Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Cristaria Schumacher, 1817: cladomics

8 Recent valid species (11 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 8 (11 total)
analyzed in 131 cladograms in 88 publications
maximum number of terminals in an analysis = 7 (mean = 1.49, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 24
maximum number of included terminals = 7, (mean = 3.67 median = 3)
proportion of cladograms that support basic monophyly = 0.96

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Cristaria cladogram distances
PCoA Ordination

NJ Tree of Cristaria cladogram distances
NJ Tree

Cladograms

Dai et al., 2026 [Publ 4757], fig. 3

** Top Monophyly Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 7 7 6 7 27 23 6 **
Cladomics cladogram Amuranodonta Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria wangyucheni Cristaria bellua Cristaria clessini Cristaria plicata Cristaria truncata Cristaria radiata 2 Cristaria radiata 3 Cristaria liboi Cristaria radiata 1

Chen et al., 2026 [Publ 4805], fig. 4

** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 28 27 8 5.0
Cladomics cladogram Cristaria clessini Cristaria bellua Cristaria plicata Cristaria truncata Cristaria radiata Cristaria liboi Beringiana Mimo Acudonta Sinanodonta Pletholophus Anemina Amuranodonta Buldowskia

Tumpeesuwan et al., 2026 [Publ 4845], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 7 3 3 3 8.0
Cladomics cladogram Beringiana Sinanodonta Buldowskia Cristaria liboi Cristaria radiata Cristaria plicata Cristaria truncata Cristaria clessini Cristaria hospes Cristaria bellua

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

cst iT tT bi riS oT roT roS iD
0.900 6 6 4 6 627 27 8 7.0
Cladomics cladogram Cristaria liboi Cristaria radiata Cristaria bellua Cristaria clessini Cristaria plicata Cristaria truncata Lepidodesma Lanceolaria Pseudoacuticosta retiaria 2 Acuticosta chinensis 2 Acuticosta chinensis 1 Pseudoacuticosta retiaria 1 Globunio Scriptolamprotula Middendorffinaia Unio Schistodesmus Nodularia Tchangsinaia Inversiunio Pseudocuneopsis Diaurora Cuneopsis Pseudodontoideus Lasmigona Pterosyna Arcidens Utterbackia Pyganodon Utterbackiana Simpsonaias Alasminota Alasmidonta Prolasmidonta Platynaias Anodontoides Strophitus Anodonta anatina Pseudanodonta Brachyanodon oregonensis 2 Anodonta cygnea Anodonta seddonae Anodonta exulcerata Brachyanodon impura Brachyanodon Brachyanodon nuttalliana 1 Brachyanodon nuttalliana 2 Brachyanodon californiensis Brachyanodon oregonensis 1

Wu et al., 2025 [Publ 4442], fig. 2

cst iT tT bi riS oT roT roS iD
0.889 6 6 4 6 29 27 7 5.0
Cladomics cladogram Cristaria clessini Cristaria bellua Cristaria plicata Cristaria truncata Cristaria radiata Cristaria liboi Beringiana Acudonta Sinanodonta Pletholophus guangzhouensis Pletholophus tenuis Pletholophus reinianus Pletholophus honglinhensis Anemina Amuranodonta Buldowskia

Konopleva et al., 2023 [Publ 4269], fig. S1_4

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 51 8 5 8.0
Cladomics cladogram Anemina Amuranodonta Buldowskia Beringiana Sinanodonta Cristaria radiata Cristaria truncata Cristaria plicata Cristaria bellua Cristaria clessini

Xiang et al., 2025 [Publ 4433], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 24 23 7 5.0
Cladomics cladogram Pletholophus Beringiana Mimo Sinanodonta Anemina Amuranodonta Buldowskia Cristaria bellua Cristaria clessini Cristaria plicata Cristaria truncata

Bogan et al., 2023 [Publ 4295], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 27 23 6 5.0
Cladomics cladogram Cristaria bellua Cristaria clessini Cristaria plicata Cristaria truncata Amuranodonta Anemina Buldowskia Pletholophus Beringiana Sinanodonta

Dai et al., 2024 [Publ 4385], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 27 23 6 5.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria bellua Cristaria clessini Cristaria plicata Cristaria truncata

Dai et al., 2024 [Publ 4385], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 27 23 6 5.0
Cladomics cladogram Amuranodonta Anemina Buldowskia Pletholophus Beringiana Sinanodonta Cristaria bellua Cristaria clessini Cristaria plicata Cristaria truncata

Bogan et al., 2023 [Publ 4295], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 3 2 2 7.0
Cladomics cladogram Sinanodonta Pletholophus Cristaria radiata Cristaria clessini Cristaria plicata Cristaria truncata

Wu et al., 2025 [Publ 4442], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 14 12 7 6.0
Cladomics cladogram Cristaria liboi Cristaria plicata Cristaria truncata Amuranodonta Anemina Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Wang et al., 2025 [Publ 4488], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 61 12 7 6.0
Cladomics cladogram Cristaria liboi Cristaria plicata Cristaria truncata Amuranodonta Anemina Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 92 12 7 6.0
Cladomics cladogram Cristaria liboi Cristaria plicata Cristaria truncata Amuranodonta Anemina Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Wu et al., 2026 [Publ 4825], fig. 6B

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 62 11 7 6.0
Cladomics cladogram Cristaria liboi Cristaria plicata Cristaria truncata Amuranodonta Anemina Buldowskia Beringiana Pletholophus Acudonta Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 4 2 1 6.0
Cladomics cladogram Sinanodonta Cristaria clessini Cristaria plicata Cristaria radiata

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 120 22 6 --
Cladomics cladogram Cristaria plicata Cristaria clessini Anemina Amuranodonta Buldowskia Pletholophus Beringiana Sinanodonta

Froufe et al., 2017 [Publ 3802], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 10 8 3 --
Cladomics cladogram Anemina Cristaria plicata Cristaria radiata Beringiana Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 12

cst iT tT bi riS oT roT roS iD
0.600 2 2 0 2 11 5 3 --
Cladomics cladogram Amuranodonta Cristaria plicata Cristaria

Klishko et al., 2014 [Publ 3666], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 2 2 2 --
Cladomics cladogram Cristaria plicata Cristaria radiata Beringiana Sinanodonta

Klishko et al., 2016 [Publ 3733], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 2 2 2 --
Cladomics cladogram Cristaria plicata Cristaria radiata Sinanodonta Beringiana

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 137 4 2 --
Cladomics cladogram Cristaria plicata Cristaria radiata Pletholophus Sinanodonta

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 137 4 2 --
Cladomics cladogram Cristaria plicata Cristaria radiata Pletholophus Sinanodonta

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
0.250 2 18 0 2 123 4 2 22.0
Cladomics cladogram Unio crassus Unio pictorum Unio tumidus Scriptolamprotula Cuneopsis szechenyii Tchangsinaia Cuneopsis rufescens Nodularia douglasiae Nodularia nuxpersicae Schistodesmus Alasmidonta Anodonta anatina Anodonta cygnea Brachyanodon Lasmigona Pseudanodonta Pyganodon Simpsonaias Strophitus Sinanodonta lauta Cristaria plicata Cristaria radiata Pletholophus Sinanodonta lucida Sinanodonta jourdyi Lanceolaria gladiola Lanceolaria grayii Lanceolaria danzhouensis

Bolotov et al., 2020 [Publ 4084], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 8 5 --
Cladomics cladogram Amuranodonta Anemina Buldowskia Cristaria plicata Beringiana Sinanodonta

Sano et al., 2020 [Publ 4138], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 68 10 5 --
Cladomics cladogram Cristaria plicata Anodonta Pyganodon Utterbackia Alasmidonta Prolasmidonta Platynaias Lanceolaria Lepidodesma Scriptolamprotula Unio Inversiunio Nodularia Tchangsinaia Schistodesmus

Sano et al., 2020 [Publ 4138], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 68 10 5 --
Cladomics cladogram Cristaria plicata Anodonta Pyganodon Utterbackia Alasmidonta Platynaias Prolasmidonta Lanceolaria Lepidodesma Scriptolamprotula Unio Inversiunio Nodularia Schistodesmus Tchangsinaia

Sano et al., 2020 [Publ 4138], fig. 4

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 68 10 5 --
Cladomics cladogram Cristaria plicata Anodonta Pyganodon Utterbackia Alasmidonta Prolasmidonta Platynaias Lepidodesma Lanceolaria Scriptolamprotula Unio Nodularia Inversiunio Schistodesmus Tchangsinaia

Sano et al., 2020 [Publ 4138], fig. 1

cst iT tT bi riS oT roT roS iD
0.286 1 22 5 1 33 9 5 36.0
Cladomics cladogram Sinohyriopsis Lamprotula Pronodularia Sinosolenaia Inversidens Obovalis omiensis 1 Obovalis omiensis 2 Ptychorhynchus Lanceolaria oxyrhyncha 1 Lanceolaria oxyrhyncha 2 Cristaria plicata 1 Cristaria plicata 2 Sinanodonta lauta Beringiana japonica Beringiana fukuharai 1 Beringiana beringiana Beringiana fukuharai 2 Beringiana fukuharai 3 Buldowskia iwakawai 1 Buldowskia iwakawai 2 Sinanodonta tumens 1 Sinanodonta tumens 2 Pletholophus Anemina Lanceolaria lanceolata Cristaria plicata 3 Lanceolaria gladiola Sinanodonta angula Lepidodesma Scriptolamprotula Acuticosta Cuneopsis Nodularia Schistodesmus Inversiunio

Chen et al., 2023 [Publ 4302], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 40 5 5 --
Cladomics cladogram Lanceolaria gladiola Acuticosta Lepidodesma Lanceolaria oxyrhyncha Cristaria plicata Alasmidonta Lasmigona Anodonta Pseudanodonta Scriptolamprotula Middendorffinaia Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia Unio

Chen et al., 2023 [Publ 4302], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 5 5 --
Cladomics cladogram Cristaria plicata Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Dai et al., 2025 [Publ 4440], fig. 1

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 43 5 5 --
Cladomics cladogram Cristaria plicata Alasmidonta Lasmigona Anodonta Pseudanodonta Lanceolaria Lepidodesma Acuticosta Globunio Scriptolamprotula Middendorffinaia Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Unio Tchangsinaia Inversiunio

Dai et al., 2025 [Publ 4440], fig. 2

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 43 5 5 --
Cladomics cladogram Lanceolaria gladiola Lanceolaria oxyrhyncha Cristaria plicata Acuticosta Lepidodesma Globunio Scriptolamprotula Middendorffinaia Nodularia Cuneopsis Diaurora Schistodesmus Pseudocuneopsis Inversiunio Tchangsinaia Unio

Chen et al., 2025 [Publ 4487], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 40 5 5 --
Cladomics cladogram Cristaria plicata Anemina Amuranodonta Pletholophus Sinanodonta Beringiana

Chen et al., 2025 [Publ 4487], fig. 3

cst iT tT bi riS oT roT roS iD
0.857 1 1 0 1 40 5 5 --
Cladomics cladogram Cristaria plicata Lanceolaria

Sano et al., 2017 [Publ 3799], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 7 4 --
Cladomics cladogram Cristaria plicata Buldowskia Pletholophus Sinanodonta Beringiana

Sano et al., 2017 [Publ 3799], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 32 8 4 --
Cladomics cladogram Cristaria plicata Buldowskia Beringiana Pletholophus Sinanodonta

Sano et al., 2022 [Publ 4311], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 11 4 --
Cladomics cladogram Beringiana Sinanodonta Pletholophus Cristaria plicata Buldowskia

Sano et al., 2022 [Publ 4311], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 17 4 --
Cladomics cladogram Beringiana Sinanodonta Pletholophus Cristaria plicata Buldowskia

Xiang et al., 2025 [Publ 4433], fig. 7

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 6 4 --
Cladomics cladogram Cristaria plicata Anemina Beringiana Mimo Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 10

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 5 4 --
Cladomics cladogram Anemina Amuranodonta Cristaria plicata Beringiana Sinanodonta

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 80 4 3 --
Cladomics cladogram Cristaria plicata Anemina Pletholophus Sinanodonta

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 200 8 3 --
Cladomics cladogram Anemina Cristaria plicata Pletholophus Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 198 8 3 --
Cladomics cladogram Anemina Cristaria plicata Pletholophus Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 200 8 3 --
Cladomics cladogram Acuticosta Alasmidonta Anodonta anatina Anemina Anodonta cygnea Cristaria plicata Brachyanodon Lasmigona Lepidodesma Pseudanodonta Pyganodon Simpsonaias Strophitus Utterbackia Schistodesmus Unio Scriptolamprotula Lanceolaria Pletholophus Sinanodonta Cuneopsis celtiformis Tchangsinaia Cuneopsis rufescens Nodularia Cuneopsis szechenyii Cuneopsis heudei

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 5 3 --
Cladomics cladogram Anemina Cristaria plicata Beringiana Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 5 3 --
Cladomics cladogram Cristaria plicata Anemina Beringiana Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 5 3 --
Cladomics cladogram Cristaria plicata Anemina Beringiana Sinanodonta

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 5 3 --
Cladomics cladogram Cristaria plicata Anemina Beringiana Sinanodonta

Wu et al., 2024 [Publ 4379], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 5 3 --
Cladomics cladogram Cristaria plicata Anemina Beringiana Sinanodonta

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 3 3 --
Cladomics cladogram Cristaria plicata Anemina Beringiana Sinanodonta

Hou et al., 2024 [Publ 4400], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 74 5 3 --
Cladomics cladogram Cristaria plicata Anemina Beringiana Sinanodonta

Hou et al., 2025 [Publ 4445], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 51 5 3 --
Cladomics cladogram Cristaria plicata Anemina Beringiana Sinanodonta

Altaba et al., 2025 [Publ 4849], fig. 11

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 3 3 --
Cladomics cladogram Cristaria plicata Amuranodonta Sinanodonta Beringiana

Xue et al., 2016 [Publ 3659], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 14 3 2 --
Cladomics cladogram Cristaria plicata Platynaias

Wu et al., 2016 [Publ 3711], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 20 2 2 --
Cladomics cladogram Cristaria plicata Anodonta Pyganodon Utterbackia

Fonseca et al., 2016 [Publ 3775], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 31 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 81 3 2 --
Cladomics cladogram Cristaria plicata Pletholophus Sinanodonta

Huang et al., 2018 [Publ 3828], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 41 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Wu et al., 2018 [Publ 3836], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 5 2 --
Cladomics cladogram Anemina Cristaria plicata Sinanodonta

Wang et al., 2018 [Publ 3846], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Aunins et al., 2018 [Publ 3863], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Wang et al., 2018 [Publ 3865], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Cha et al., 2018 [Publ 3916], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Wu et al., 2018 [Publ 4143], fig. 2

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 21 2 2 --
Cladomics cladogram Cristaria plicata Anodonta Pyganodon Utterbackia

Huang et al., 2019 [Publ 3843], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 45 7 2 --
Cladomics cladogram Cristaria plicata Lanceolaria

Huang et al., 2019 [Publ 3843], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 55 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Huang et al., 2019 [Publ 3843], fig. S1

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 46 7 2 --
Cladomics cladogram Lepidodesma Cristaria plicata

Wu et al., 2019 [Publ 3864], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Froufe et al., 2019 [Publ 3936], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Froufe et al., 2019 [Publ 3936], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 43 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 2 2 --
Cladomics cladogram Cristaria radiata Anemina Sinanodonta

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 3 2 --
Cladomics cladogram Cristaria plicata Sinanodonta Pletholophus

Riccardi et al., 2020 [Publ 4089], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Fukata & Iigo, 2020 [Publ 4106], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 53 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

cst iT tT bi riS oT roT roS iD
0.500 1 1 0 1 11 4 2 --
Cladomics cladogram Cristaria plicata Sinanodonta tumens Sinanodonta lauta 5 Sinanodonta lauta 6 Beringiana japonica 15 Beringiana japonica 16 Beringiana japonica 17 Beringiana japonica 18 Beringiana japonica 19 Beringiana japonica 20 Beringiana japonica 1 Beringiana japonica 2 Beringiana japonica 3 Beringiana japonica 4 Beringiana japonica 5 Sinanodonta lauta 1 Beringiana japonica 6 Sinanodonta lauta 2 Sinanodonta calipygos 1 Beringiana japonica 7 Sinanodonta lauta 4 Beringiana japonica 8 Beringiana japonica 9 Beringiana japonica 10 Beringiana japonica 11 Beringiana japonica 12 Beringiana japonica 13 Beringiana japonica 14 Sinanodonta lauta 3 Sinanodonta calipygos 2

Wu et al., 2020 [Publ 4121], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Wu et al., 2020 [Publ 4147], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 29 2 2 --
Cladomics cladogram Cristaria plicata Sinanodonta Anemina

Chen et al., 2020 [Publ 4170], fig. 1

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 33 2 2 --
Cladomics cladogram Cristaria plicata Anodonta

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 3 2 --
Cladomics cladogram Anemina Cristaria plicata Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 67 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 3 2 --
Cladomics cladogram Anemina Cristaria plicata Sinanodonta

Wu et al., 2022 [Publ 4238], fig. 2d

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Wu et al., 2022 [Publ 4238], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 72 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 4 2 --
Cladomics cladogram Cristaria plicata Pletholophus Sinanodonta

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 4 2 --
Cladomics cladogram Cristaria plicata Pletholophus Sinanodonta

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 36 2 2 --
Cladomics cladogram Cristaria plicata Sinanodonta Anemina

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 38 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 2 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Bogan et al., 2023 [Publ 4295], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 4 2 --
Cladomics cladogram Cristaria truncata Sinanodonta Pletholophus

Bogan et al., 2023 [Publ 4295], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 12 11 2 --
Cladomics cladogram Pletholophus Cristaria truncata Sinanodonta

Wu et al., 2023 [Publ 4339], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 18 3 2 --
Cladomics cladogram Anemina Cristaria plicata Sinanodonta

Dai et al., 2024 [Publ 4358], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 2 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Dai et al., 2024 [Publ 4389], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 2 2 --
Cladomics cladogram Sinanodonta Anemina Cristaria plicata

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Dai et al., 2025 [Publ 4441], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 2 2 --
Cladomics cladogram Sinanodonta Anemina Cristaria plicata

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 4 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Chen et al., 2025 [Publ 4663], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 2 2 --
Cladomics cladogram Sinanodonta Anemina Cristaria plicata

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 139 3 2 --
Cladomics cladogram Cristaria plicata Anemina Sinanodonta

Huang et al., 2002 [Publ 2657], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 1 1 --
Cladomics cladogram Sinanodonta Cristaria plicata

Huang et al., 2002 [Publ 2657], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 1 1 --
Cladomics cladogram Sinanodonta Cristaria plicata

Zhang et al., 2004 [Publ 2735], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 13 1 1 --
Cladomics cladogram Sinanodonta Cristaria plicata

Zhou et al., 2007 [Publ 2658], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 20 1 1 --
Cladomics cladogram Sinanodonta Cristaria plicata

Zhou et al., 2007 [Publ 2658], fig. 2

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 17 1 1 --
Cladomics cladogram Cristaria plicata Scriptolamprotula Acuticosta Nodularia Cuneopsis Lamprotula Theliderma Quadrula quadrula Pustulosa infucata Quadrula verrucosa Pustulosa kieneriana Pustulosa succissa

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 1 1 --
Cladomics cladogram Cristaria plicata Sinanodonta

Huang et al., 2013 [Publ 3629], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 1 1 --
Cladomics cladogram Sinanodonta Cristaria plicata

Zhang et al., 2013 [Publ 4214], fig. 5

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 3 1 1 --
Cladomics cladogram Cristaria plicata Sinanodonta Margaritifera Lamprotula

Zhou et al., 2016 [Publ 3710], fig. 1

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 19 1 1 --
Cladomics cladogram Cristaria plicata Platynaias Pyganodon Utterbackia

Xia et al., 2016 [Publ 3712], fig. 7B

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 2 1 1 --
Cladomics cladogram Cristaria plicata Sinohyriopsis

Guerra et al., 2017 [Publ 3772], fig. 2a

cst iT tT bi riS oT roT roS iD
0.667 1 1 0 1 24 1 1 --
Cladomics cladogram Cristaria plicata Anodonta Platynaias Pyganodon Utterbackia

Guerra et al., 2017 [Publ 3772], fig. 2b

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 24 1 1 --
Cladomics cladogram Cristaria plicata Sinanodonta

Wen et al., 2017 [Publ 3795], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 15 1 1 --
Cladomics cladogram Sinanodonta Cristaria plicata

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 1 1 --
Cladomics cladogram Cristaria plicata Sinanodonta

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 1 1 --
Cladomics cladogram Cristaria plicata Sinanodonta

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 11 1 1 --
Cladomics cladogram Cristaria plicata Sinanodonta

Wu et al., 2024 [Publ 4379], fig. 4_6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 2 1 --
Cladomics cladogram Cristaria plicata Sinanodonta

Jiang et al., 2010 [Publ 3438], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram Lampsilis Cristaria plicata Pronodularia Sinohyriopsis

Breton et al., 2011 [Publ 3376], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 37 0 0 --
Cladomics cladogram Cristaria plicata Pterosyna Lasmigona Strophitus Platynaias Alasmidonta Pyganodon Utterbackia

Plazzi et al., 2011 [Publ 3676], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Cristaria plicata Lanceolaria Anodonta Unio Pyganodon Pseudanodonta Alathyria Pronodularia Sinohyriopsis Quadrula Parvaspina Lampsilis Epioblasma Venustaconcha

Chen et al., 2012 [Publ 3624], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 0 0 --
Cladomics cladogram Pyganodon Cristaria plicata

Peng et al., 2012 [Publ 3689], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 2 0 0 --
Cladomics cladogram Cristaria plicata Sinohyriopsis

Sheng et al., 2014 [Publ 3704], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 7 0 0 --
Cladomics cladogram Cristaria plicata Pyganodon Lampsilis Venustaconcha Quadrula Lamprotula Sinohyriopsis

Wei et al., 2016 [Publ 3665], fig. 0

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 5 0 0 --
Cladomics cladogram Cristaria plicata Lampsilis Pyganodon Quadrula

Froufe et al., 2017 [Publ 3802], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 3 0 0 --
Cladomics cladogram Cristaria plicata Anodonta

Smith et al., 2018 [Publ 3790], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Cristaria plicata Utterbackia Pyganodon Anodontoides Strophitus Alasmidonta Alasminota Pseudodontoideus Utterbackiana

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Cristaria plicata Pyganodon Utterbackia