Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Limnoscaphini | Contradentina

genus Physunio Simpson, 1900: cladomics

4 Recent valid species (4 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (4 total)
analyzed in 65 cladograms in 43 publications
maximum number of terminals in an analysis = 5 (mean = 1.62, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 22
maximum number of included terminals = 5, (mean = 2.82 median = 3)
proportion of cladograms that support basic monophyly = 0.86

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Physunio cladogram distances
PCoA Ordination

NJ Tree of Physunio cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram **

cst iT tT bi riS oT roT roS iD
0.800 5 31 30 5 628 26 3 **
Cladomics cladogram Physunio modelli Physunio pellucidus Physunio massini Physunio superbus Physunio Trapezoideus Yaukthwa Lens Ensidens Rectidens Hyriopsis delaportei Hyriopsis desowitzi Hyriopsis panhai Hyriopsis bialata Hyriopsis khoratensis Hyriopsis kratiensis Hyriopsis phuphaniensis Hyriopsis myersiana Hyriopsis sakhonensis

Pfeiffer et al., 2021 [Publ 4126], fig. 9

** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 26 12 4 29.0
Cladomics cladogram Yaukthwa Solenaia Lens Trapezoideus Physunio superbus Physunio modelli Physunio pellucidus

Pfeiffer et al., 2021 [Publ 4126], fig. 18

cst iT tT bi riS oT roT roS iD
1.000 4 4 3 4 4 4 2 29.0
Cladomics cladogram Yaukthwa Trapezoideus Physunio massini Physunio superbus Physunio modelli 3 Physunio modelli 1 Physunio modelli 2 Physunio pellucidus

Konopleva et al., 2019 [Publ 3900], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 48 13 3 31.0
Cladomics cladogram Yaukthwa Trapezoideus Lens Physunio superbus Physunio modelli Physunio

Konopleva et al., 2019 [Publ 4041], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 31 16 3 31.0
Cladomics cladogram Yaukthwa Lens Trapezoideus Physunio superbus Physunio modelli Physunio

Bolotov et al., 2020 [Publ 4100], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 114 20 3 31.0
Cladomics cladogram Yaukthwa Lens Trapezoideus Physunio superbus Physunio modelli Physunio

Bolotov et al., 2020 [Publ 4100], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 108 20 3 31.0
Cladomics cladogram Physunio Physunio modelli Physunio superbus Trapezoideus Lens Yaukthwa

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 18 7 3 29.0
Cladomics cladogram Yaukthwa Lens Trapezoideus Physunio superbus Physunio pellucidus Physunio modelli

Konopleva et al., 2023 [Publ 4262], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 79 16 3 29.0
Cladomics cladogram Yaukthwa Lens Trapezoideus Physunio modelli Physunio superbus Physunio massini

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 136 9 2 31.0
Cladomics cladogram Yaukthwa Lens Physunio Physunio modelli Physunio superbus

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 136 8 2 31.0
Cladomics cladogram Lens Yaukthwa Physunio Physunio modelli Physunio superbus

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 122 9 2 31.0
Cladomics cladogram Yaukthwa Lens Physunio Physunio superbus Physunio modelli

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 198 9 2 31.0
Cladomics cladogram Yaukthwa Lens Physunio Physunio modelli Physunio superbus

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 196 9 2 31.0
Cladomics cladogram Yaukthwa Lens Physunio Physunio superbus Physunio modelli

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.750 3 12 7 3 198 9 2 26.0
Cladomics cladogram Physunio modelli Physunio Physunio superbus Yaukthwa Lens Hyriopsis desowitzi Hyriopsis myersiana Hyriopsis panhai Hyriopsis bialata Hyriopsis khoratensis Rectidens Ensidens

Zieritz et al., 2020 [Publ 4102], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 12 3 --
Cladomics cladogram Physunio modelli Physunio superbus Yaukthwa Trapezoideus Lens

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
0.250 2 10 0 2 80 8 2 32.0
Cladomics cladogram Ensidens telus Ensidens ingallsianus Ensidens jaculus Hyriopsis desowitzi Hyriopsis myersiana Hyriopsis khoratensis Ensidens spiculus Ensidens sagittarius Lens eximius Physunio modelli 1 Yaukthwa nesemanni Yaukthwa panhai Lens comptus Lens novoselovi Physunio modelli 2 Physunio Yaukthwa inlenensis Lens contradens Lens micropterus

Bolotov et al., 2022 [Publ 4250], fig. S5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 13 13 2 --
Cladomics cladogram Yaukthwa Trapezoideus Physunio modelli Physunio superbus

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 38 1 1 --
Cladomics cladogram Lens Physunio Physunio modelli

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 79 2 1 --
Cladomics cladogram Lens Physunio modelli Physunio

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 1 1 --
Cladomics cladogram Lens Physunio Physunio modelli

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 7 4 1 --
Cladomics cladogram Lens Physunio modelli Physunio superbus

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 4 4 --
Cladomics cladogram Trapezoideus Physunio superbus Yaukthwa Lens Solenaia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 4 4 --
Cladomics cladogram Trapezoideus Physunio superbus Yaukthwa Lens Solenaia

Zieritz et al., 2021 [Publ 4218], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 6 3 --
Cladomics cladogram Physunio superbus Trapezoideus Lens Yaukthwa

Wu et al., 2024 [Publ 4350], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 6 3 --
Cladomics cladogram Physunio superbus Trapezoideus Lens Yaukthwa

Wu et al., 2024 [Publ 4350], fig. 5b

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 35 6 3 --
Cladomics cladogram Yaukthwa elongatula Trapezoideus Physunio superbus Yaukthwa inlenensis Yaukthwa paiensis Lens

Wu et al., 2024 [Publ 4350], fig. 5c

cst iT tT bi riS oT roT roS iD
0.800 1 1 0 1 35 6 3 --
Cladomics cladogram Physunio superbus Yaukthwa paiensis Yaukthwa inlenensis Trapezoideus Yaukthwa elongatula Lens

Wu et al., 2024 [Publ 4350], fig. 6-7

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 6 3 --
Cladomics cladogram Physunio superbus Trapezoideus Lens Yaukthwa

Liu et al., 2024 [Publ 4380], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 6 3 --
Cladomics cladogram Physunio superbus Trapezoideus Lens Yaukthwa

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 7 3 --
Cladomics cladogram Physunio superbus Trapezoideus Yaukthwa Lens

Wu et al., 2024 [Publ 4388], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 6 3 --
Cladomics cladogram Physunio superbus Trapezoideus Lens Yaukthwa

Pfeiffer et al., 2018 [Publ 3827], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 6 3 2 --
Cladomics cladogram Physunio Trapezoideus Lens

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 121 2 2 --
Cladomics cladogram Lens Ensidens Physunio modelli Rectidens Yaukthwa

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 2 2 --
Cladomics cladogram Lens Trapezoideus Physunio superbus

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 34 2 2 --
Cladomics cladogram Lens Trapezoideus Physunio superbus

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 3 2 --
Cladomics cladogram Physunio modelli Lens Yaukthwa

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 3 2 --
Cladomics cladogram Physunio modelli Lens Yaukthwa

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 2 2 --
Cladomics cladogram Lens Trapezoideus Physunio superbus

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 2 2 --
Cladomics cladogram Lens Trapezoideus Physunio superbus

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 58 2 2 --
Cladomics cladogram Physunio superbus Lens Trapezoideus

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 1 1 --
Cladomics cladogram Lens Physunio superbus

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 1 1 --
Cladomics cladogram Lens Physunio superbus

Dai et al., 2023 [Publ 4285], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 2 1 --
Cladomics cladogram Physunio superbus Lens

Dai et al., 2023 [Publ 4285], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 44 2 1 --
Cladomics cladogram Physunio superbus Lens

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Lens Physunio superbus

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Lens Physunio superbus

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Lens Physunio superbus

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Lens Physunio superbus

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Lens Physunio superbus

Matos et al., 2024 [Publ 4383], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 1 1 --
Cladomics cladogram Physunio superbus Lens

Matos et al., 2024 [Publ 4384], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 1 1 --
Cladomics cladogram Physunio superbus Lens

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Lens Physunio superbus

Dai et al., 2024 [Publ 4389], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 2 1 --
Cladomics cladogram Physunio superbus Lens

Zhang et al., 2024 [Publ 4397], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 26 1 1 --
Cladomics cladogram Lens Physunio superbus

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 1 1 --
Cladomics cladogram Lens Physunio superbus

Dai et al., 2025 [Publ 4441], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 2 1 --
Cladomics cladogram Physunio superbus Lens

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 1 1 --
Cladomics cladogram Lens Physunio superbus

Chen et al., 2025 [Publ 4663], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 2 1 --
Cladomics cladogram Physunio superbus Lens

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 1 1 --
Cladomics cladogram Lens Physunio superbus

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 1 1 --
Cladomics cladogram Lens Physunio superbus

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 139 1 1 --
Cladomics cladogram Lens Physunio superbus

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 1 1 --
Cladomics cladogram Lens Physunio superbus

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Lens Physunio superbus

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 0 0 --
Cladomics cladogram Physunio Pronodularia Potomida Lamprotula