     |  |                                 
     |  |  +----- Beringiana_japonica__1  
     |  |  |                              
     |  |  +----- Beringiana_japonica__2  
     |  |  |                              
     +--+  +----- Beringiana_japonica__3  
        |  |                              
        |  +----- Beringiana_japonica__4  
        |  |                              
        |  +----- Beringiana_japonica__5  
        |  |                              
        |  +----- Sinanodonta_lauta__1    
        |  |                              
        |  +----- Beringiana_japonica__6  
        |  |                              
        |  +----- Sinanodonta_lauta__2    
        |  |                              
        |  +----- Sinanodonta_calipygos__1
        |  |                              
        +--+----- Beringiana_japonica__7  
           |                              
           +----- Sinanodonta_lauta__4    
           |                              
           +----- Beringiana_japonica__8  
           |                              
           +----- Beringiana_japonica__9  
           |                              
           +----- Beringiana_japonica__10 
           |                              
           +----- Beringiana_japonica__11 
           |                              
           +----- Beringiana_japonica__12 
           |                              
           +----- Beringiana_japonica__13 
           |                              
           +----- Beringiana_japonica__14 
           |                              
           |  +-- Sinanodonta_lauta__3    
           +--+                           
              +-- Sinanodonta_calipygos__2

Xia et al., 2016 [Publ 3712], fig. 7B

iTtTbiriSoTroTroSiD
1101211--
-- Cristaria_plicata **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019032--
  +----- Anemina          
--+                       
  |  +-- Cristaria_plicata **
  +--+                    
     +-- Sinanodonta      

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
11016732--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
11015253--
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta      

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
11011911--
-- Cristaria_plicata **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
110167105--
-- Cristaria_plicata **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011900--
-- Cristaria_plicata **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
11013600--
-- Cristaria_plicata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017542--
  +----- Cristaria_plicata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta      

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
11015232--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013585--
     +----- Amuranodonta     
  +--+                       
  |  |  +-- Anemina          
  |  +--+                    
--+     +-- Buldowskia       
  |                          
  |  +----- Cristaria_plicata **
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta      

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019053--
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta      

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
11011511--
  +-- Sinanodonta      
--+                    
  +-- Cristaria_plicata **

Sano et al., 2022 [Publ 4311], fig. 2

iTtTbiriSoTroTroSiD
110111114--
        +-- Beringiana       
  +-----+                    
  |     +-- Sinanodonta      
--+                          
  |  +----- Pletholophus     
  +--+                       
     |  +-- Cristaria_plicata **
     +--+                    
        +-- Buldowskia       

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
11014132--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
11011711--
-- Cristaria_plicata **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
11014455--
-- Cristaria_plicata **

Peng et al., 2012 [Publ 3689], fig. 1

iTtTbiriSoTroTroSiD
1101200--
-- Cristaria_plicata **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015122--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019053--
  +-------- Cristaria_plicata **
--+                          
  |  +----- Anemina          
  +--+                       
     |  +-- Beringiana       
     +--+                    
        +-- Sinanodonta      

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017232--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11018232--
  +----- Cristaria_plicata **
--+                       
  |  +-- Pletholophus     
  +--+                    
     +-- Sinanodonta      

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
11013932--
  +----- Cristaria_plicata **
--+                       
  |  +-- Anemina          
  +--+                    
     +-- Sinanodonta      
 
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