Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Lamproscaphini | Contradentina

genus Lens Simpson, 1900: cladomics

13 Recent valid species (13 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 13 (13 total)
analyzed in 73 cladograms in 47 publications
maximum number of terminals in an analysis = 11 (mean = 2.89, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 35
maximum number of included terminals = 11, (mean = 4.94 median = 4)
proportion of cladograms that support basic monophyly = 0.97

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination of Lens cladogram distances
PCoA Ordination

NJ Tree of Lens cladogram distances
NJ Tree

Cladograms

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram **

cst iT tT bi riS oT roT roS iD
0.800 11 11 10 11 622 20 3 **
Cladomics cladogram Physunio modelli Physunio pellucidus Physunio massini Physunio superbus Physunio Trapezoideus Yaukthwa Lens inornatus Lens misellus Lens contradens Lens micropterus Lens lugens Lens pallegoixi Lens comptus Lens novoselovi Lens maenamensis Lens eximius 2 Lens eximius 1 Lens rolfbrandti

Pfeiffer et al., 2021 [Publ 4126], fig. 9

** Top Sister-Group Cladogram **

cst iT tT bi riS oT roT roS iD
1.000 8 8 6 8 21 7 4 6.0
Cladomics cladogram Trapezoideus Physunio Yaukthwa Solenaia Lens misellus Lens inornatus Lens contradens Lens micropterus Lens maenamensis Lens eximius Lens pallegoixi Lens comptus

Zieritz et al., 2024 [Publ 4381], fig. 7A

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 11 1 1 1 3.0
Cladomics cladogram Solenaia Lens contradens Lens lugens Lens micropterus Lens inornatus Lens misellus Lens maenamensis Lens eximius Lens rolfbrandti Lens pallegoixi Lens comptus Lens novoselovi

Zieritz et al., 2026 [Publ 4812], fig. 2a

cst iT tT bi riS oT roT roS iD
1.000 11 11 9 11 1 1 1 3.0
Cladomics cladogram Solenaia Lens inornatus Lens misellus Lens contradens Lens lugens Lens micropterus Lens pallegoixi Lens comptus Lens novoselovi Lens maenamensis Lens rolfbrandti Lens eximius

Pfeiffer et al., 2021 [Publ 4126], fig. 12

cst iT tT bi riS oT roT roS iD
1.000 10 10 7 10 1 1 1 3.0
Cladomics cladogram Solenaia Lens contradens Lens micropterus Lens inornatus Lens misellus Lens maenamensis Lens rolfbrandti Lens eximius Lens pallegoixi Lens comptus Lens novoselovi

Konopleva et al., 2019 [Publ 4041], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 9 25 10 3 3.0
Cladomics cladogram Yaukthwa Trapezoideus Physunio Lens misellus Lens contradens Lens micropterus Lens pallegoixi Lens novoselovi Lens comptus Lens maenamensis Lens eximius Lens rolfbrandti

Bolotov et al., 2020 [Publ 4100], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 9 108 14 3 3.0
Cladomics cladogram Trapezoideus Physunio Yaukthwa Lens misellus Lens contradens Lens micropterus Lens pallegoixi Lens comptus Lens novoselovi Lens maenamensis Lens eximius Lens rolfbrandti

Bolotov et al., 2020 [Publ 4100], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 9 102 14 3 3.0
Cladomics cladogram Physunio Trapezoideus Yaukthwa Lens misellus Lens contradens Lens micropterus Lens maenamensis Lens eximius Lens rolfbrandti Lens pallegoixi Lens novoselovi Lens comptus

Konopleva et al., 2023 [Publ 4262], fig. 8

cst iT tT bi riS oT roT roS iD
1.000 9 9 7 9 73 10 3 3.0
Cladomics cladogram Physunio Trapezoideus Yaukthwa Lens misellus Lens contradens Lens micropterus Lens pallegoixi Lens novoselovi Lens comptus Lens maenamensis Lens eximius Lens rolfbrandti

Konopleva et al., 2019 [Publ 3900], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 7 7 5 7 44 9 3 7.0
Cladomics cladogram Yaukthwa Physunio Trapezoideus Lens contradens Lens micropterus Lens misellus Lens eximius Lens pallegoixi Lens comptus Lens novoselovi

Zieritz et al., 2020 [Publ 4102], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 6 6 4 6 25 8 3 6.0
Cladomics cladogram Physunio Yaukthwa Trapezoideus Lens contradens Lens misellus Lens eximius Lens pallegoixi Lens comptus Lens novoselovi

Bolotov et al., 2017 [Publ 3761], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 134 7 2 7.0
Cladomics cladogram Physunio Yaukthwa Lens contradens Lens micropterus Lens eximius Lens comptus Lens novoselovi

Bolotov et al., 2017 [Publ 3764], fig. S1

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 120 7 2 7.0
Cladomics cladogram Physunio Yaukthwa Lens contradens Lens micropterus Lens eximius Lens comptus Lens novoselovi

Bolotov et al., 2018 [Publ 3826], fig. 2

cst iT tT bi riS oT roT roS iD
0.250 5 10 0 5 77 5 2 15.0
Cladomics cladogram Ensidens telus Ensidens ingallsianus Ensidens jaculus Hyriopsis desowitzi Hyriopsis myersiana Hyriopsis khoratensis Ensidens spiculus Ensidens sagittarius Lens eximius Physunio modelli 1 Yaukthwa nesemanni Yaukthwa panhai Lens comptus Lens novoselovi Physunio modelli 2 Physunio Yaukthwa inlenensis Lens contradens Lens micropterus

Huang et al., 2019 [Publ 3843], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 196 7 2 7.0
Cladomics cladogram Physunio Yaukthwa Lens contradens Lens micropterus Lens eximius Lens comptus Lens novoselovi

Huang et al., 2019 [Publ 3843], fig. S3

cst iT tT bi riS oT roT roS iD
1.000 5 5 3 5 194 7 2 7.0
Cladomics cladogram Physunio Yaukthwa Lens contradens Lens micropterus Lens eximius Lens novoselovi Lens comptus

Huang et al., 2019 [Publ 3843], fig. S4

cst iT tT bi riS oT roT roS iD
0.750 5 5 3 5 196 7 2 9.0
Cladomics cladogram Physunio modelli Physunio Physunio superbus Yaukthwa Lens comptus Lens novoselovi Lens eximius Lens contradens Lens micropterus

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 17 6 3 8.0
Cladomics cladogram Trapezoideus Physunio Yaukthwa Lens contradens Lens novoselovi Lens rolfbrandti Lens maenamensis

Bolotov et al., 2017 [Publ 3761], fig. S2

cst iT tT bi riS oT roT roS iD
1.000 4 4 2 4 135 7 2 8.0
Cladomics cladogram Physunio Yaukthwa Lens contradens Lens eximius Lens comptus Lens novoselovi

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 4 4 9 4 5 2 1 14.0
Cladomics cladogram Physunio Lens maenamensis Lens rolfbrandti Lens contradens 12 Lens micropterus 3 Lens contradens 10 Lens micropterus 2 Lens contradens 11 Lens contradens 9 Lens contradens 8 Lens contradens 7 Lens contradens 6 Lens contradens 1 Lens contradens 2 Lens contradens 3 Lens contradens 4 Lens micropterus 1 Lens contradens 5

Zieritz et al., 2024 [Publ 4381], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 3 3 1 3 42 5 3 9.0
Cladomics cladogram Physunio Trapezoideus Yaukthwa Lens eximius Lens contradens Lens lugens

Zieritz et al., 2021 [Publ 4218], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 29 5 3 --
Cladomics cladogram Physunio Trapezoideus Yaukthwa Lens contradens Lens eximius

Wu et al., 2024 [Publ 4350], fig. 5a

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 34 5 3 --
Cladomics cladogram Physunio Trapezoideus Yaukthwa Lens contradens Lens eximius

Wu et al., 2024 [Publ 4350], fig. 5b

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 2 34 5 3 --
Cladomics cladogram Yaukthwa elongatula Trapezoideus Physunio Yaukthwa inlenensis Yaukthwa paiensis Lens contradens Lens eximius

Wu et al., 2024 [Publ 4350], fig. 5c

cst iT tT bi riS oT roT roS iD
0.800 2 2 0 2 34 5 3 --
Cladomics cladogram Physunio Yaukthwa paiensis Yaukthwa inlenensis Trapezoideus Yaukthwa elongatula Lens contradens Lens eximius

Wu et al., 2024 [Publ 4350], fig. 6-7

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 34 5 3 --
Cladomics cladogram Physunio Trapezoideus Yaukthwa Lens contradens Lens eximius

Liu et al., 2024 [Publ 4380], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 34 5 3 --
Cladomics cladogram Physunio Trapezoideus Yaukthwa Lens contradens Lens eximius

Wu et al., 2024 [Publ 4388], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 34 5 3 --
Cladomics cladogram Physunio Trapezoideus Yaukthwa Lens contradens Lens eximius

Pfeiffer et al., 2018 [Publ 3827], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 5 2 2 --
Cladomics cladogram Physunio Trapezoideus Lens comptus Lens contradens

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 79 2 1 --
Cladomics cladogram Physunio Lens contradens Lens comptus

Dai et al., 2023 [Publ 4285], fig. 3-BI

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 43 1 1 --
Cladomics cladogram Physunio Lens contradens Lens comptus

Dai et al., 2023 [Publ 4285], fig. 3-ML

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 43 1 1 --
Cladomics cladogram Physunio Lens contradens Lens comptus

Dai et al., 2024 [Publ 4389], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 44 1 1 --
Cladomics cladogram Physunio Lens comptus Lens contradens

Dai et al., 2025 [Publ 4441], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 45 1 1 --
Cladomics cladogram Physunio Lens comptus Lens contradens

Chen et al., 2025 [Publ 4663], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 2 2 0 2 46 1 1 --
Cladomics cladogram Physunio Lens comptus Lens contradens

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 4 4 --
Cladomics cladogram Trapezoideus Physunio Yaukthwa Lens contradens Solenaia

Pfeiffer et al., 2019 [Publ 3937], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 56 4 4 --
Cladomics cladogram Trapezoideus Physunio Yaukthwa Lens contradens Solenaia

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

cst iT tT bi riS oT roT roS iD
0.750 1 1 0 1 121 2 2 --
Cladomics cladogram Lens contradens Ensidens Physunio Rectidens Yaukthwa

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 33 2 2 --
Cladomics cladogram Lens contradens Trapezoideus Physunio

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 34 2 2 --
Cladomics cladogram Lens contradens Trapezoideus Physunio

Wu et al., 2021 [Publ 4366], fig. 2_I

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 3 2 --
Cladomics cladogram Physunio Lens eximius Yaukthwa

Wu et al., 2021 [Publ 4366], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 3 2 --
Cladomics cladogram Physunio Lens eximius Yaukthwa

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 2 2 --
Cladomics cladogram Lens contradens Trapezoideus Physunio

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 75 2 2 --
Cladomics cladogram Lens contradens Trapezoideus Physunio

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 58 2 2 --
Cladomics cladogram Physunio Lens contradens Trapezoideus

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 39 2 1 --
Cladomics cladogram Lens micropterus Physunio

Konopleva et al., 2017 [Publ 3807], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 30 2 1 --
Cladomics cladogram Lens micropterus Physunio

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 47 1 1 --
Cladomics cladogram Lens contradens Physunio

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 22 1 1 --
Cladomics cladogram Lens contradens Physunio

Dai et al., 2023 [Publ 4285], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Lens contradens Physunio

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Lens contradens Physunio

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Lens contradens Physunio

Wu et al., 2024 [Publ 4350], fig. 8a_S1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Lens contradens Physunio

Wu et al., 2024 [Publ 4350], fig. 8b_S2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 90 1 1 --
Cladomics cladogram Lens contradens Physunio

Matos et al., 2024 [Publ 4383], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 1 1 --
Cladomics cladogram Physunio Lens contradens

Matos et al., 2024 [Publ 4384], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 1 1 --
Cladomics cladogram Physunio Lens contradens

Wu et al., 2024 [Publ 4388], fig. 6

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 52 1 1 --
Cladomics cladogram Lens contradens Physunio

Zhang et al., 2024 [Publ 4397], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 26 1 1 --
Cladomics cladogram Lens contradens Physunio

Dai et al., 2025 [Publ 4440], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 86 1 1 --
Cladomics cladogram Lens contradens Physunio

Chen et al., 2025 [Publ 4448], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 84 1 1 --
Cladomics cladogram Lens contradens Physunio

Klymus et al., 2025 [Publ 4814], fig. 3A

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 1 1 --
Cladomics cladogram Lens contradens Physunio

Klymus et al., 2025 [Publ 4814], fig. 3B

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 1 1 --
Cladomics cladogram Lens contradens Physunio

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 139 1 1 --
Cladomics cladogram Lens contradens Physunio

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 1 1 --
Cladomics cladogram Lens contradens Physunio

Wu et al., 2026 [Publ 4811], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 94 1 1 --
Cladomics cladogram Lens contradens Physunio

Graf, 2002 [Publ 2651], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 17 0 0 --
Cladomics cladogram Lens contradens Unio Alasmidonta Pyganodon Gonidea Lampsilis Obliquaria Eurynia Amblema

Graf & Cummings, 2006 [Publ 1968], fig. 2

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 0 0 --
Cladomics cladogram Coelatura Lens contradens Unio Lasmigona Alasmidonta Pyganodon Strophitus Gonidea Quadrula Amblema Eurynia Fusconaia Pleurobema Obliquaria Truncilla Ptychobranchus Ligumia Cambarunio Epioblasma Lampsilis Ortmanniana

Hoeh et al., 2009 [Publ 2716], fig. 3

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 46 0 0 --
Cladomics cladogram Lens contradens Gonidea Pilsbryoconcha Pseudodon Amblema Obliquaria Quadrula Eurynia Pleurobema Fusconaia Ptychobranchus Truncilla Ortmanniana Lampsilis Cambarunio Ligumia Epioblasma

Hoeh et al., 2009 [Publ 2716], fig. 4

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 0 0 --
Cladomics cladogram Lens contradens Unio Lasmigona Alasmidonta Strophitus Pyganodon Gonidea Obliquaria Quadrula Eurynia Pleurobema Fusconaia Ptychobranchus Truncilla Ortmanniana Lampsilis Cambarunio Ligumia Epioblasma

Hoeh et al., 2009 [Publ 2716], fig. 5

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 45 0 0 --
Cladomics cladogram Lens contradens Unio Lasmigona Alasmidonta Strophitus Pyganodon Gonidea Obliquaria Quadrula Amblema Eurynia Pleurobema Fusconaia Truncilla Ptychobranchus Ortmanniana Cambarunio Ligumia Epioblasma

Whelan et al., 2011 [Publ 3380], fig. 1-ML

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Pseudodon Lens contradens

Whelan et al., 2011 [Publ 3380], fig. 1-MP

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 35 0 0 --
Cladomics cladogram Lens contradens Pseudodon Unio Pyganodon Lasmigona Gonidea Pronodularia Potomida Quadrula Pleurobema Amblema Lampsilis Truncilla

Smith, 2021 [Publ 4331], fig. 1

cst iT tT bi riS oT roT roS iD
1.000 1 1 0 1 19 0 0 --
Cladomics cladogram Lens contradens Lamprotula