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Data last updated
Mon 1 Dec 2025
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Publication Pfeiffer & Graf (2015): summarypublication summary | cladograms
Full CitationPfeiffer, J.M. & D.L. Graf. 2015. Evolution of bilaterally asymmetrical larvae in freshwater mussels (Bivalvia: Unionoida: Unionidae). Zoological Journal of the Linnean Society 175(2): 307-318. Available electronically.
Attributed Nominal Species
Radiatula bonneaudii Whelan et al., 2011 = Radiatula chaudhurii
‘Eydoux’ of Authors. Whelan et al., 2011. MPE 61: 506. Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): [4]. Bolotov et al., 2017. Sci. Repts. 7: 10, Table 3 [as ‘sp. 1’]. Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. Bolotov et al., 2017. Sci. Repts. 7: [5]. Bolotov et al., 2018. Sci. Repts. 8: Table S1. Pfeiffer et al., 2018. J. Moll. Studies 84(4): [4]. [in part] Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25. Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Radiatula bonneaudii sp. 1’]. Huang et al., 2019. MPE 130: Table S4. Ortiz-Sepulveda et al., 2020. MPE 148: Table S1. (misidentified reference to Unio bonneaudii Eydoux, 1838)
 Pseudodon cambodjensis Pfeiffer & Graf, 2013 = Pseudodon cambodjensis
‘Petit’ of Authors. Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251. Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. Zieritz et al., 2016. Sci. Total Environ. 571: 1071. Bolotov et al., 2017. Sci. Repts. 7: Table S1. Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’]. Lopes-Lima et al., 2017. MPE 106: 177. Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’]. Razak et al., 2018. Conserv. Genet. Resour. 11: 459. Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina cambodjensis’]. Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’]. Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)]. Konopleva et al., 2021. Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)]. Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)]. Pfeiffer et al., 2021. Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’]. Wu et al., 2021. ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’]. Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1. Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’]. Dai et al., 2023. Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’]. Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2. Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1. (reference to Monocondylea cambodjensis Petit de la Saussaye, 1865) Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.
Contradens contradens Pfeiffer & Graf, 2015 = Lens micropterus
‘Lea’ of Authors. Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’]. Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’]. Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Contradens aff. contradens sp. 1’]. Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1 [as ‘Contradens aff. contradens sp. 1’]. Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Contradens aff. contradens sp. 1’]. Huang et al., 2019. MPE 130: Table S4 [as ‘Contradens aff. contradens sp. 1’]. Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Contradens aff. contradens sp. 1’]. Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Lens aff. contradens’]. (misidentified reference to Unio contradens Lea, 1838)
‘Lea’ of Authors. Whelan et al., 2011. MPE 61: 506. Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Lamellidens corrianus sp. 1]. Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. Pfeiffer et al., 2018. J. Moll. Studies 84(4): [3]. Pfeiffer et al., 2019. MPE 137: 117. Lopes-Lima et al., 2020. MPE 146: Table 1. (misidentified reference to Unio corrianus Lea, 1834) Type(s): SEQUENCED SPECIMEN UMMZ_304642, Ayeryarwaddy River, mud bank on E side of river, Mandalay Division, Burma. 21.48187, 95.30501.
There are 15 nominal species attributed
to this publication (none available).
 Scabies crispata Pfeiffer & Graf, 2015 = Scabies mandarinus
‘Gould’ of Authors. Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): [4]. Ng et al., 2016. PLoS One 11(8), e0161130: [5]. Bolotov et al., 2017. Sci. Repts. 7: [6], Table S1 [as ‘Scabies aff. crispata’]. Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. (misidentified reference to Unio crispata Gould, 1843) Type(s): SEQUENCED SPECIMEN UMMZ_304646, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.
‘Lea’ of Authors. Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. Lopes-Lima et al., 2017. MPE 106: 177. Bolotov et al., 2017. Sci. Repts. 7: Table S1. Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘”Bineurus” cumingii’]. Huang et al., 2019. MPE 130: Table S4 [as ‘”Bineurus” cumingii’]. Lopes-Lima et al., 2020. MPE 146: Table 1. (misidentified reference to Anodonta cumingii Lea, 1851) Type(s): SPECIMEN UMMZ_304648, Nam Phiat, Phonethong, ca. 2 km from confluence, Bolikhamxai Province, Laos. 18.08389, 104.97806.
‘Lea’ of Authors. Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. Ng et al., 2016. PLoS One 11(8), e0161130: [5]. [in part] Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’]. Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’]. Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘P. aff. exilis sp. 1’]. Huang et al., 2019. MPE 130: Table S4 [as ‘P. aff. exilis sp. 1’]. Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘P. aff. exilis sp. 1’]. Lopes-Lima et al., 2020. MPE 146: Table 1 [as ‘P. aff. exilis sp. 1’]. Zieritz et al., 2020. Biodivers. Conserv. 29: 2244. Wu et al., 2021. ZooKeys 1054: 87. Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1. Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1. Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘P. aff. exilis’]. Dai et al., 2023. Invert. Syst. 37(2): Tables 1-2. [in part] (misidentified reference to Anodonta exilis Lea, 1838) Type(s): SEQUENCED SPECIMEN UMMZ_304647, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.
Pseudodon mouhotii Pfeiffer & Graf, 2015 = Bineurus exilis
‘Lea’ of Authors. Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. Lopes-Lima et al., 2017. MPE 106: 177. Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. mouhotii sp. 3’]. Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’]. Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’]. Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’]. Huang et al., 2019. MPE 130: Table S4 [as ‘Bineurus aff. mouhotii sp. 3’]. (misidentified reference to Monocondyloea mouhotii Lea, 1863)
Ensidens sagittarius Pfeiffer & Graf, 2015 = Ensidens telus
‘Lea’ of Authors. Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. Lopes-Lima et al., 2017. MPE 106: 177. Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1]. Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1]. Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Ensidens aff. sagittarius sp. 1]. Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 1]. Huang et al., 2019. MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 1]. Zieritz et al., 2020. Biodivers. Conserv. 29: 2242. Lopes-Lima et al., 2020. MPE 146: Table 1. Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1. (misidentified reference to Unio sagittarius Lea, 1856)
‘Lea’ of Authors. Graf, 2002. J. Moll. Studies 68: 66. Graf & Cummings, 2006. Zool. J. Linn. Soc. 148: 348. Hoeh et al., 2009. Malacologia 51(2): 310. Whelan et al., 2011. MPE 61: 506. [in part] Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. [in part] Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part] Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 2’]. [in part] Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part] Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part] Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part] Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part] (reference to Margaritana vondembuschiana Lea, 1840)
Taxonomic OpinionsAmblema plicata (Say, 1817) Castalia ambigua Lamarck, 1819 Chambardia wahlbergi (Krauss, 1848) Coelatura aegyptiaca (Cailliaud, 1827) Contradens contradens (Lea, 1838) = Lens contradens + Physunio modelli Brandt, 1974 Contradens sp. = Physunio modelli Ensidens sagittarius (Lea, 1856) Etheria elliptica Lamarck, 1807 Gonidea angulata (Lea, 1838) Hyridella australis (Lamarck, 1819) Lamellidens corrianus (Lea, 1834) Lamellidens generosus (Gould, 1847) Lamproscapha ensiformis (Spix & Wagner, 1827) Lamprotula caveata (Heude, 1877) Lampsilis cardium Rafinesque, 1820 Lasmigona costata + Lasmigona compressa Graf & Ó Foighil, 2000 Margaritifera margaritifera (Linnaeus, 1758) Margaritifera monodonta (Say, 1829) = Cumberlandia monodonta Neotrigonia margaritacea (Lamarck, 1804) Oxynaia pugio (Benson, 1862) = Indochinella pugio Parreysia olivacea Prashad, 1930 = Leoparreysia olivacea Parreysia tavoyensis (Gould, 1843) = Leoparreysia tavoyensis Pilsbryoconcha exilis (Lea, 1838) Potamilus alatus (Say, 1817) Potomida littoralis (Cuvier, 1798) Pronodularia japanensis (Lea, 1859) Pseudodon cambodjensis (Petit, 1865) Pseudodon cumingii (Lea, 1850) = Sundadontina cumingii Pseudodon inoscularis (Gould, 1844) Pseudodon mouhotii (Lea, 1863) = Bineurus mouhotii Pseudodon vondembuschianus (Lea, 1840) Pyganodon grandis (Say, 1829) Quadrula quadrula (Rafinesque, 1820) Radiatula bonneaudii (Eydoux, 1838) = Indonaia bonneaudii Scabies crispata (Gould, 1843) Trapezoideus exolescens (Gould, 1843) = Trapezidens exolescens Triplodon corrugatus (Lamarck, 1819) Truncilla truncata Rafinesque, 1820 Unio pictorum (Linnaeus, 1758) Velesunio ambiguous [sic] (Philippi, 1847) = Velesunio ambiguus
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