Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Rectidentini | Contradentina

genus Physunio Simpson, 1900: cladomics

4 Recent valid species (4 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 4 (4 total)
analyzed in 65 cladograms in 43 publications
maximum number of terminals in an analysis = 5 (mean = 1.62, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 22
maximum number of included terminals = 5, (mean = 2.82 median = 3)
proportion of cladograms that support basic monophyly = 0.86

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Physunio cladogram distances
PCoA Ordination

NJ Tree of Physunio cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram **

cstiTtTbiriSoTroTroSiD
0.800 5 31 30 5 627 26 3 ** 
                    +-- Physunio_modelli **       
        +-----------+                          
        |           +-- Physunio_pellucidus **    
        |                                      
  +-----+           +-- Physunio_massini **       
  |     |  +--------+                          
  |     |  |        +-- Physunio_superbus **      
  |     +--+                                   
  |        |  +-------- Physunio **               
  |        +--+                                
  |           |  +----- Trapezoideus           
--+           +--+                             
  |              |  +-- Yaukthwa               
  |              +--+                          
  |                 +-- Lens                   
  |                                            
  |  +----------------- Ensidens               
  |  |                                         
  |  |           +----- Rectidens              
  +--+  +--------+                             
     |  |        |  +-- Hyriopsis_delaportei   
     |  |        +--+                          
     |  |           +-- Hyriopsis_desowitzi    
     +--+                                      
        |  +----------- Hyriopsis_panhai       
        |  |                                   
        |  |        +-- Hyriopsis_bialata      
        +--+  +-----+                          
           |  |     +-- Hyriopsis_khoratensis  
           |  |                                
           +--+     +-- Hyriopsis_kratiensis   
              |  +--+                          
              |  |  +-- Hyriopsis_phuphaniensis
              +--+                             
                 |  +-- Hyriopsis_myersiana    
                 +--+                          
                    +-- Hyriopsis_sakhonensis

Pfeiffer et al., 2021 [Publ 4126], fig. 9

** Top Sister-Group Cladogram **

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 26 12 4 29.0 
        +----- Yaukthwa           
  +-----+                         
  |     |  +-- Solenaia           
  |     +--+                      
--+        +-- Lens               
  |                               
  |  +-------- Trapezoideus       
  +--+                            
     |  +----- Physunio_superbus **  
     +--+                         
        |  +-- Physunio_modelli **   
        +--+                      
           +-- Physunio_pellucidus **

Pfeiffer et al., 2021 [Publ 4126], fig. 18

cstiTtTbiriSoTroTroSiD
1.000 4 4 3 4 4 4 2 29.0 
  +-------------- Yaukthwa           
  |                                  
--+  +----------- Trapezoideus       
  |  |                               
  +--+        +-- Physunio_massini **   
     |  +-----+                      
     |  |     +-- Physunio_superbus **  
     +--+                            
        |  +----- Physunio_modelli__3 **
        |  |                         
        +--+  +-- Physunio_modelli__1 **
           |  |                      
           +--+-- Physunio_modelli__2 **
              |                      
              +-- Physunio_pellucidus **

Konopleva et al., 2019 [Publ 3900], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 48 13 3 31.0 
  +----------- Yaukthwa         
  |                             
--+        +-- Trapezoideus     
  |  +-----+                    
  |  |     +-- Lens             
  +--+                          
     |  +----- Physunio_superbus **
     +--+                       
        |  +-- Physunio_modelli ** 
        +--+                    
           +-- Physunio **

Konopleva et al., 2019 [Publ 4041], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 31 16 3 31.0 
  +-------------- Yaukthwa         
--+                                
  |  +----------- Lens             
  +--+                             
     |  +-------- Trapezoideus     
     +--+                          
        |  +----- Physunio_superbus **
        +--+                       
           |  +-- Physunio_modelli ** 
           +--+                    
              +-- Physunio **

Bolotov et al., 2020 [Publ 4100], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 114 20 3 31.0 
           +-- Yaukthwa         
  +--------+                    
  |        +-- Lens             
--+                             
  |  +-------- Trapezoideus     
  +--+                          
     |  +----- Physunio_superbus **
     +--+                       
        |  +-- Physunio_modelli ** 
        +--+                    
           +-- Physunio **

Bolotov et al., 2020 [Publ 4100], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 108 20 3 31.0 
     +----- Physunio **         
  +--+                       
  |  |  +-- Physunio_modelli ** 
  |  +--+                    
--+     +-- Physunio_superbus **
  |                          
  |  +----- Trapezoideus     
  +--+                       
     |  +-- Lens             
     +--+                    
        +-- Yaukthwa

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 18 7 3 29.0 
           +-- Yaukthwa           
  +--------+                      
  |        +-- Lens               
--+                               
  |  +-------- Trapezoideus       
  +--+                            
     |  +----- Physunio_superbus **  
     +--+                         
        |  +-- Physunio_pellucidus **
        +--+                      
           +-- Physunio_modelli **

Konopleva et al., 2023 [Publ 4262], fig. 8

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 79 16 3 29.0 
           +-- Yaukthwa         
  +--------+                    
  |        +-- Lens             
--+                             
  |  +-------- Trapezoideus     
  +--+                          
     |  +----- Physunio_modelli ** 
     +--+                       
        |  +-- Physunio_superbus **
        +--+                    
           +-- Physunio_massini **

Bolotov et al., 2017 [Publ 3761], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 136 9 2 31.0 
        +-- Yaukthwa         
  +-----+                    
  |     +-- Lens             
--+                          
  |  +----- Physunio **         
  +--+                       
     |  +-- Physunio_modelli ** 
     +--+                    
        +-- Physunio_superbus **

Bolotov et al., 2017 [Publ 3761], fig. S2

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 136 8 2 31.0 
  +----------- Lens             
--+                             
  |  +-------- Yaukthwa         
  +--+                          
     |  +----- Physunio **         
     +--+                       
        |  +-- Physunio_modelli ** 
        +--+                    
           +-- Physunio_superbus **

Bolotov et al., 2017 [Publ 3764], fig. S1

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 122 9 2 31.0 
        +-- Yaukthwa         
  +-----+                    
  |     +-- Lens             
--+                          
  |  +----- Physunio **         
  +--+                       
     |  +-- Physunio_superbus **
     +--+                    
        +-- Physunio_modelli **

Huang et al., 2019 [Publ 3843], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 198 9 2 31.0 
        +-- Yaukthwa         
  +-----+                    
  |     +-- Lens             
--+                          
  |  +----- Physunio **         
  +--+                       
     |  +-- Physunio_modelli ** 
     +--+                    
        +-- Physunio_superbus **

Huang et al., 2019 [Publ 3843], fig. S3

cstiTtTbiriSoTroTroSiD
1.000 3 3 1 3 196 9 2 31.0 
        +-- Yaukthwa         
  +-----+                    
  |     +-- Lens             
--+                          
  |  +----- Physunio **         
  +--+                       
     |  +-- Physunio_superbus **
     +--+                    
        +-- Physunio_modelli **

Huang et al., 2019 [Publ 3843], fig. S4

cstiTtTbiriSoTroTroSiD
0.750 3 12 7 3 198 9 2 26.0 
        +-- Physunio_modelli **     
        |                        
        +-- Physunio **             
        |                        
  +-----+-- Physunio_superbus **    
  |     |                        
  |     +-- Yaukthwa             
  |     |                        
  |     +-- Lens                 
--+                              
  |  +----- Hyriopsis_desowitzi  
  |  |                           
  |  +----- Hyriopsis_myersiana  
  |  |                           
  |  +----- Hyriopsis_panhai     
  +--+                           
     |  +-- Hyriopsis_bialata    
     +--+                        
     |  +-- Hyriopsis_khoratensis
     |                           
     |  +-- Rectidens            
     +--+                        
        +-- Ensidens

Zieritz et al., 2020 [Publ 4102], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 29 12 3 -- 
        +-- Physunio_modelli ** 
  +-----+                    
  |     +-- Physunio_superbus **
--+                          
  |  +----- Yaukthwa         
  +--+                       
     |  +-- Trapezoideus     
     +--+                    
        +-- Lens

Bolotov et al., 2018 [Publ 3826], fig. 2

cstiTtTbiriSoTroTroSiD
0.250 2 10 0 2 80 8 2 32.0 
     +-- Ensidens_telus       
     |                        
     +-- Ensidens_ingallsianus
     |                        
     +-- Ensidens_jaculus     
     |                        
     +-- Hyriopsis_desowitzi  
  +--+                        
  |  +-- Hyriopsis_myersiana  
  |  |                        
  |  +-- Hyriopsis_khoratensis
  |  |                        
  |  +-- Ensidens_spiculus    
  |  |                        
  |  +-- Ensidens_sagittarius 
  |                           
--+  +-- Lens_eximius         
  |  |                        
  |  +-- Physunio_modelli__1 **  
  |  |                        
  |  +-- Yaukthwa_nesemanni   
  |  |                        
  |  +-- Yaukthwa_panhai      
  |  |                        
  |  +-- Lens_comptus         
  |  |                        
  +--+-- Lens_novoselovi      
     |                        
     +-- Physunio_modelli__2 **  
     |                        
     +-- Physunio **             
     |                        
     +-- Yaukthwa_inlenensis  
     |                        
     +-- Lens_contradens      
     |                        
     +-- Lens_micropterus

Bolotov et al., 2022 [Publ 4250], fig. S5

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 13 13 2 -- 
  +-------- Yaukthwa         
--+                          
  |  +----- Trapezoideus     
  +--+                       
     |  +-- Physunio_modelli ** 
     +--+                    
        +-- Physunio_superbus **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 38 1 1 -- 
  +----- Lens            
--+                      
  |  +-- Physunio **        
  +--+                   
     +-- Physunio_modelli **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 79 2 1 -- 
  +----- Lens            
--+                      
  |  +-- Physunio_modelli **
  +--+                   
     +-- Physunio **

Konopleva et al., 2017 [Publ 3807], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 29 1 1 -- 
  +----- Lens            
--+                      
  |  +-- Physunio **        
  +--+                   
     +-- Physunio_modelli **

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 2 2 0 2 7 4 1 -- 
  +----- Lens             
--+                       
  |  +-- Physunio_modelli ** 
  +--+                    
     +-- Physunio_superbus **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 4 4 -- 
        +-- Trapezoideus     
  +-----+                    
  |     +-- Physunio_superbus **
--+                          
  |  +----- Yaukthwa         
  +--+                       
     |  +-- Lens             
     +--+                    
        +-- Solenaia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 56 4 4 -- 
        +-- Trapezoideus     
  +-----+                    
  |     +-- Physunio_superbus **
--+                          
  |  +----- Yaukthwa         
  +--+                       
     |  +-- Lens             
     +--+                    
        +-- Solenaia

Zieritz et al., 2021 [Publ 4218], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 6 3 -- 
  +-------- Physunio_superbus **
--+                          
  |  +----- Trapezoideus     
  +--+                       
     |  +-- Lens             
     +--+                    
        +-- Yaukthwa

Wu et al., 2024 [Publ 4350], fig. 5a

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 6 3 -- 
     +-- Physunio_superbus **
  +--+                    
  |  +-- Trapezoideus     
--+                       
  |  +-- Lens             
  +--+                    
     +-- Yaukthwa

Wu et al., 2024 [Publ 4350], fig. 5b

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 35 6 3 -- 
  +----------- Yaukthwa_elongatula
  |                               
--+  +-------- Trapezoideus       
  |  |                            
  +--+     +-- Physunio_superbus **  
     |  +--+                      
     |  |  +-- Yaukthwa_inlenensis
     +--+                         
        |  +-- Yaukthwa_paiensis  
        +--+                      
           +-- Lens

Wu et al., 2024 [Publ 4350], fig. 5c

cstiTtTbiriSoTroTroSiD
0.800 1 1 0 1 35 6 3 -- 
  +-- Physunio_superbus **  
  |                      
  +-- Yaukthwa_paiensis  
  |                      
  +-- Yaukthwa_inlenensis
--+                      
  +-- Trapezoideus       
  |                      
  +-- Yaukthwa_elongatula
  |                      
  +-- Lens

Wu et al., 2024 [Publ 4350], fig. 6-7

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 6 3 -- 
     +-- Physunio_superbus **
  +--+                    
  |  +-- Trapezoideus     
--+                       
  |  +-- Lens             
  +--+                    
     +-- Yaukthwa

Liu et al., 2024 [Publ 4380], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 6 3 -- 
     +-- Physunio_superbus **
  +--+                    
  |  +-- Trapezoideus     
--+                       
  |  +-- Lens             
  +--+                    
     +-- Yaukthwa

Zieritz et al., 2024 [Publ 4381], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 44 7 3 -- 
     +-- Physunio_superbus **
  +--+                    
  |  +-- Trapezoideus     
--+                       
  |  +-- Yaukthwa         
  +--+                    
     +-- Lens

Wu et al., 2024 [Publ 4388], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 6 3 -- 
     +-- Physunio_superbus **
  +--+                    
  |  +-- Trapezoideus     
--+                       
  |  +-- Lens             
  +--+                    
     +-- Yaukthwa

Pfeiffer et al., 2018 [Publ 3827], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 6 3 2 -- 
  +-- Physunio **    
  |               
--+-- Trapezoideus
  |               
  +-- Lens

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cstiTtTbiriSoTroTroSiD
0.750 1 1 0 1 121 2 2 -- 
  +-- Lens            
  |                   
  +-- Ensidens        
  |                   
--+-- Physunio_modelli **
  |                   
  +-- Rectidens       
  |                   
  +-- Yaukthwa

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 33 2 2 -- 
  +----- Lens             
--+                       
  |  +-- Trapezoideus     
  +--+                    
     +-- Physunio_superbus **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 34 2 2 -- 
  +----- Lens             
--+                       
  |  +-- Trapezoideus     
  +--+                    
     +-- Physunio_superbus **

Wu et al., 2021 [Publ 4366], fig. 2_I

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 3 2 -- 
  +----- Physunio_modelli **
--+                      
  |  +-- Lens            
  +--+                   
     +-- Yaukthwa

Wu et al., 2021 [Publ 4366], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 3 2 -- 
  +----- Physunio_modelli **
--+                      
  |  +-- Lens            
  +--+                   
     +-- Yaukthwa

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 75 2 2 -- 
  +----- Lens             
--+                       
  |  +-- Trapezoideus     
  +--+                    
     +-- Physunio_superbus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 75 2 2 -- 
  +----- Lens             
--+                       
  |  +-- Trapezoideus     
  +--+                    
     +-- Physunio_superbus **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 58 2 2 -- 
  +----- Physunio_superbus **
--+                       
  |  +-- Lens             
  +--+                    
     +-- Trapezoideus

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 22 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Dai et al., 2023 [Publ 4285], fig. 3-BI

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 44 2 1 -- 
  +-- Physunio_superbus **
--+                    
  +-- Lens

Dai et al., 2023 [Publ 4285], fig. 3-ML

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 44 2 1 -- 
  +-- Physunio_superbus **
--+                    
  +-- Lens

Dai et al., 2023 [Publ 4285], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 90 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Matos et al., 2024 [Publ 4383], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 1 1 -- 
  +-- Physunio_superbus **
--+                    
  +-- Lens

Matos et al., 2024 [Publ 4384], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 19 1 1 -- 
  +-- Physunio_superbus **
--+                    
  +-- Lens

Wu et al., 2024 [Publ 4388], fig. 6

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 52 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Dai et al., 2024 [Publ 4389], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 45 2 1 -- 
  +-- Physunio_superbus **
--+                    
  +-- Lens

Zhang et al., 2024 [Publ 4397], fig. 5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 26 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Dai et al., 2025 [Publ 4440], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 86 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Dai et al., 2025 [Publ 4441], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 46 2 1 -- 
  +-- Physunio_superbus **
--+                    
  +-- Lens

Chen et al., 2025 [Publ 4448], fig. 3

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 84 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Chen et al., 2025 [Publ 4663], fig. 1

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 47 2 1 -- 
  +-- Physunio_superbus **
--+                    
  +-- Lens

Klymus et al., 2025 [Publ 4814], fig. 3A

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Klymus et al., 2025 [Publ 4814], fig. 3B

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 35 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 139 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 46 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Wu et al., 2026 [Publ 4811], fig. 4

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 94 1 1 -- 
  +-- Lens             
--+                    
  +-- Physunio_superbus **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

cstiTtTbiriSoTroTroSiD
1.000 1 1 0 1 30 0 0 -- 
  +-------- Physunio **    
--+                     
  |  +----- Pronodularia
  +--+                  
     |  +-- Potomida    
     +--+               
        +-- Lamprotula