Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Rectidentini | Contradentina

genus Lens Simpson, 1900: cladomics

13 Recent valid species (13 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 13 (13 total)
analyzed in 71 cladograms in 46 publications
maximum number of terminals in an analysis = 11 (mean = 2.94, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 35
maximum number of included terminals = 11, (mean = 4.94 median = 4)
proportion of cladograms that support basic monophyly = 0.97

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lens cladogram distances
PCoA Ordination

NJ Tree of Lens cladogram distances
NJ Tree

Cladograms

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
4 4 9 17 5 2 1 0 
  +----------------------- Physunio           
  |                                           
  |                    +-- Lens_maenamensis **   
--+  +-----------------+                      
  |  |                 +-- Lens_rolfbrandti **   
  |  |                                        
  |  |  +----------------- Lens_contradens__12 **
  +--+  |                                     
     |  |           +----- Lens_micropterus__3 **
     |  |           |                         
     |  +-----------+  +-- Lens_contradens__10 **
     |  |           |  |                      
     +--+           +--+-- Lens_micropterus__2 **
        |              |                      
        |              +-- Lens_contradens__11 **
        |                                     
        |  +-------------- Lens_contradens__9 ** 
        |  |                                  
        +--+  +----------- Lens_contradens__8 ** 
           |  |                               
           +--+  +-------- Lens_contradens__7 ** 
              |  |                            
              +--+  +----- Lens_contradens__6 ** 
                 |  |                         
                 |  |  +-- Lens_contradens__1 ** 
                 +--+  |                      
                    |  +-- Lens_contradens__2 ** 
                    |  |                      
                    |  +-- Lens_contradens__3 ** 
                    +--+                      
                       +-- Lens_contradens__4 ** 
                       |                      
                       +-- Lens_micropterus__1 **
                       |                      
                       +-- Lens_contradens__5 **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
8 8 6 8 21 7 4 15 
                    +-- Trapezoideus    
  +-----------------+                   
  |                 +-- Physunio        
  |                                     
--+  +----------------- Yaukthwa        
  |  |                                  
  |  |  +-------------- Solenaia        
  +--+  |                               
     |  |     +-------- Lens_misellus **   
     |  |  +--+                         
     +--+  |  |  +----- Lens_inornatus **  
        |  |  +--+                      
        |  |     |  +-- Lens_contradens ** 
        |  |     +--+                   
        +--+        +-- Lens_micropterus **
           |                            
           |        +-- Lens_maenamensis **
           |     +--+                   
           |     |  +-- Lens_eximius **    
           +-----+                      
                 |  +-- Lens_pallegoixi ** 
                 +--+                   
                    +-- Lens_comptus **

Zieritz et al., 2026 [Publ 4812], fig. 2a

iTtTbiriSoTroTroSiD
11 11 9 11 1 1 1 14 
  +----------------- Solenaia        
  |                                  
  |              +-- Lens_inornatus **  
--+  +-----------+                   
  |  |           +-- Lens_misellus **   
  |  |                               
  |  |        +----- Lens_contradens ** 
  +--+  +-----+                      
     |  |     |  +-- Lens_lugens **     
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_comptus **    
        |  |  +--+                   
        +--+     +-- Lens_novoselovi ** 
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_rolfbrandti **
              +--+                   
                 +-- Lens_eximius **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
11 11 10 12 620 20 3 14 
                             +-- Physunio_modelli   
  +--------------------------+                      
  |                          +-- Physunio_pellucidus
  |                                                 
--+                          +-- Physunio_massini   
  |  +-----------------------+                      
  |  |                       +-- Physunio_superbus  
  +--+                                              
     |  +----------------------- Physunio           
     |  |                                           
     +--+  +-------------------- Trapezoideus       
        |  |                                        
        |  |  +----------------- Yaukthwa           
        +--+  |                                     
           |  |              +-- Lens_inornatus **     
           |  |        +-----+                      
           |  |        |     +-- Lens_misellus **      
           +--+  +-----+                            
              |  |     |  +----- Lens_contradens **    
              |  |     +--+                         
              |  |        |  +-- Lens_micropterus **   
              |  |        +--+                      
              +--+           +-- Lens_lugens **        
                 |                                  
                 |        +----- Lens_pallegoixi **    
                 |  +-----+                         
                 |  |     |  +-- Lens_comptus **       
                 |  |     +--+                      
                 +--+        +-- Lens_novoselovi **    
                    |                               
                    |  +-------- Lens_maenamensis **   
                    +--+                            
                       |  +----- Lens_eximius__2 **    
                       +--+                         
                          |  +-- Lens_eximius__1 **    
                          +--+                      
                             +-- Lens_rolfbrandti **

Zieritz et al., 2024 [Publ 4381], fig. 7A

iTtTbiriSoTroTroSiD
11 11 9 11 1 1 1 14 
  +----------------- Solenaia        
  |                                  
  |           +----- Lens_contradens ** 
--+  +--------+                      
  |  |        |  +-- Lens_lugens **     
  |  |        +--+                   
  |  |           +-- Lens_micropterus **
  +--+                               
     |           +-- Lens_inornatus **  
     |  +--------+                   
     |  |        +-- Lens_misellus **   
     |  |                            
     +--+     +----- Lens_maenamensis **
        |  +--+                      
        |  |  |  +-- Lens_eximius **    
        |  |  +--+                   
        +--+     +-- Lens_rolfbrandti **
           |                         
           |  +----- Lens_pallegoixi ** 
           +--+                      
              |  +-- Lens_comptus **    
              +--+                   
                 +-- Lens_novoselovi **

Pfeiffer et al., 2021 [Publ 4126], fig. 12

iTtTbiriSoTroTroSiD
10 10 7 10 1 1 1 14 
  +-------------- Solenaia        
  |                               
  |           +-- Lens_contradens ** 
  |  +--------+                   
--+  |        +-- Lens_micropterus **
  |  |                            
  |  |        +-- Lens_inornatus **  
  |  +--------+                   
  +--+        +-- Lens_misellus **   
     |                            
     |     +----- Lens_maenamensis **
     |  +--+                      
     |  |  |  +-- Lens_rolfbrandti **
     |  |  +--+                   
     +--+     +-- Lens_eximius **    
        |                         
        |  +----- Lens_pallegoixi ** 
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
9 9 7 9 73 10 3 14 
                 +-- Physunio        
  +--------------+                   
  |              +-- Trapezoideus    
  |                                  
--+  +-------------- Yaukthwa        
  |  |                               
  |  |        +----- Lens_misellus **   
  +--+  +-----+                      
     |  |     |  +-- Lens_contradens ** 
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_novoselovi ** 
        |  |  +--+                   
        +--+     +-- Lens_comptus **    
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_eximius **    
              +--+                   
                 +-- Lens_rolfbrandti **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
9 9 7 9 108 14 3 14 
                 +-- Trapezoideus    
  +--------------+                   
  |              +-- Physunio        
  |                                  
--+  +-------------- Yaukthwa        
  |  |                               
  |  |        +----- Lens_misellus **   
  +--+  +-----+                      
     |  |     |  +-- Lens_contradens ** 
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_comptus **    
        |  |  +--+                   
        +--+     +-- Lens_novoselovi ** 
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_eximius **    
              +--+                   
                 +-- Lens_rolfbrandti **

Konopleva et al., 2019 [Publ 4041], fig. 3

iTtTbiriSoTroTroSiD
9 9 7 9 25 10 3 14 
  +----------------- Yaukthwa        
  |                                  
  |              +-- Trapezoideus    
--+  +-----------+                   
  |  |           +-- Physunio        
  |  |                               
  |  |        +----- Lens_misellus **   
  +--+  +-----+                      
     |  |     |  +-- Lens_contradens ** 
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_novoselovi ** 
        |  |  +--+                   
        +--+     +-- Lens_comptus **    
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_eximius **    
              +--+                   
                 +-- Lens_rolfbrandti **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
9 9 7 9 102 14 3 14 
  +-------------------- Physunio        
  |                                     
--+  +----------------- Trapezoideus    
  |  |                                  
  +--+  +-------------- Yaukthwa        
     |  |                               
     |  |        +----- Lens_misellus **   
     +--+  +-----+                      
        |  |     |  +-- Lens_contradens ** 
        |  |     +--+                   
        |  |        +-- Lens_micropterus **
        +--+                            
           |     +----- Lens_maenamensis **
           |  +--+                      
           |  |  |  +-- Lens_eximius **    
           |  |  +--+                   
           +--+     +-- Lens_rolfbrandti **
              |                         
              |  +----- Lens_pallegoixi ** 
              +--+                      
                 |  +-- Lens_novoselovi ** 
                 +--+                   
                    +-- Lens_comptus **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
7 7 5 7 44 9 3 16 
  +----------------------- Yaukthwa        
  |                                        
--+  +-------------------- Physunio        
  |  |                                     
  +--+  +----------------- Trapezoideus    
     |  |                                  
     +--+              +-- Lens_contradens ** 
        |  +-----------+                   
        |  |           +-- Lens_micropterus **
        +--+                               
           |  +----------- Lens_misellus **   
           +--+                            
              |  +-------- Lens_eximius **    
              +--+                         
                 |  +----- Lens_pallegoixi ** 
                 +--+                      
                    |  +-- Lens_comptus **    
                    +--+                   
                       +-- Lens_novoselovi **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
6 6 4 6 25 8 3 18 
  +-------------------- Physunio       
  |                                    
--+  +----------------- Yaukthwa       
  |  |                                 
  +--+  +-------------- Trapezoideus   
     |  |                              
     +--+           +-- Lens_contradens **
        |  +--------+                  
        |  |        +-- Lens_misellus **  
        +--+                           
           |  +-------- Lens_eximius **   
           +--+                        
              |  +----- Lens_pallegoixi **
              +--+                     
                 |  +-- Lens_comptus **   
                 +--+                  
                    +-- Lens_novoselovi **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 119 7 2 16 
  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 3 5 195 7 2 16 
  +----------- Physunio_modelli 
  |                             
  +----------- Physunio         
  |                             
  +----------- Physunio_superbus
--+                             
  +----------- Yaukthwa         
  |                             
  |        +-- Lens_comptus **     
  |  +-----+                    
  |  |     +-- Lens_novoselovi **  
  +--+                          
     |  +----- Lens_eximius **     
     +--+                       
        |  +-- Lens_contradens **  
        +--+                    
           +-- Lens_micropterus **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 5 193 7 2 16 
  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_novoselovi ** 
           +--+                   
              +-- Lens_comptus **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 133 7 2 16 
  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
5 10 0 5 77 5 2 16 
  +-- Lens_eximius **       
  |                      
  +-- Physunio_modelli__1
  |                      
  +-- Yaukthwa_nesemanni 
  |                      
  +-- Yaukthwa_panhai    
  |                      
  +-- Lens_comptus **       
  |                      
--+-- Lens_novoselovi **    
  |                      
  +-- Physunio_modelli__2
  |                      
  +-- Physunio           
  |                      
  +-- Yaukthwa_inlenensis
  |                      
  +-- Lens_contradens **    
  |                      
  +-- Lens_micropterus **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 5 195 7 2 16 
  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
4 4 2 4 134 7 2 17 
           +-- Physunio       
  +--------+                  
  |        +-- Yaukthwa       
--+                           
  |  +-------- Lens_contradens **
  +--+                        
     |  +----- Lens_eximius **   
     +--+                     
        |  +-- Lens_comptus **   
        +--+                  
           +-- Lens_novoselovi **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 4 17 6 3 15 
              +-- Trapezoideus    
  +-----------+                   
  |           +-- Physunio        
--+                               
  |  +----------- Yaukthwa        
  +--+                            
     |  +-------- Lens_contradens ** 
     +--+                         
        |  +----- Lens_novoselovi ** 
        +--+                      
           |  +-- Lens_rolfbrandti **
           +--+                   
              +-- Lens_maenamensis **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 42 5 3 17 
           +-- Physunio       
  +--------+                  
  |        +-- Trapezoideus   
--+                           
  |  +-------- Yaukthwa       
  +--+                        
     |  +----- Lens_eximius **   
     +--+                     
        |  +-- Lens_contradens **
        +--+                  
           +-- Lens_lugens **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 43 1 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Lens_comptus **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 29 5 3 -- 
  +----------- Physunio       
--+                           
  |  +-------- Trapezoideus   
  +--+                        
     |  +----- Yaukthwa       
     +--+                     
        |  +-- Lens_contradens **
        +--+                  
           +-- Lens_eximius **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 5 2 2 -- 
  +----- Physunio       
  |                     
--+----- Trapezoideus   
  |                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 45 1 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 44 1 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 46 1 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 43 1 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Lens_comptus **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
  +-------------- Yaukthwa_elongatula
  |                                  
--+  +----------- Trapezoideus       
  |  |                               
  +--+        +-- Physunio           
     |  +-----+                      
     |  |     +-- Yaukthwa_inlenensis
     +--+                            
        |  +----- Yaukthwa_paiensis  
        +--+                         
           |  +-- Lens_contradens **    
           +--+                      
              +-- Lens_eximius **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
2 2 0 2 78 2 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Lens_comptus **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
  +----- Physunio           
  |                         
  +----- Yaukthwa_paiensis  
  |                         
  +----- Yaukthwa_inlenensis
--+                         
  +----- Trapezoideus       
  |                         
  +----- Yaukthwa_elongatula
  |                         
  |  +-- Lens_contradens **    
  +--+                      
     +-- Lens_eximius **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 34 2 2 -- 
  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Lens_contradens **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 2 1 -- 
  +-- Lens_micropterus **
--+                   
  +-- Physunio

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 140 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Lens_contradens **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 2 2 -- 
-- Lens_contradens **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Lens_contradens **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Lens_contradens **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Lens_contradens **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Physunio       
--+                  
  +-- Lens_contradens **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 4 4 -- 
        +-- Trapezoideus   
  +-----+                  
  |     +-- Physunio       
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Solenaia

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 26 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Lens_contradens **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 3 2 -- 
  +----- Physunio    
--+                  
  |  +-- Lens_eximius **
  +--+               
     +-- Yaukthwa

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 1 -- 
  +-- Lens_micropterus **
--+                   
  +-- Physunio

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 58 2 2 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Trapezoideus

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Physunio       
--+                  
  +-- Lens_contradens **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 4 4 -- 
        +-- Trapezoideus   
  +-----+                  
  |     +-- Physunio       
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Solenaia

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Lens_contradens **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 3 2 -- 
  +----- Physunio    
--+                  
  |  +-- Lens_eximius **
  +--+               
     +-- Yaukthwa

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 94 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Lens_contradens **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 2 -- 
  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio