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Taxonomy

all genera

Unionidae

Margaritiferidae

Hyriidae

Etheriidae

Mycetopodidae

Iridinidae


Species & Genus Richness by
Taxonomy

Geography

Summary

North America

Central America

South America

Afrotropics

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East Asia

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Species Richness & Endemism by
Geography

Data last updated
Mon 1 Dec 2025

Publication Pfeiffer & Graf (2015): summary

publication summary | cladograms

Full Citation

Pfeiffer, J.M. & D.L. Graf. 2015. Evolution of bilaterally asymmetrical larvae in freshwater mussels (Bivalvia: Unionoida: Unionidae). Zoological Journal of the Linnean Society 175(2): 307-318. Available electronically.

Attributed Nominal Species

Velesunio ambiguous Graf & Cummings, 2006 = Velesunio ambiguus

‘Philippi’ Graf & Cummings, 2006. Zool. J. Linn. Soc. 148: 348, table 2.
‘Philippi’ Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): [4].
(error for Unio ambiguus Philippi, 1847)

Radiatula bonneaudii Whelan et al., 2011 = Radiatula chaudhurii

‘Eydoux’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): [4].
Bolotov et al., 2017. Sci. Repts. 7: 10, Table 3 [as ‘sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: [5].
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Pfeiffer et al., 2018. J. Moll. Studies 84(4): [4]. [in part]
Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25.
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Radiatula bonneaudii sp. 1’].
Huang et al., 2019. MPE 130: Table S4.
Ortiz-Sepulveda et al., 2020. MPE 148: Table S1.
(misidentified reference to Unio bonneaudii Eydoux, 1838)

image of type lot

Pseudodon cambodjensis Pfeiffer & Graf, 2013 = Pseudodon cambodjensis

‘Petit’ of Authors.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Zieritz et al., 2016. Sci. Total Environ. 571: 1071.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’].
Lopes-Lima et al., 2017. MPE 106: 177.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’].
Razak et al., 2018. Conserv. Genet. Resour. 11: 459.
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina cambodjensis’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Konopleva et al., 2021. Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)].
Pfeiffer et al., 2021. Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’].
Dai et al., 2023. Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’].
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
(reference to Monocondylea cambodjensis Petit de la Saussaye, 1865)
Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.

Contradens contradens Pfeiffer & Graf, 2015 = Lens micropterus

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Lens aff. contradens’].
(misidentified reference to Unio contradens Lea, 1838)

image of type lot

Lamellidens corrianus Whelan et al., 2011 = Trapezidens dolichorhynchus

‘Lea’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Lamellidens corrianus sp. 1].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Pfeiffer et al., 2018. J. Moll. Studies 84(4): [3].
Pfeiffer et al., 2019. MPE 137: 117.
Lopes-Lima et al., 2020. MPE 146: Table 1.
(misidentified reference to Unio corrianus Lea, 1834)
Type(s): SEQUENCED SPECIMEN UMMZ_304642, Ayeryarwaddy River, mud bank on E side of river, Mandalay Division, Burma. 21.48187, 95.30501.

There are 15 nominal species attributed to this publication (none available).
image of type lot

Scabies crispata Pfeiffer & Graf, 2015 = Scabies mandarinus

‘Gould’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): [4].
Ng et al., 2016. PLoS One 11(8), e0161130: [5].
Bolotov et al., 2017. Sci. Repts. 7: [6], Table S1 [as ‘Scabies aff. crispata’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
(misidentified reference to Unio crispata Gould, 1843)
Type(s): SEQUENCED SPECIMEN UMMZ_304646, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.

image of type lot

Pseudodon cumingii Pfeiffer & Graf, 2015 = Nyeinchanconcha nyeinchani

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘”Bineurus” cumingii’].
Huang et al., 2019. MPE 130: Table S4 [as ‘”Bineurus” cumingii’].
Lopes-Lima et al., 2020. MPE 146: Table 1.
(misidentified reference to Anodonta cumingii Lea, 1851)
Type(s): SPECIMEN UMMZ_304648, Nam Phiat, Phonethong, ca. 2 km from confluence, Bolikhamxai Province, Laos. 18.08389, 104.97806.

image of type lot

Pilsbryoconcha exilis Pfeiffer & Graf, 2015 = Pilsbryoconcha carinifera

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Ng et al., 2016. PLoS One 11(8), e0161130: [5]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘P. aff. exilis sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘P. aff. exilis sp. 1’].
Lopes-Lima et al., 2020. MPE 146: Table 1 [as ‘P. aff. exilis sp. 1’].
Zieritz et al., 2020. Biodivers. Conserv. 29: 2244.
Wu et al., 2021. ZooKeys 1054: 87.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘P. aff. exilis’].
Dai et al., 2023. Invert. Syst. 37(2): Tables 1-2. [in part]
(misidentified reference to Anodonta exilis Lea, 1838)
Type(s): SEQUENCED SPECIMEN UMMZ_304647, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.

Trapezoideus exolescens Pfeiffer & Graf, 2013 = unionoid species

‘Gould’ of Authors.
Brandt, 1974. Arch. Moll. 105: 299.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): [4].
Lopes-Lima et al., 2017. MPE 106: 177.
(misidentified reference to Unio exolescens Gould, 1843)

Lamellidens generosus Whelan et al., 2011 = Lamellidens savadiensis

‘Gould’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Araujo et al., 2017. Zool. J. Linn. Soc. 179(2): 280.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Pfeiffer et al., 2018. J. Moll. Studies 84(4): [3].
Pfeiffer et al., 2019. MPE 137: 117.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio generosus Gould, 1847)

Pseudodon inoscularis Pfeiffer & Graf, 2013 = Sundadontina tumida

‘Gould’ of Authors.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon cf. inoscularis’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
Huang et al., 2019. MPE 130: Table S4 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
Lopes-Lima et al., 2020. MPE 146: Table 1 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
(misidentified reference to Anodon inoscularis Gould, 1844)

Pseudodon mouhotii Pfeiffer & Graf, 2015 = Bineurus exilis

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. mouhotii sp. 3’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Bineurus aff. mouhotii sp. 3’].
(misidentified reference to Monocondyloea mouhotii Lea, 1863)

Ensidens sagittarius Pfeiffer & Graf, 2015 = Ensidens telus

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 1].
Huang et al., 2019. MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 1].
Zieritz et al., 2020. Biodivers. Conserv. 29: 2242.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio sagittarius Lea, 1856)

Pseudodon vondembuschianus Pfeiffer & Graf, 2015 = Pseudodon vondembuschianus

‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 2’]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
(reference to Margaritana vondembuschiana Lea, 1840)

Pseudodon vondembuschianus Graf, 2002 = Pseudodon vondembuschianus

‘Lea’ of Authors.
Graf, 2002. J. Moll. Studies 68: 66.
Graf & Cummings, 2006. Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009. Malacologia 51(2): 310.
Whelan et al., 2011. MPE 61: 506. [in part]
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 2’]. [in part]
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
(reference to Margaritana vondembuschiana Lea, 1840)

Taxonomic Opinions

Amblema plicata (Say, 1817)

Castalia ambigua Lamarck, 1819

Chambardia wahlbergi (Krauss, 1848)

Coelatura aegyptiaca (Cailliaud, 1827)

Contradens contradens (Lea, 1838) = Lens contradens
+ Physunio modelli Brandt, 1974

Contradens sp. = Physunio modelli

Ensidens sagittarius (Lea, 1856)

Etheria elliptica Lamarck, 1807

Gonidea angulata (Lea, 1838)

Hyridella australis (Lamarck, 1819)

Lamellidens corrianus (Lea, 1834)

Lamellidens generosus (Gould, 1847)

Lamproscapha ensiformis (Spix & Wagner, 1827)

Lamprotula caveata (Heude, 1877)

Lampsilis cardium Rafinesque, 1820

Lasmigona costata
+ Lasmigona compressa Graf & Ó Foighil, 2000

Margaritifera margaritifera (Linnaeus, 1758)

Margaritifera monodonta (Say, 1829) = Cumberlandia monodonta

Neotrigonia margaritacea (Lamarck, 1804)

Oxynaia pugio (Benson, 1862) = Indochinella pugio

Parreysia olivacea Prashad, 1930 = Leoparreysia olivacea

Parreysia tavoyensis (Gould, 1843) = Leoparreysia tavoyensis

Pilsbryoconcha exilis (Lea, 1838)

Potamilus alatus (Say, 1817)

Potomida littoralis (Cuvier, 1798)

Pronodularia japanensis (Lea, 1859)

Pseudodon cambodjensis (Petit, 1865)

Pseudodon cumingii (Lea, 1850) = Sundadontina cumingii

Pseudodon inoscularis (Gould, 1844)

Pseudodon mouhotii (Lea, 1863) = Bineurus mouhotii

Pseudodon vondembuschianus (Lea, 1840)

Pyganodon grandis (Say, 1829)

Quadrula quadrula (Rafinesque, 1820)

Radiatula bonneaudii (Eydoux, 1838) = Indonaia bonneaudii

Scabies crispata (Gould, 1843)

Trapezoideus exolescens (Gould, 1843) = Trapezidens exolescens

Triplodon corrugatus (Lamarck, 1819)

Truncilla truncata Rafinesque, 1820

Unio pictorum (Linnaeus, 1758)

Velesunio ambiguous [sic] (Philippi, 1847) = Velesunio ambiguus

 
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