Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Rectidentini | Contradentina

genus Lens Simpson, 1900: cladomics

13 Recent valid species (13 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 13 (13 total)
analyzed in 73 cladograms in 47 publications
maximum number of terminals in an analysis = 11 (mean = 2.89, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 35
maximum number of included terminals = 11, (mean = 4.94 median = 4)
proportion of cladograms that support basic monophyly = 0.97

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lens cladogram distances
PCoA Ordination

NJ Tree of Lens cladogram distances
NJ Tree

Cladograms

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
4 4 9 17 5 2 1 0 
  +----------------------- Physunio           
  |                                           
  |                    +-- Lens_maenamensis **   
--+  +-----------------+                      
  |  |                 +-- Lens_rolfbrandti **   
  |  |                                        
  |  |  +----------------- Lens_contradens__12 **
  +--+  |                                     
     |  |           +----- Lens_micropterus__3 **
     |  |           |                         
     |  +-----------+  +-- Lens_contradens__10 **
     |  |           |  |                      
     +--+           +--+-- Lens_micropterus__2 **
        |              |                      
        |              +-- Lens_contradens__11 **
        |                                     
        |  +-------------- Lens_contradens__9 ** 
        |  |                                  
        +--+  +----------- Lens_contradens__8 ** 
           |  |                               
           +--+  +-------- Lens_contradens__7 ** 
              |  |                            
              +--+  +----- Lens_contradens__6 ** 
                 |  |                         
                 |  |  +-- Lens_contradens__1 ** 
                 +--+  |                      
                    |  +-- Lens_contradens__2 ** 
                    |  |                      
                    |  +-- Lens_contradens__3 ** 
                    +--+                      
                       +-- Lens_contradens__4 ** 
                       |                      
                       +-- Lens_micropterus__1 **
                       |                      
                       +-- Lens_contradens__5 **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
8 8 6 8 21 7 4 15 
                    +-- Trapezoideus    
  +-----------------+                   
  |                 +-- Physunio        
  |                                     
--+  +----------------- Yaukthwa        
  |  |                                  
  |  |  +-------------- Solenaia        
  +--+  |                               
     |  |     +-------- Lens_misellus **   
     |  |  +--+                         
     +--+  |  |  +----- Lens_inornatus **  
        |  |  +--+                      
        |  |     |  +-- Lens_contradens ** 
        |  |     +--+                   
        +--+        +-- Lens_micropterus **
           |                            
           |        +-- Lens_maenamensis **
           |     +--+                   
           |     |  +-- Lens_eximius **    
           +-----+                      
                 |  +-- Lens_pallegoixi ** 
                 +--+                   
                    +-- Lens_comptus **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
11 11 10 12 621 20 3 14 
                             +-- Physunio_modelli   
  +--------------------------+                      
  |                          +-- Physunio_pellucidus
  |                                                 
--+                          +-- Physunio_massini   
  |  +-----------------------+                      
  |  |                       +-- Physunio_superbus  
  +--+                                              
     |  +----------------------- Physunio           
     |  |                                           
     +--+  +-------------------- Trapezoideus       
        |  |                                        
        |  |  +----------------- Yaukthwa           
        +--+  |                                     
           |  |              +-- Lens_inornatus **     
           |  |        +-----+                      
           |  |        |     +-- Lens_misellus **      
           +--+  +-----+                            
              |  |     |  +----- Lens_contradens **    
              |  |     +--+                         
              |  |        |  +-- Lens_micropterus **   
              |  |        +--+                      
              +--+           +-- Lens_lugens **        
                 |                                  
                 |        +----- Lens_pallegoixi **    
                 |  +-----+                         
                 |  |     |  +-- Lens_comptus **       
                 |  |     +--+                      
                 +--+        +-- Lens_novoselovi **    
                    |                               
                    |  +-------- Lens_maenamensis **   
                    +--+                            
                       |  +----- Lens_eximius__2 **    
                       +--+                         
                          |  +-- Lens_eximius__1 **    
                          +--+                      
                             +-- Lens_rolfbrandti **

Zieritz et al., 2026 [Publ 4812], fig. 2a

iTtTbiriSoTroTroSiD
11 11 9 11 1 1 1 14 
  +----------------- Solenaia        
  |                                  
  |              +-- Lens_inornatus **  
--+  +-----------+                   
  |  |           +-- Lens_misellus **   
  |  |                               
  |  |        +----- Lens_contradens ** 
  +--+  +-----+                      
     |  |     |  +-- Lens_lugens **     
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_comptus **    
        |  |  +--+                   
        +--+     +-- Lens_novoselovi ** 
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_rolfbrandti **
              +--+                   
                 +-- Lens_eximius **

Zieritz et al., 2024 [Publ 4381], fig. 7A

iTtTbiriSoTroTroSiD
11 11 9 11 1 1 1 14 
  +----------------- Solenaia        
  |                                  
  |           +----- Lens_contradens ** 
--+  +--------+                      
  |  |        |  +-- Lens_lugens **     
  |  |        +--+                   
  |  |           +-- Lens_micropterus **
  +--+                               
     |           +-- Lens_inornatus **  
     |  +--------+                   
     |  |        +-- Lens_misellus **   
     |  |                            
     +--+     +----- Lens_maenamensis **
        |  +--+                      
        |  |  |  +-- Lens_eximius **    
        |  |  +--+                   
        +--+     +-- Lens_rolfbrandti **
           |                         
           |  +----- Lens_pallegoixi ** 
           +--+                      
              |  +-- Lens_comptus **    
              +--+                   
                 +-- Lens_novoselovi **

Pfeiffer et al., 2021 [Publ 4126], fig. 12

iTtTbiriSoTroTroSiD
10 10 7 10 1 1 1 14 
  +-------------- Solenaia        
  |                               
  |           +-- Lens_contradens ** 
  |  +--------+                   
--+  |        +-- Lens_micropterus **
  |  |                            
  |  |        +-- Lens_inornatus **  
  |  +--------+                   
  +--+        +-- Lens_misellus **   
     |                            
     |     +----- Lens_maenamensis **
     |  +--+                      
     |  |  |  +-- Lens_rolfbrandti **
     |  |  +--+                   
     +--+     +-- Lens_eximius **    
        |                         
        |  +----- Lens_pallegoixi ** 
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
9 9 7 9 73 10 3 14 
                 +-- Physunio        
  +--------------+                   
  |              +-- Trapezoideus    
  |                                  
--+  +-------------- Yaukthwa        
  |  |                               
  |  |        +----- Lens_misellus **   
  +--+  +-----+                      
     |  |     |  +-- Lens_contradens ** 
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_novoselovi ** 
        |  |  +--+                   
        +--+     +-- Lens_comptus **    
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_eximius **    
              +--+                   
                 +-- Lens_rolfbrandti **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
9 9 7 9 102 14 3 14 
  +-------------------- Physunio        
  |                                     
--+  +----------------- Trapezoideus    
  |  |                                  
  +--+  +-------------- Yaukthwa        
     |  |                               
     |  |        +----- Lens_misellus **   
     +--+  +-----+                      
        |  |     |  +-- Lens_contradens ** 
        |  |     +--+                   
        |  |        +-- Lens_micropterus **
        +--+                            
           |     +----- Lens_maenamensis **
           |  +--+                      
           |  |  |  +-- Lens_eximius **    
           |  |  +--+                   
           +--+     +-- Lens_rolfbrandti **
              |                         
              |  +----- Lens_pallegoixi ** 
              +--+                      
                 |  +-- Lens_novoselovi ** 
                 +--+                   
                    +-- Lens_comptus **

Konopleva et al., 2019 [Publ 4041], fig. 3

iTtTbiriSoTroTroSiD
9 9 7 9 25 10 3 14 
  +----------------- Yaukthwa        
  |                                  
  |              +-- Trapezoideus    
--+  +-----------+                   
  |  |           +-- Physunio        
  |  |                               
  |  |        +----- Lens_misellus **   
  +--+  +-----+                      
     |  |     |  +-- Lens_contradens ** 
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_novoselovi ** 
        |  |  +--+                   
        +--+     +-- Lens_comptus **    
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_eximius **    
              +--+                   
                 +-- Lens_rolfbrandti **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
9 9 7 9 108 14 3 14 
                 +-- Trapezoideus    
  +--------------+                   
  |              +-- Physunio        
  |                                  
--+  +-------------- Yaukthwa        
  |  |                               
  |  |        +----- Lens_misellus **   
  +--+  +-----+                      
     |  |     |  +-- Lens_contradens ** 
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_comptus **    
        |  |  +--+                   
        +--+     +-- Lens_novoselovi ** 
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_eximius **    
              +--+                   
                 +-- Lens_rolfbrandti **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
7 7 5 7 44 9 3 16 
  +----------------------- Yaukthwa        
  |                                        
--+  +-------------------- Physunio        
  |  |                                     
  +--+  +----------------- Trapezoideus    
     |  |                                  
     +--+              +-- Lens_contradens ** 
        |  +-----------+                   
        |  |           +-- Lens_micropterus **
        +--+                               
           |  +----------- Lens_misellus **   
           +--+                            
              |  +-------- Lens_eximius **    
              +--+                         
                 |  +----- Lens_pallegoixi ** 
                 +--+                      
                    |  +-- Lens_comptus **    
                    +--+                   
                       +-- Lens_novoselovi **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
6 6 4 6 25 8 3 18 
  +-------------------- Physunio       
  |                                    
--+  +----------------- Yaukthwa       
  |  |                                 
  +--+  +-------------- Trapezoideus   
     |  |                              
     +--+           +-- Lens_contradens **
        |  +--------+                  
        |  |        +-- Lens_misellus **  
        +--+                           
           |  +-------- Lens_eximius **   
           +--+                        
              |  +----- Lens_pallegoixi **
              +--+                     
                 |  +-- Lens_comptus **   
                 +--+                  
                    +-- Lens_novoselovi **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5 5 3 5 195 7 2 16 
  +----------- Physunio_modelli 
  |                             
  +----------- Physunio         
  |                             
  +----------- Physunio_superbus
--+                             
  +----------- Yaukthwa         
  |                             
  |        +-- Lens_comptus **     
  |  +-----+                    
  |  |     +-- Lens_novoselovi **  
  +--+                          
     |  +----- Lens_eximius **     
     +--+                       
        |  +-- Lens_contradens **  
        +--+                    
           +-- Lens_micropterus **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
5 10 0 5 77 5 2 16 
  +-- Lens_eximius **       
  |                      
  +-- Physunio_modelli__1
  |                      
  +-- Yaukthwa_nesemanni 
  |                      
  +-- Yaukthwa_panhai    
  |                      
  +-- Lens_comptus **       
  |                      
--+-- Lens_novoselovi **    
  |                      
  +-- Physunio_modelli__2
  |                      
  +-- Physunio           
  |                      
  +-- Yaukthwa_inlenensis
  |                      
  +-- Lens_contradens **    
  |                      
  +-- Lens_micropterus **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 133 7 2 16 
  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5 5 3 5 195 7 2 16 
  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5 5 3 5 193 7 2 16 
  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_novoselovi ** 
           +--+                   
              +-- Lens_comptus **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
5 5 3 5 119 7 2 16 
  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
4 4 2 4 134 7 2 17 
           +-- Physunio       
  +--------+                  
  |        +-- Yaukthwa       
--+                           
  |  +-------- Lens_contradens **
  +--+                        
     |  +----- Lens_eximius **   
     +--+                     
        |  +-- Lens_comptus **   
        +--+                  
           +-- Lens_novoselovi **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iTtTbiriSoTroTroSiD
4 4 2 4 17 6 3 15 
              +-- Trapezoideus    
  +-----------+                   
  |           +-- Physunio        
--+                               
  |  +----------- Yaukthwa        
  +--+                            
     |  +-------- Lens_contradens ** 
     +--+                         
        |  +----- Lens_novoselovi ** 
        +--+                      
           |  +-- Lens_rolfbrandti **
           +--+                   
              +-- Lens_maenamensis **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
3 3 1 3 42 5 3 17 
           +-- Physunio       
  +--------+                  
  |        +-- Trapezoideus   
--+                           
  |  +-------- Yaukthwa       
  +--+                        
     |  +----- Lens_eximius **   
     +--+                     
        |  +-- Lens_contradens **
        +--+                  
           +-- Lens_lugens **

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
2 2 0 2 78 2 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Lens_comptus **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
  +-------------- Yaukthwa_elongatula
  |                                  
--+  +----------- Trapezoideus       
  |  |                               
  +--+        +-- Physunio           
     |  +-----+                      
     |  |     +-- Yaukthwa_inlenensis
     +--+                            
        |  +----- Yaukthwa_paiensis  
        +--+                         
           |  +-- Lens_contradens **    
           +--+                      
              +-- Lens_eximius **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 29 5 3 -- 
  +----------- Physunio       
--+                           
  |  +-------- Trapezoideus   
  +--+                        
     |  +----- Yaukthwa       
     +--+                     
        |  +-- Lens_contradens **
        +--+                  
           +-- Lens_eximius **

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
2 2 0 2 44 1 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
2 2 0 2 43 1 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Lens_comptus **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
2 2 0 2 43 1 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Lens_comptus **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 5 2 2 -- 
  +----- Physunio       
  |                     
--+----- Trapezoideus   
  |                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 45 1 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
2 2 0 2 46 1 1 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
2 2 0 2 34 5 3 -- 
  +----- Physunio           
  |                         
  +----- Yaukthwa_paiensis  
  |                         
  +----- Yaukthwa_inlenensis
--+                         
  +----- Trapezoideus       
  |                         
  +----- Yaukthwa_elongatula
  |                         
  |  +-- Lens_contradens **    
  +--+                      
     +-- Lens_eximius **

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Physunio       
--+                  
  +-- Lens_contradens **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 39 2 1 -- 
  +-- Lens_micropterus **
--+                   
  +-- Physunio

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
1 1 0 1 90 3 2 -- 
  +----- Physunio    
--+                  
  |  +-- Lens_eximius **
  +--+               
     +-- Yaukthwa

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 30 2 1 -- 
  +-- Lens_micropterus **
--+                   
  +-- Physunio

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 90 3 2 -- 
  +----- Physunio    
--+                  
  |  +-- Lens_eximius **
  +--+               
     +-- Yaukthwa

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 87 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 58 2 2 -- 
  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Trapezoideus

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 17 0 0 -- 
-- Lens_contradens **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 4 4 -- 
        +-- Trapezoideus   
  +-----+                  
  |     +-- Physunio       
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Solenaia

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Lens_contradens **

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 2 -- 
  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 95 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
1 1 0 1 35 0 0 -- 
-- Lens_contradens **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Lens_contradens **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 46 0 0 -- 
-- Lens_contradens **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 85 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Lens_contradens **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 4 4 -- 
        +-- Trapezoideus   
  +-----+                  
  |     +-- Physunio       
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Solenaia

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 2 2 -- 
-- Lens_contradens **

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Physunio       
--+                  
  +-- Lens_contradens **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 140 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 34 2 2 -- 
  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 19 0 0 -- 
-- Lens_contradens **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 26 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 45 0 0 -- 
-- Lens_contradens **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 91 1 1 -- 
  +-- Lens_contradens **
--+                  
  +-- Physunio