Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Rectidentini | Rectidentina

genus Rectidens Simpson, 1900: cladomics

3 Recent valid species (3 total species)

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Cladomics Summary

number of species = 3 (3 total)
analyzed in 56 cladograms in 35 publications
maximum number of terminals in an analysis = 3 (mean = 1.09, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 4
maximum number of included terminals = 3, (mean = 2.25 median = 2)
proportion of cladograms that support basic monophyly = 1

Cladograms

Zieritz et al., 2026 [Publ 4812], fig. 7a

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
3 3 0 3 1 1 1 0 
  +----- Hyriopsis            
  |                           
--+  +-- Rectidens_sumatrensis **
  |  |                        
  +--+-- Rectidens_orientalis ** 
     |                        
     +-- Rectidens_lauris **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
2 2 0 2 628 15 2 -- 
  +----------------- Ensidens               
  |                                         
  |              +-- Hyriopsis_delaportei   
  |           +--+                          
--+           |  +-- Hyriopsis_desowitzi    
  |  +--------+                             
  |  |        |  +-- Rectidens_lauris **       
  |  |        +--+                          
  |  |           +-- Rectidens_sumatrensis **  
  +--+                                      
     |  +----------- Hyriopsis_panhai       
     |  |                                   
     |  |        +-- Hyriopsis_bialata      
     +--+  +-----+                          
        |  |     +-- Hyriopsis_khoratensis  
        |  |                                
        +--+     +-- Hyriopsis_kratiensis   
           |  +--+                          
           |  |  +-- Hyriopsis_phuphaniensis
           +--+                             
              |  +-- Hyriopsis_myersiana    
              +--+                          
                 +-- Hyriopsis_sakhonensis

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
2 2 0 2 43 3 2 -- 
  +-------- Ensidens             
--+                              
  |  +----- Hyriopsis            
  +--+                           
     |  +-- Rectidens_sumatrensis **
     +--+                        
        +-- Rectidens_lauris **

Zieritz et al., 2024 [Publ 4381], fig. 6A

iTtTbiriSoTroTroSiD
2 2 0 2 1 1 1 -- 
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Rectidens_lauris **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 94 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 26 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
-- Rectidens_sumatrensis **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 34 2 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens_sumatrensis **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 58 2 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens_sumatrensis **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 30 10 2 -- 
  +----------- Ensidens             
  |                                 
--+  +-------- Hyriopsis_desowitzi  
  |  |                              
  |  +-------- Rectidens_sumatrensis **
  +--+                              
     |  +----- Hyriopsis_panhai     
     |  |                           
     +--+  +-- Hyriopsis_khoratensis
        |  |                        
        +--+-- Hyriopsis_bialata    
           |                        
           +-- Hyriopsis_myersiana

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 20 10 2 -- 
  +----------------- Ensidens               
  |                                         
  |              +-- Hyriopsis_desowitzi    
--+  +-----------+                          
  |  |           +-- Hyriopsis_delaportei   
  |  |                                      
  +--+  +----------- Rectidens_sumatrensis **  
     |  |                                   
     |  |     +----- Hyriopsis_myersiana    
     +--+  +--+                             
        |  |  |  +-- Hyriopsis_bialata      
        |  |  +--+                          
        +--+     +-- Hyriopsis_panhai       
           |                                
           |  +----- Hyriopsis_khoratensis  
           +--+                             
              |  +-- Hyriopsis_phuphaniensis
              +--+                          
                 +-- Hyriopsis_sakhonensis

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
-- Rectidens_sumatrensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 27

iTtTbiriSoTroTroSiD
1 1 0 1 10 9 1 -- 
  +-- Rectidens_sumatrensis **
--+                        
  +-- Hyriopsis

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1 1 0 1 121 1 1 -- 
-- Rectidens_sumatrensis **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 86 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 33 2 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens_sumatrensis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 124 10 2 -- 
  +----------- Ensidens             
  |                                 
--+  +-------- Hyriopsis_desowitzi  
  |  |                              
  |  +-------- Rectidens_sumatrensis **
  +--+                              
     |  +----- Hyriopsis_panhai     
     |  |                           
     +--+  +-- Hyriopsis_myersiana  
        |  |                        
        +--+-- Hyriopsis_bialata    
           |                        
           +-- Hyriopsis_khoratensis

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
1 1 0 1 94 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
1 1 0 1 200 10 2 -- 
  +----- Hyriopsis_desowitzi  
  |                           
  +----- Hyriopsis_myersiana  
  |                           
  +----- Hyriopsis_panhai     
--+                           
  |  +-- Hyriopsis_bialata    
  +--+                        
  |  +-- Hyriopsis_khoratensis
  |                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Ensidens

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
1 1 0 1 198 10 2 -- 
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+  +-------- Rectidens_sumatrensis **
     |  |                              
     +--+     +-- Hyriopsis_panhai     
        |  +--+                        
        |  |  +-- Hyriopsis_khoratensis
        +--+                           
           |  +-- Hyriopsis_bialata    
           +--+                        
              +-- Hyriopsis_myersiana

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +-- Ensidens             
  |                        
--+-- Hyriopsis            
  |                        
  +-- Rectidens_sumatrensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 30

iTtTbiriSoTroTroSiD
1 1 0 1 6 6 1 -- 
  +-- Rectidens_sumatrensis **
--+                        
  +-- Ensidens

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
1 1 0 1 28 12 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
1 1 0 1 47 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 30 3 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
1 1 0 1 46 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 50 10 2 -- 
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+----------- Rectidens_sumatrensis **
     |                                 
     |  +-------- Hyriopsis_panhai     
     +--+                              
        |  +----- Hyriopsis_myersiana  
        +--+                           
           |  +-- Hyriopsis_khoratensis
           +--+                        
              +-- Hyriopsis_bialata

Zieritz et al., 2016 [Publ 3804], fig. S2-1

iTtTbiriSoTroTroSiD
1 1 0 1 5 5 1 -- 
  +-- Rectidens_sumatrensis **
--+                        
  +-- Hyriopsis

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 116 11 2 -- 
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+  +-------- Rectidens_sumatrensis **
     |  |                              
     +--+     +-- Hyriopsis_khoratensis
        |  +--+                        
        |  |  +-- Hyriopsis_bialata    
        +--+                           
           |  +-- Hyriopsis_myersiana  
           +--+                        
              +-- Hyriopsis_panhai

Konopleva et al., 2019 [Publ 4041], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 33 11 2 -- 
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_dugasti     
  |                        
  +-- Hyriopsis_khoratensis
  |                        
  +-- Hyriopsis_bialata    
  |                        
  +-- Hyriopsis_myersiana  
  |                        
--+-- Hyriopsis_desowitzi  
  |                        
  +-- Rectidens_sumatrensis **
  |                        
  +-- Hyriopsis_panhai     
  |                        
  +-- Ensidens_jaculus__1  
  |                        
  +-- Ensidens_jaculus__2  
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
1 1 0 1 94 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iTtTbiriSoTroTroSiD
1 1 0 1 6 2 2 -- 
  +-- Ensidens    
  |               
  +-- Hyriopsis__1
--+               
  +-- Hyriopsis__2
  |               
  +-- Rectidens **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
1 1 0 1 90 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1 1 0 1 139 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
1 1 0 1 81 15 2 -- 
  +----------------------- Ensidens               
  |                                               
--+                    +-- Hyriopsis_desowitzi    
  |  +-----------------+                          
  |  |                 +-- Hyriopsis_delaportei   
  +--+                                            
     |  +----------------- Rectidens_sumatrensis **  
     |  |                                         
     +--+  +-------------- Hyriopsis_panhai       
        |  |                                      
        +--+  +----------- Hyriopsis_myersiana    
           |  |                                   
           +--+        +-- Hyriopsis_khoratensis  
              |  +-----+                          
              |  |     +-- Hyriopsis_bialata      
              +--+                                
                 |  +----- Hyriopsis_sakhonensis  
                 +--+                             
                    |  +-- Hyriopsis_phuphaniensis
                    +--+                          
                       +-- Hyriopsis_kratiensis

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 84 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens_sumatrensis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
1 1 0 1 80 6 2 -- 
  +----- Hyriopsis_desowitzi  
  |                           
  +----- Rectidens_sumatrensis **
--+                           
  +----- Ensidens             
  |                           
  |  +-- Hyriopsis_khoratensis
  +--+                        
     +-- Hyriopsis_myersiana

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
1 1 0 1 19 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 110 11 2 -- 
  +----------------- Ensidens             
--+                                       
  |  +-------------- Hyriopsis_desowitzi  
  +--+                                    
     |  +----------- Rectidens_sumatrensis **
     +--+                                 
        |  +-------- Hyriopsis_panhai     
        +--+                              
           |  +----- Hyriopsis_myersiana  
           +--+                           
              |  +-- Hyriopsis_khoratensis
              +--+                        
                 +-- Hyriopsis_bialata

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
  +-------- Ensidens             
--+                              
  |  +----- Hyriopsis_bialata    
  +--+                           
     |  +-- Hyriopsis_desowitzi  
     +--+                        
        +-- Rectidens_sumatrensis **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
1 1 0 1 35 3 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
1 1 0 1 138 10 2 -- 
  +-------------- Ensidens             
  |                                    
--+           +-- Hyriopsis_bialata    
  |  +--------+                        
  |  |        +-- Hyriopsis_khoratensis
  +--+                                 
     |  +-------- Hyriopsis_panhai     
     +--+                              
        |  +----- Hyriopsis_myersiana  
        +--+                           
           |  +-- Rectidens_sumatrensis **
           +--+                        
              +-- Hyriopsis_desowitzi

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
1 1 0 1 138 10 2 -- 
        +-- Hyriopsis_desowitzi  
        |                        
  +-----+-- Rectidens_sumatrensis **
  |     |                        
  |     +-- Ensidens             
--+                              
  |  +----- Hyriopsis_panhai     
  |  |                           
  +--+----- Hyriopsis_khoratensis
     |                           
     |  +-- Hyriopsis_myersiana  
     +--+                        
        +-- Hyriopsis_bialata

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
1 1 0 1 35 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
1 1 0 1 200 10 2 -- 
  +----- Hyriopsis_desowitzi  
  |                           
  +----- Rectidens_sumatrensis **
  |                           
--+----- Ensidens             
  |                           
  |  +-- Hyriopsis_bialata    
  |  |                        
  |  +-- Hyriopsis_myersiana  
  +--+                        
     +-- Hyriopsis_panhai     
     |                        
     +-- Hyriopsis_khoratensis

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
1 1 0 1 90 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
1 1 0 1 22 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
1 1 0 1 56 2 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens_sumatrensis **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
1 1 0 1 52 1 1 -- 
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
1 1 0 1 75 2 2 -- 
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis